A phylogenetic analysis of the subfamilies of Anyphaenidae (Arachnida, Araneae)

1995 ◽  
Vol 26 (4) ◽  
pp. 361-384 ◽  
Author(s):  
Martin J. Ramírez

AbstractAnyphaenidae are diagnosed by the pattern of tracheal system, the forwardly advanced tracheal spiracle, and the lamelliform claw tuft setae. Cladistic analyses of the relationships of the anyphaenid genera are made from a selection of 9 exemplar taxa scored for 20 characters. In result, three subfamilies are proposed. Malenellinae subfam. n. is erected for Malenella nana gen. & sp. n. from southern Chile. This subfamily is considered as the sister group of all other anyphaenids. Anyphaeninae and Amaurobioidinae form a monophyletic group united by the orientation of the claw tuft setae and the grooved cymbial tip. Anyphaeninae are diagnosed by three characters associated with the advancement of the spiracle toward the epigastric furrow. Amaurobioidinae are diagnosed by the ingression of the median hematodocha in the male tegulum, and a characteristic secondary conductor. The relationships among the genera of Amaurobioidinae are discussed. The complex tracheal system of anyphaenids develops, during ontogeny, from a simple four-branched system. The genera Aysenia (from Clubionidae), Philisca (from Miturgidae) and Sanogasta (from Corinnidae) are transferred to Anyphaenidae: Amaurobioidinae.

Zootaxa ◽  
2002 ◽  
Vol 87 (1) ◽  
pp. 1 ◽  
Author(s):  
FRANK LOUIS CARLE ◽  
KARL M. KJER

Phylogenetic analysis was performed on a set of 242 morphological characters. The taxon sample included 31 Libellula, and representative species from selected libeluline tribes, from all libellulid subfamilies, from all libelluloid families, from all anisopteran superfamilies, and Epiophlebia. Corduliinae was shown to be paraphyletic even among genera characterized by a well developed anal loop bisector. Sympetrini was found to be polyphyletic with Crocothemis the sister group to Libellulini. The traditional placement of Trameini, far from Libellulini is in doubt, because it is here placed as the sister group to Crocothemis + Libellulini. Kennedy’s phylogeny of Libellula was largely corroborated, with the following exceptions: the subgenera Libellula, Eolibellula, and Syntetrum form a monophyletic group which is the sister group to a clade including Belonia, Holotania, Neotetrum, and Eotainia subgenus nov. [type species Mesothemis composita Hagen]; and Eurothemis is determined to be the sister group of Ladona instead of Neotetrum. In addition we confirm Belonia to be monophyletic, and find Platetrum + Plathemis to form a monophyletic group, sister to Ladona + Eurothemis; these four subgenera together form the sister group to Libellula sensu stricto (s.s.).


2011 ◽  
Vol 42 (3) ◽  
pp. 237-275 ◽  
Author(s):  
Celso Azevedo ◽  
Diego Barbosa

AbstractAs currently recognised, Laelius Ashmead contains 47 known species worldwide and is found on all continents except Australia (until this study). In this paper, we examined the taxonomic limits of this genus and tested whether species of the genera Laelius and Prolaelius Kieffer constitute a monophyletic group. We also tested whether the junior synonyms Allepyris Kieffer and Paralaelius Kieffer are supported by cladistic analyses. Our initial analysis indicated that these genera form a single clade. Cladistic analyses were based on 108 female structural characters. The data set was analyzed under equal-weights parsimony and implied weighting. In both analyses, Laelius was retrieved as a polyphyletic group because Prolaelius Kieffer was always nested within Laelius and Allepyris berlandi Benoit always remained in the outgroup and was retrieved as the sister group of Disepyris sp. Based on our analyses, we propose nine nomenclatural acts and recognise 52 valid species of Laelius. The following four new species are described: Laelius haplos sp.n. and L. quadrangulus sp.n. from Australia and L. ogmos sp.n. and L. titanokkos sp.n. from ailand. The following new generic synonym is proposed: Prolaelius Kieffer syn.n. of Laelius Ashmead. The following three new combinations are proposed: Disepyris berlandi (Benoit) comb.n. from Laelius, Laelius firmipennis (Cameron) comb.n. and Laelius glossinae (Turner & Waterston) comb.n. from Prolaelius. Additionally, a lectotype is designated for Laelius fulvipes Kieffer.


1992 ◽  
Vol 5 (6) ◽  
pp. 717 ◽  
Author(s):  
PA Gradek ◽  
CJ Quinn ◽  
JE Rodman ◽  
KG Karol ◽  
E Conti ◽  
...  

The affinities of the Australian monotypic endemic family Akaniaceae, traditionally assigned to the Sapindales, are reassessed on the basis of comparative sequence data for the chloroplast encoded gene, rbcL. Cladistic analyses show Akania to cluster robustly with Bretschneidera and then Tropaeolum, within the clade of glucosinolate Capparalean families. Eight species representing six other families assigned to the Sapindales, plus Leitneria, formed a monophyletic cluster in 100% of trees in a bootstrap analysis with 500 replicates. This Sapindalean clade is shown to be supported by 17 synapomorphs, only one of which occurs in Akania. Relationships at the ordinal level, among the Sapindalean, Malvalean, Capparalean and Myrtalean clades, are, however, not well resolved. While the most parsimonious arrangement has the Malvales as sister-group to the Sapindales, with the Capparalean and Myrtalean clades joining in sequence, the occurrence of an apomorphic triplet of bases at positions 294–6 in all members of the Malvales, Myrtales and Sapindales so far examined is tentative evidence that these orders may constitute a monophyletic group.


2017 ◽  
Vol 31 (5) ◽  
pp. 519 ◽  
Author(s):  
Arnaud Henrard ◽  
Rudy Jocqué

The new zodariid genus Suffascar, gen. nov. is described from Madagascar. It is apparently endemic to the island, common and speciose. Twelve new species were discovered and are here described: Suffascar albolineatus, sp. nov., S. fianara, sp. nov., S. fisheri, sp. nov., S. fitzpatrickae, sp. nov., S. gigas, sp. nov., S. griswoldi, sp. nov., S. macromma, sp. nov., S. micromma, sp. nov., S. nonus, sp. nov., S. scutatus, sp. nov., S. sufficiens, sp. nov. and S. tofti, sp. nov. It is well known that Zodariidae with femoral organs are specialised consumers of ants or termites, although the exact role of these glands still remains obscure. Suffascar is part of the ‘dual femoral organ clade’ (DF), which also includes Asceua Thorell, 1887, Suffasia Jocqué, 1991, and Suffrica Henrard & Jocqué, 2015. Using a morphology-based phylogenetic analysis, we test whether zodariids sharing the dual femoral organ are part of the same group as the ‘single femoral organ clade’ (SF) or form a separate monophyletic group. It appears that SF and DF constitute clearly monophyletic entities with a sister-group relation. These groups may be considered as distinct subfamilies and, together, they may turn out to be sister to the Storeninae (which then could be re-established). An analysis that focuses on these questions is underway. http://zoobank.org/urn:lsid:zoobank.org:pub:281E1357-FAEE-4300-B1BB-F0D43E724E91


1997 ◽  
Vol 67 (2) ◽  
pp. 125-141 ◽  
Author(s):  
Christopher C. Tudge

A phylogenetic analysis of selected anomuran, thalassinidean, and other decapod crustacean taxa, based on spermatozoal ultrastructural characters and spermatophore morphological characters, was performed and the following relationships of the taxa are elucidated from the trees produced. The Anomura are not a monophyletic assemblage, with the lomoid Lomis being exclusive of the remainder of the anomuran taxa, and the thalassinid Thalassina included in the anomuran clade. The synapomorphy joining the majority of the conventional anomuran taxa (Lomis excluded) is the cytoplasmic origin of the microtubular arms. When the palinurid and thalassinoid representatives are separately designated as outgroups, the Astacidea and Brachyura jointly formed a sister group to the Anomura. The superfamilies Thalassinoidea, Paguroidea, and Galatheoidea are not monophyletic groups. In all analyses the anomuran families Coenobitidae and Porcellanidae each form a monophyletic group. The paguroid family Diogenidae is paraphyletic, with the genera Clibanarius and Cancellus separate from a single clade containing the remaining diogenid genera. The families Paguridae and Parapaguridae form a monophyletic clade with the exception of Porcellanopagurus. The two representatives of the family Chirostylidae (Eumunida and Uroptychus) fail to associate with the other species in the Galatheoidea. The taxa in the family Galatheidae are not a monophyletic assemblage. The only investigated hippoid Hippa is portrayed as the sister group to the remainder of the anomuran taxa (with the exception of Lomis).


1989 ◽  
Vol 67 (11) ◽  
pp. 2731-2739 ◽  
Author(s):  
R. O. Brinkhurst

Though the Lumbriculidae is a monotypic taxon including Kurenkovia, there is no support from cladistic analyses for the Kurenkovidae. The unusual nature of the vasa deferentia, which run between the epithelial and muscular layers of the body wall as in some Haplotaxidae and Glossoscolecidae, may affect that decision. The Dorydrilidae is a sister-group to the Lumbriculidae. Lamprodrilus and the Lake Baikal endemic genera are now seen as advanced forms which may have given rise to some Asian and Pacific rim North American taxa. Several of the genera best represented in Europe have the plesiomorphic pattern of the reproductive system, with gonads in GI–GIII but atria limited to GII, so that two pairs of vasa deferentia are associated with them. This pattern is shared with the Branchiobdellida, and whereas median fusion of the atria has happened independently in the two taxa, these two taxa may arise from a common ancestor despite the degree of specialization to an epizooic habit in the branchiobdellidans. Most subgeneric taxa entered into the analyses are grouped at least close to their congeners, with the exception of Neoscolex, which may be more closely related to the pair of Kincaidiana species than to Styloscolex. Rhynchelmis may have an origin distinct from that of Lamprodrilus, Kincaidiana, and Neoscolex, as the loss of the anterior testes may be convergent and not monophyletic as suggested here.


2015 ◽  
Vol 28 (4) ◽  
pp. 203 ◽  
Author(s):  
Xénia A. Weber ◽  
Alexander N. Schmidt-Lebuhn

The Australasian clade of Gnaphalieae (Asteraceae) is an ecologically diverse group of species whose generic limits and phylogenetic relationships are still partly unresolved. Previous studies including hybridisation trials, morphological characterisation and preliminary phylogenetic analyses within the Gnaphalieae have suggested that two genera, namely, Waitzia and Leucochrysum, require further investigation into their generic boundaries. To explore the phylogenetic relationships of both genera, the present study used a combination of morphological and molecular approaches. The character traits of herbarium specimens from 14 species and six infraspecific taxa of Waitzia, Leucochrysum and Anemocarpa were examined. Chloroplast (psbA–trnH) and nuclear ribosomal ITS and ETS sequences were generated for phylogenetic analysis. Our findings support Waitzia in its current circumscription as a monophyletic group, whereas Leucochrysum was found to be polyphyletic. Leucochrysum fitzgibbonii was found to be the sister group of Waitzia and shares characters of the involucral bracts, indumentum and growth habit with that genus. The species is formally transferred to the genus Waitzia as W. fitzgibbonii.


2005 ◽  
Vol 36 (2) ◽  
pp. 121-143 ◽  
Author(s):  
Pekka Vilkamaa ◽  
Heikki Hippa

AbstractThe phylogeny of the main groups of the Sciaroidea, including the fossil Antefungivoridae, Archizelmiridae, Mesosciophilidae, Pleciofungivoridae, Pleciomimidae, Protopleciidae and Bolitophilidae: Mangasinae, and an extant new taxon, was studied by parsimony analysis. Two cladistic analyses of seventy-eight morphological characters from adults were made. One analysis, with forty-one extant taxa in the ingroup and the other, with the addition of twelve fossil taxa, both produced two most parsimonious cladograms. The phylogenetic hypotheses obtained differed from each other, and in part also to a great extent from previous ones although most of the traditionally recognized groups appeared monophyletic, including the speciose Cecidomyiidae and Sciaridae. The Cecidomyiidae (fossil analysis) or the Keroplatidae-Ditomyiidae (extant analysis) appeared as the sister-group of the rest of the Sciaroidea. Following on from these analyses, we propose emending the current Sciaridae to include the following subfamilies: Archizelmirinae stat. n., Rangomaraminae stat. n., Sciarinae, Sciarosominae subfam. n. and Sciarotrichinae subfam. n. A new taxon from Namibia, Sciarotricha biloba gen. n., sp. n. is described, and, according to the phylogenetic analysis, is placed in the Sciaridae (Sciarotrichinae). The sister-group of the Sciaridae as newly defined is the Mycetophilidae group, in the extant analysis including the Mycetophilidae, Manotidae, Lygistorrhinidae, Pterogymnus and Sciaropota, and in the fossil analysis even including the Mesosciophilidae and the Ohakunea group (Ohakunea + Colonomyia).


Parasitology ◽  
2018 ◽  
Vol 146 (5) ◽  
pp. 596-603 ◽  
Author(s):  
Sergey G. Sokolov ◽  
Dmitry M. Atopkin ◽  
Misako Urabe ◽  
Ilya I. Gordeev

AbstractIn the present paper, the phylogenetic relationships between genera, subfamilies and families of the Hemiuroidea are explored. Twelve new sequences of 28 rDNA and data taken from GenBank (NSBI) on 43 species affiliated to 34 genera were included in the analysis. Most of the hemiuroidean trematodes form two highly supported clades (A and B), which are sister groups to each other.Hemipera manterijoined withGonocercaspp. with moderate statistical support. This clade is basal relative to the clades A and B. Сlade A is polytomic and contains representatives of the families Accacoeliidae, Syncoeliidae, Didymozoidae, Hirudinellidae and Sclerodistomidae, and derogenid subfamilies Derogeninae and Halipeginae. At the same time, the Syncoeliidae, Hirudinellidae and Accacoeliidae form a well-supported monophyletic group. The phylogenetic relationship between Derogeninae and Halipeginae is poorly resolved. Сlade B unites the isoparorchiid, bunocotylid, lecithasterid and hemiurid trematodes. Our data re-establishes the family Bunocotylidae, which consists of two subfamilies, Opisthadeninae and Bunocotylinae, and theMachidatrema chilostoma+Hysterolecithoides frontilatusgroup. The Bunocotylidae is the sister group to the Hemiuridae + Lecithasteridae group and the Isoparorchiidae is a basal relative to the representatives of these three hemiuroid families.


Zootaxa ◽  
2008 ◽  
Vol 1686 (1) ◽  
pp. 1 ◽  
Author(s):  
KEVIN W CONWAY ◽  
WEI-JEN CHEN ◽  
RICHARD L MAYDEN

The osteology of the miniature cyprinid Celestichthys margaritatus Roberts, type species of Celestichthys Roberts, is described in detail and briefly compared with that of other members of the Rasborinae (notably Danio (s.s), Danionella, Devario, Esomus, Microrasbora, Paedocypris and Sundadanio). Celestichthys margaritatus possesses an “A” stripe on the anal fin and two pigment stripes on the caudal fin (apomorphic features of Danio sensu Fang, 2003). In addition, C. margaritatus exhibits a median projection on the outer arm of the os suspensorium, a derived feature, present only in species of Danio (including D. erythromicron) amongst the Cyprinidae, and a lateral projection on the lateral face of the dentary (present only in Danio, Sundadanio and Paedocypris). Phylogenetic analysis of 1,494 bp of the RAG1 nuclear gene for 31 rasborine taxa, including 5 species of Danio, places C. margaritatus as the sister group to D. erythromicron, and part of a larger monophyletic group including all other species of Danio included for analysis. Based on characters of morphology and its position in a molecular phylogeny of the Rasborinae it is proposed that Celestichthys be placed in the synonymy of Danio, its only member referred to as Danio margaritatus new combination.


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