scholarly journals The pasture ryegrass plant, what is it?

Author(s):  
J.L. Brock ◽  
V.J. Thomas

Detailed studies of plant growth processes are important in understanding the performance and persistence of species in pastures, particularly in response to uncharacteristic environmental stress. The morphology of perennial ryegrass plants in mixed sheep grazed pastures was determined in self contained farmlets undercontrasting managements of rotational grazing, set stocking or a combination of both. Average size was 90 mg total DW, with 4- 5 tillers and 12-13 leaves, little different to white clover. Ryegrass exhibits strong clonal growth, with extension at the apex and death of the basal stem releasing branches to form new plants at regular intervals, maintaining a stable population structure of small plants all year. The normally short internodes on ryegrass stems can elongate to form stolon and elevate the apex to a more favourable position if survival is threatened. On average only 2530% of plants contained stolon at any one time. Because of high plant density the quantities of ryegrass stem present was often in excess of that produced by white clover in the same swards. Grazing management did not affect plant structure (numbers of tillers, leaves etc) only their size (dry weight), but had marked effects on pasture structure and subsequent survival of plants under stress. Keywords perennial ryegrass, plant morphology, grazing management, seasonal variation, plant survival, stolon formation

1993 ◽  
Vol 120 (3) ◽  
pp. 301-310 ◽  
Author(s):  
J. L. Brock ◽  
R. H. Fletcher

SUMMARYThe morphology of independent plants of perennial ryegrass (Lolium perenne L.) in mixed swards under intensive sheep grazing was studied at Palmerston North, New Zealand in 1988/89. Intact plants of two cultivars, ‘Grasslands Nui’ and ‘Grasslands Ariki’, were sampled from pastures under rotational grazing, set stocking, and a combination of both systems, every 2 months for 16 months, by taking turves (250 × 250 mm) and washing out the plants. Characters measured for each plant were: number of flower heads, leaves and tillers; basal stem and internode stolon length; flower head, leaf blade, leaf sheath and stolon dry weight (DW). Additional pasture data on tiller density and dispersion and herbage DW to ground level were also collected.Perennial rye grass exhibited a strong clonal growth pattern similar to that of white clover (Trifolium repens L.) but with a better balance between growth at the apex and death of old basal stem. Release of branch stolons to form new plants was regular, which maintained population stability. Formation of internode (I) stolon appeared to be a response of heavily shaded tillers for survival by elevation of the growing point to a more favourable position. The total DW of ryegrass stolon in pastures was greater than that of the companion white clover.There was little seasonal variation in number of tillers or basal (B) stem length per plant. During the reproductive period (late spring/early summer) plants were heavier and had more leaves and internode stolon than at other times of the year. Grazing management had little effect on plant structure, but plants under rotational grazing had 75% greater DW, less basal stem and reduced probability of internode stolon formation than plants under set stocking. Nui plants had heavier but fewer tillers than Ariki did, resulting in little difference in plant DW. Ariki had fewer single-tiller plants and a higher proportion of multiple-tiller (> 7) plants than Nui had. The structure of the populations was not affected by season, management or cultivar and remained stable throughout the year, with a log-normal distribution of plant size. Because of the uniformity of plant structure, plant density in pastures paralleled tiller density, averaging 1900, 3360 and 2330 plants/m2 for rotationally grazed, set stocked and combined grazing treatments respectively.In view of the strong influence that grazing management can have on ryegrass performance and persistence in pastures, the lack of response at the level of plant and population structure to grazing management suggests that some other level of organization within the ecology of the pasture is of greater importance.


1997 ◽  
Vol 129 (1) ◽  
pp. 19-31 ◽  
Author(s):  
D. E. HUME ◽  
J. L. BROCK

The effects of contrasting management systems either of infrequent rotational grazing by town milk supply dairy cattle, or of frequent defoliation by continuously grazing sheep and beef cattle, on the morphology of independent plants and populations of ‘Grasslands Roa’ tall fescue in mixed pastures, were measured over 1 year (1992/93) in New Zealand. Volunteer perennial ryegrass plants were also measured for comparison.While both species exhibited a similar pattern of clonal growth, tall fescue developed more plants of higher branching complexity than perennial ryegrass, chiefly through maintaining more connective stems, as herbage production was confined to the three youngest branching orders in both species. Greater resistance to microbial degradation of old stems through poorer quality organic matter (wide C[ratio ]N ratio) compared to perennial ryegrass may be responsible for the greater complexity of fescue plants. In addition, tillering rates in tall fescue were three times lower which was offset by greater longevity and size of leaves and tillers, compared to perennial ryegrass. As a result, seasonal fluctuation in the distribution of plants among the various branching orders in tall fescue was small, producing a more stable population relative to the distinct seasonal changes in the population of perennial ryegrass plants. Grazing management had no effect on the seasonal population structure in either species.Differences in plant structure due to grazing management were small, with only slightly more tillers on sheep-grazed than on cattle-grazed tall fescue plants. The major effect of grazing management was on dry weight or size of plant components. Cattle-grazed tall fescue plants were 120% heavier, with greater numbers and lengths of stolons and flower heads than those under sheep grazing. For the volunteer perennial ryegrass, the difference was only 65%, possibly due to competition from the more vigorous tall fescue under rotational cattle grazing.Both species produced stolons throughout the year, although these were primarily associated with reproductive growth in spring. In tall fescue, an additional distinction was made between stolon and rhizome, the latter occurring mainly in the summer–autumn. Their possible functions in plant growth are discussed.


Author(s):  
S.L. Harris

Seasonal changes in white clover plant morphology were investigated in Waikato dairy pasture. Plants were classified according to the degree of branching, 2nd-order plants comprising the bulk of the population (47%) throughout the year. There was a slightly greater proportion of lst-order plants than found in similar studies on sheep-grazed swards in Manawahr. Plant fragmentation occurred in late spring, increasing the proportion of lstorder plants. A corresponding decrease in plant size was observed, with minimum values measured for most characteristics, i.e., stolon, leaf and root dry weight (DW); stolon length, and number of leaves, roots and growing points. Overall, white clover plants in Waikato dairy pastures are considerably larger than plants in Manawatu sheep pastures. The trial highlighted that because white clover plants in spring are small, they are vulnerable to mismanagement in dairy pastures, just as has been demonstrated in pasture grazed by sheep. There were, however, some significant differences in white clover plant structure in dairy pastures compared with sheep-grazed swards. Keywords: dairy pasture, morphology, plant order, plant size, seasonal variation, Trifolium repens


Author(s):  
J.L. Brock ◽  
Moon Chul Kim

Field observations indicate that conditions at the stolen/soil surface interface are important for the survival of white clover (Trifolium repens) during drought. Mini-swards of clonal material of two contrasting white clover cultivars (Grasslands Tahora, small-leaved, densely branched; Dusi, large-leaved, tap-rooted) were established in boxes (300 x 420 x 290 mm deep) for 6 months, then subjected to a 6-week drought. Stolen/soil surface conditions were modified to have stolons either, 1) fully exposed, 2) shaded, or 3) buried. Numbers and, characteristics of surviving plants were measured. The major effect was on the number of plants surviving. More plants survived under shading (55%) than burial (25%) and full exposure (17%). Numbers of plants surviving drought was related to pre-drought plant density, hence Tahora had higher plant survival than Dusi, particularly when fully exposed. Although initially Dusi plants were 25 times heavier, with 5 times the number of stolons and roots of Tahora, surviving plants of both cultivars were small and similar in dry weight, although Tahora had higher numbers of growing points and roots thanDusi. The tap-rooted character of Dusi did not. confer any advantage during drought. It was concluded that the main factor influencing drought survival of white clover appears to be grazing management. Systems that can produce dense white clover populations and maintain high pasture density (grass) capable of providing shelter from direct solar radiation during drought stress, are superior. Keywords: drought survival, plant density, plant morphology, stolon burial, stolon shading, Trifolium repens


2003 ◽  
Vol 9 ◽  
pp. 25-40
Author(s):  
A.M. Nicol ◽  
T.N. Barry

Pasture is the primary feed source for NZ deer production with the greatest proportion grazed in situ. The quantity and quality of the pastures available to grazing deer varies markedly as a result of environmental factors and feed planning decisions by deer farmers. The grazing behaviour of deer responds to changes in pasture height and mass and the effect of pasture variables (height, pre -and post-grazing pasture mass and pasture allowance) on deer productivity are presented. These show that maximum levels of deer production from pasture will be achieved at a pasture height of around 8 cm (continuously stocked or post-grazing) although there is some evidence that for large genotypes, higher pasture availability is required. The relationship of liveweight gain of young deer with pasture availability shows marked seasonal effects. At the same level of pasture availability liveweight gain in spring is about twice that in winter, with autumn and summer intermediate. Increasing pasture availability cannot compensate for seasonal differences in liveweight gain. Furthermore, liveweight gain increases at a greater rate in spring than winter to increasing pasture availability, thus it is more important that appropriate pasture allowances are provided in spring than in winter. There is more variability in liveweight gain at a similar pasture availability in summer than in other seasons because of the greater variation in pasture quality in summer with the potential accumulation of seedheads and dead material. Alternative forage species are used in deer production for times of the year when quantity and quality of perennial ryegrass-based pastures limit productivity. Relative to weaner red deer grazed on perennial ryegrass/ white clover pasture, grazing on pure swards of red clover or chicory increased growth during autumn by 26-47% and during spring by 10-14%. The proportion of stags attaining target slaughter liveweight at 12 months of age increased from 75 to 94%. Pre-weaning growth during lactation was increased by approximately 20%. Red clover and chicory produce a greater proportion of their total DM during late summer and autumn than does perennial ryegrass/white clover pasture, and are therefore better aligned with deer feed requirements, particularly those of lactating hinds. Grazing on sulla in autumn and spring increased the growth of weaner deer by 33 and 10%, relative to pasture fed deer. Indoor studies showed that relative to perennial ryegrass, chicory was of higher organic matter digestibility, disintegrated more rapidly in the rumen with a low rumination time and had shorter mean retention time of material in the rumen. This explains differences in voluntary feed intake which were 56, 26 and 15% higher for deer grazing chicory than perennial ryegrass/white clover pastures during summer, autumn and spring respectively. Similar r esults have been found for the digestion of red clover versus perennial ryegrass by red deer. Plant density in stands of both chicory and red clover declines with time, with their lifetime under deer grazing being approximately 4 years. A mixture of both plants offers a food option as specialist forage for increasing deer growth and also fixing nitrogen. To ensure good persistence such forages should not be grazed in periods of prolonged wet weather. Best persistence is obtained when these are managed as specialist forages for increasing deer growth on a small area of the farm, (10- 20% total area), rather than being sown as a mixture with grasses over large areas of the farm. There is no specific comparison of deer production under different stocking systems and both continuous grazing and rotational grazing are used. Based on evidence and practices with other species, choice of stocking system has more to do with pasture/forage species, feed budgeting, pasture management and animal behaviour than with productivity. At high stocking densities (150 deer/ha), the grazing time of subordinate animals is reduced. Where possible, without inducing undue stress (e.g. at weaning), young deer should be grouped for grazing by liveweight. Deer production systems have a seasonal pattern of energy demand that does not match that of pasture growth in most NZ environments. This calls for manipulation of the feed demand by integration of livestock systems and/ or modification of the feed supply through conservation and supplementation. The most commonly used supplements are pasture and lucerne silage/baleage and grain. The quality (ME/kg DM) has a significant impact on the resulting liveweight gain.


1988 ◽  
Vol 111 (2) ◽  
pp. 273-283 ◽  
Author(s):  
J. L. Brock ◽  
M. J. M. Hay ◽  
V. J. Thomas ◽  
J. R. Sedcole

SummaryThere has been little study on the growth and morphology of individual plants constituting the population of white clover in mixed swards under grazing. Such information is required if the mechanisms governing plant productivity and persistence are to be understood.Intact white clover plants were sampled from intensively sheep-grazed pastures under set stocking, rotational grazing, and a combination of both systems, by taking turves (250 × 250 mm), and washing out the plants, every month for a year. Characters measured for every stolon of each plant were: presence of a growing point; numbers of leaves, roots and axillary buds; stolon length. Total plant leaf and stolon dry weight were also recorded. Plants were classified according to degree of branching, and the contribution of each branching order to the population determined.There were strong seasonal variations in plant size (leaf and stolon dry weight, stolon length, and numbers of stolons and leaves per plant) which showed a significant decrease in spring with recovery over the following summer. This was paralleled by a rapid increase in the proportion of less branched plants (1st and 2nd branching order) in the population from 60 to 80% in spring, as higher-order plants broke up into smaller- and lower-ordered plants at this time. Numbers of roots per plant increased over winter to peak in early spring then declined in the following summer-autumn. While system of grazing management had no significant effect on branching structure of plants, it had a large effect on plant dry weight; rotationally grazed plants were 2·5 times larger than set stocked plants (0·182 cf. 0·073 g respectively).Other general features of plant morphology were that each successive order of branch stolons was shorter and length before branching was less than that of their preceding parent stolon. The highest branching order observed was 6th order. There was no relationship between branching and numbers of roots; in branched plants only 55% of stolons were rooted regardless of plant order, but rooted stolons accounted for 85% of total stolon length and carried 62, 48 and 90% of the leaves, growing points and axillary buds per plant, respectively.Comparison with other studies suggests that the processes outlined in this report may be common to white clover growth under grazing over a wide range of favourable environments.


Author(s):  
R.J. Townsend ◽  
J.E. Radcliffe

Lambs were rotationally grazed with ewes in a 'sheep only' system and with low, medium, and high proportions,of wether goats from October to March on dryland perennial ryegrass-white clover pastures in Canterbury. Lambs grazed with ewes and goats until they were weaned; then they grazed ahead of other stock. Treatment ratios of adult stock on a per head basis were 3 ewes:1 goat ('low goats'), 3 ewes:2 goats ('medium goats') and 3 ewes:3 goats ('high goats'). The ewe:lamb ratio was 1:1.35 on all treatments. Lamb growth rates (g/day) were 152 for 'sheep only', and 169, 185 and 203 for low, medium and high goat treatments respectively. As each treatment lamb reached a target draft weight, it was replaced by a lighter lamb for finishing. In comparison with the 'sheep only' treatment, an extra 1.6 lambs/ha were drafted off the 'medium goat' and an additional 2.6-lambs/ha off the 'high goat' treatment by the end of January. Increasing the proportion of goats:ewes increased the supply of white clover in pastures especially after lambs were weaned. Examination of faecal plant material showed 30% clover in the diet of lambs in a 'sheep only' system, increasing as the goat:sheep ratio increased to 60% clover in the 'high goat' treatment. Keywords white clover, perennial ryegrass, goats, sheep,grazing management, diet, lamb


HortScience ◽  
1995 ◽  
Vol 30 (2) ◽  
pp. 238-241 ◽  
Author(s):  
Pat Bowen ◽  
Stan Freyman

Raspberries (Rubus idaeus L.) were grown for 5 years with three floor management treatments: clean cultivation (CC) and ground covers of white clover (Trifolium repens L.) (WC) and perennial ryegrass (Lolium perenne L.) (PR). Primocane growth was strongest with WC and weakest with PR. Fruit production did not differ between WC and CC treatments and was higher than with PR. On 22 Aug. and 10 Sept. in the last year, primocanes grown with WC were taller, had more nodes and a higher dry weight, contained more N, and had retained more leaves than those grown with PR. Net CO2 assimilation per unit leaf area was higher with WC than with PR, and the difference was greater at the more proximal position. The estimated net CO2 assimilation rate per primocane was more than twice as high with WC than with PR.


Author(s):  
C.J. Korte

The patterns of herbage accumulation and the tiller dynamics of a "Grasslands Nui" perennial ryegrass dominant pasture were compared under two grazing intensity treatments in late spring. Rank stemmy herbage developed with lax grazing whereas dense leafy pasture developed with close grazing. Dead herbage and uneaten stems in rank pastures shaded photosynthetic tissue. Leafy swards had a higher herbage accumulation rate and a higher tiller density than stemmy swards. Methods of achieving leafy pasture on farms by combinations of hard grazing, forage conservation and mechanical topping are discussed briefly.


Author(s):  
G.P. Cosgrove ◽  
R.W. Brougham

Grazing management influences on the production and species composition of a ryegrass, red and white clover pasture were evaluated over 3 years. Pastures were grazed by cattle frequently in either winter or summer and infrequently for the remainder of the year or grazed infrequently all year. Frequency was determined by the height of herbage pre-grazing and was lo-12 cm for frequent grazing and 15-20 cm for infrequent grazing. Annual herbage production of 15.1 t DM/ha was not influenced by grazing management but seasonal production was. Management caused both instantaneous and longer term effects on production of component species. Lower summer production because of frequent grazing in that season resulted from a reduction in red clover production from 2.75 to 1.3 t DM/ha. However, ryegrass production in spring was increased by 60% and in summer it was more than doubled by this grazing treatment. Frequent grazing in winter also increased ryegrass and reduced red clover production in summer compared with infrequent grazing all year but had no effect on ryegrass production in spring. Red clover plant density was unaffected by the different grazing managements imposed for two and a half years but subsequently was lower under frequent summer grazing. Red clover dominance in summer and its effect on ryegrass production in spring can be controlled by more frequent grazing in summer and a more even species balance results. Emphasis on spring or summer production can be achieved through grazing management without affecting total production. Keywords: Red clover, ryegrass, grazing frequency, cattle grazing, herbage production, seasonal production


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