New Early Eocene brown lacewings (Neuroptera: Hemerobiidae) from western North America

2003 ◽  
Vol 135 (5) ◽  
pp. 637-653 ◽  
Author(s):  
Vladimir N. Makarkin ◽  
S. Bruce Archibald ◽  
John D. Oswald
Keyword(s):  
British Columbia ◽  
North America ◽  
Sister Group ◽  
Systematic Position ◽  
Early Eocene ◽  
Western North America ◽  
Fossil Species ◽  
Incertae Sedis ◽  
Extant Genus ◽  
Poorly Differentiated

AbstractTwo Early Eocene hemerobiid species from the Okanagan Highland deposits of western North America are described: Wesmaelius mathewesisp. nov. (Quilchena, British Columbia, Canada) and Cretomerobius wehrisp. nov. (Republic, Washington, United States of America.). A poorly preserved specimen, possibly Hemerobiidae (Driftwood Canyon, British Columbia, Canada) is discussed. The systematic position of the genus Cretomerobius within the Hemerobiidae is unclear, but it appears to belong within (or possibly as the sister group to) the poorly differentiated Drepanacrinae–Megalominae–Drepanepteryginae grade of subfamilies. Wesmaelius mathewesi is the first fossil species to be assigned, although with some uncertainty, to this otherwise speciose and widely distributed extant genus. It also represents the oldest known material of the subfamily Hemerobiinae. Meso hemerobius jeholensis Ping is removed from the Hemerobiidae and treated as Neuroptera incertae sedis.

New Protosmylinae (Neuroptera: Osmylidae) from the early Eocene of western North America, with taxonomic remarks

Zootaxa ◽  
2021 ◽  
Vol 4980 (1) ◽  
pp. 142-156
Author(s):  
VLADIMIR N. MAKARKIN ◽  
S. BRUCE ARCHIBALD ◽  
ROLF W. MATHEWES
Keyword(s):  
North America ◽  
Hind Wing ◽  
Late Eocene ◽  
Early Eocene ◽  
Western North America ◽  
Green River ◽  
Incertae Sedis ◽  
Three New Species ◽  
Okanagan Highlands ◽  
Provincial Park

The osmylid subfamily Protosmylinae is revised based on our emended diagnosis: Petrushevskia Martynova, 1958 and Mesosmylidus Jepson et al., 2012 are excluded (both considered Osmylidae incertae sedis), and Sogjuta Martynova, 1958 is transferred to it from the Mesosmylininae. The late Eocene genus Protosmylus Krüger, 1913 is considered a junior synonym of Osmylidia Cockerell, 1908 based on a distinct apomorphy (deeply forked MA in the hind wing), syn. nov. Three new species of Osmylidia from the early Eocene of North America are described: O. donnae sp. nov. from Quilchena, O. glastrai sp. nov. from Republic, Washington, USA, and an unnamed species of Osmylidia is reported from Driftwood Canyon Provincial Park, British Columbia (all localities of the Okanagan Highlands series), and O. taliae sp. nov. from the Green River Formation of Colorado, USA. 

Early Eocene (Ypresian) birds from the Okanagan Highlands, British Columbia (Canada) and Washington State (USA)

2019 ◽  
Vol 56 (8) ◽  
pp. 803-813
Author(s):  
Gerald Mayr ◽  
S. Bruce Archibald ◽  
Gary W. Kaiser ◽  
Rolf W. Mathewes
Keyword(s):  
British Columbia ◽  
North America ◽  
Fossil Record ◽  
Washington State ◽  
Fossil Birds ◽  
Early Eocene ◽  
Western North America ◽  
Postcranial Skeleton ◽  
The North ◽  
Okanagan Highlands

We survey the known avian fossils from Ypresian (early Eocene) fossil sites of the North American Okanagan Highlands, mainly in British Columbia (Canada). All specimens represent taxa that were previously unknown from the Eocene of far-western North America. Wings from the McAbee site are tentatively referred to the Gaviiformes and would constitute the earliest fossil record of this group of birds. A postcranial skeleton from Driftwood Canyon is tentatively assigned to the Songziidae, a taxon originally established for fossils from the Ypresian of China. Two skeletons from Driftwood Canyon and the McAbee site are tentatively referred to Coliiformes and Zygodactylidae, respectively, whereas three further fossils from McAbee, Blakeburn, and Republic (Washington, USA) are too poorly preserved for even a tentative assignment. The specimens from the Okanagan Highlands inhabited relatively high paleoaltitudes with microthermal climates (except Quilchena: lower mesothermal) and mild winters, whereas most other Ypresian fossil birds are from much warmer lowland paleoenvironments with upper mesothermal to megathermal climates. The putative occurrence of a gaviiform bird is particularly noteworthy because diving birds are unknown from other lacustrine Ypresian fossil sites of the Northern Hemisphere. The bones of the putative zygodactylid show a sulphurous colouration, and we hypothesize that this highly unusual preservation may be due to the metabolic activity of sulphide-oxidizing bacteria.

Early Eocene snakeflies (Raphidioptera) of western North America from the Okanagan Highlands and Green River Formation

Zootaxa ◽  
2021 ◽  
Vol 4951 (1) ◽  
pp. 41-79
Author(s):  
S. BRUCE ARCHIBALD ◽  
VLADIMIR N. MAKARKIN
Keyword(s):  
United States ◽  
British Columbia ◽  
North America ◽  
United States Of America ◽  
Green River Formation ◽  
Early Eocene ◽  
Western North America ◽  
Green River ◽  
The Family ◽  
Okanagan Highlands

Four new species of Raphidiidae are described from the early Eocene of western North America: Megaraphidia antiquissima sp. nov. from McAbee, M. ootsa sp. nov. from Driftwood Canyon, M. hopkinsi sp. nov. from the Allenby Formation (all from British Columbia, Canada), M. klondika sp. nov. from Republic (Washington, United States of America). Archiinocellia Handlirsch, 1910, Archiinocellia oligoneura Handlirsch, 1910 from Horsefly River (British Columbia, Canada), and A. protomaculata (Engel, 2011), comb. nov., from the Green River Formation (Colorado, United States of America) are redescribed. Archiinocellia is assigned to Raphidiidae, sit. nov. The apparent absence of sclerotized gonocoxites 9 in the Archiinocellia protomaculata male is probably plesiomorphic at the family level. As some modern snakeflies do not require a cold interval to complete their development and Eocene Inocelliidae and Raphidiidae lived in regions of warm winters (especially A. protomaculata), adaptation to cold winters in many modern snakeflies is a post-Eocene phenomenon. Eocene Raphidiidae of Europe (Priabonian) differ greatly from those of North America (Ypresian and Priabonian). This pattern might reflect dispersal in either direction or ranges established prior to continental separation. Eocene Inocelliidae of Europe (Priabonian), however, are more similar to those of North America (Ypresian and Lutetian). 

New species of latest Eocene/earliest Oligocene microgastropods (Heterobranchia: Orbitestellidae and Omalogyridae) from the Gries Ranch Formation, Lewis County, Washington State, USA

Zootaxa ◽  
2021 ◽  
Vol 4981 (3) ◽  
pp. 469-480
Author(s):  
ALEXEI V. CHERNYSHEV ◽  
JAMES L. GOEDERT
Keyword(s):  
New Species ◽  
North America ◽  
Fossil Record ◽  
Washington State ◽  
Shell Morphology ◽  
Early Eocene ◽  
Western North America ◽  
Crescent Formation ◽  
Fossil Species ◽  
Latest Eocene

Continued sampling of the latest Eocene to earliest Oligocene Gries Ranch Formation in Lewis County, Washington State, has yielded new heterobranch microgastropod species. Orbitestella kieli sp. nov., is the third fossil species of this genus and family Orbitestellidae from western North America. Two new species of Ammonicera, A. rolani sp. nov. and A. danieli sp. nov., are together only the second fossil record of this genus and the family Omalogyridae from the northeastern Pacific Ocean. New specimens of two previously recorded species, O. palaiopacifica Squires & Goedert and A. benhami Squires & Goedert, from early Eocene rocks of the Crescent Formation provide new data regarding shell morphology. The fossil record of both Ammonicera and Orbitestella in western North America is restricted to early Eocene to earliest Oligocene age rocks in Washington State. 

A diverse new assemblage of green lacewings (Insecta, Neuroptera, Chrysopidae) from the early Eocene Okanagan Highlands, western North America

2013 ◽  
Vol 87 (1) ◽  
pp. 123-146 ◽  
Author(s):  
Vladimir N. Makarkin ◽  
S. Bruce Archibald
Keyword(s):  
North America ◽  
Fossil Record ◽  
Large Species ◽  
Early Eocene ◽  
Western North America ◽  
Fossil Species ◽  
Green Lacewings ◽  
The Family ◽  
Okanagan Highlands ◽  
First Time

The early Eocene green lacewings (Neuroptera, Chrysopidae) of the Okanagan Highlands deposits of McAbee, and Driftwood Canyon, British Columbia (Canada) and Republic, Washington (U.S.A.) are treated in detail for the first time. At least six genera are present, one unnamed, three new, with at least 10 new species, six named:Protochrysa fuscobasalisn. sp. (McAbee) (Limaiinae, the youngest known record of the subfamily),Okanaganochrysa coltsunaen. gen. n. sp. (McAbee),Adamsochrysa asperan. gen. n. sp. (McAbee),A. wilsonin. gen. n. sp. (Republic),Archaeochrysa profractan. sp. (McAbee), andPseudochrysopa harveyin. gen. n. sp. (Driftwood Canyon) (all Nothochrysinae, the latter provisionally). The four unnamed species include one assigned toPseudochrysopa, two likely belonging toAdamsochrysa, and one of an unknown nothochrysine genus. Microtholi are detected on the abdominal sclerites ofAdamsochrysa wilsoni, and the spermatheca and spermathecal duct in the abdomen ofPseudochrysopa harveyi, the first reported occurrences of these preserved in fossil Chrysopidae. Structures were detected on the apical wing margins of some species that appear similar to trichosors, which are unknown in Chrysopidae, but are present in some other neuropteran families. This is the richest described assemblage of the family anywhere in the fossil record. Okanagan chrysopids were also morphologically and presumably ecologically diverse, including large species with rich venation and well as those with simplified venation and the smallest known fossil species. This is the oldest reported occurrence of the family in North America.

Review of Eohiodon (Teleostei: Osteoglossomorpha) from western North America, with a phylogenetic reassessment of Hiodontidae

1997 ◽  
Vol 71 (6) ◽  
pp. 1109-1124 ◽  
Author(s):  
Li Guo-Qing ◽  
Mark V. H. Wilson ◽  
Lance Grande
Keyword(s):  
North America ◽  
Sister Group ◽  
Sensu Stricto ◽  
Valid Species ◽  
Western North America ◽  
Extant Species ◽  
The Family ◽  
Fossil Records ◽  
Fossil Genus ◽  
Freshwater Teleosts

Review of recently collected material of Eohiodon from North America suggests that there are two valid species, E. rosei (Hussakof) and E. woodroffi Wilson. Eohiodon falcatus Grande is identical to E. woodruffi in known skeletal features and nearly all meristic features and is treated as a junior synonym of the latter. The fossil genus Eohiodon Cavender differs from Hiodon Lesueur, which is known from both fossil and extant species, in numerous meristic and osteological features. The caudal skeleton in Eohiodon is nearly identical to that in Hiodon.The traditionally accepted Notopteroidei, containing Lycopteridae, Hiodontidae, and Notopteridae, is a polypheletic group. The Asian fossil family Lycopteridae is not more closely related to Hiodontidae than it is to other taxa in the Osteoglossomorpha, but is sister to all other Osteoglossomorpha. The Hiodontiformes sensu stricto, including only the family Hiodontidae, is the sister-group of the Osteoglossiformes. This family is not more closely related to notopterids than to other taxa in Osteoglossiformes. The Notopteridae are most closely related to the Mormyroidea; together they and the fossil family Ostariostomidae constitute the sister-group of the Osteoglossoidei.Fossil records of Hiodontiformes sensu stricto and Notopteroidei indicate a widespread pre-Neogene biogeographic range of these freshwater teleosts, suggesting that extinction must have been involved in the Cenozoic evolution of these two osteoglossomorph sublineages.

Japewia Subaurifera, A New Lichen Genus and Species From North-West Europe and Western North America

1990 ◽  
Vol 22 (3) ◽  
pp. 205-212 ◽  
Author(s):  
T. Tønsberg
Keyword(s):  
British Columbia ◽  
North America ◽  
Western North America ◽  
Coniferous Trees ◽  
North West

AbstractThe genus Japewia Tønsb. is introduced to accommodate three species including J. subaurifera Muhr & Tønsb. sp. nov. based on material from Norway, Sweden, Finland, Scotland, Canada (British Columbia) and U.S.A. (Washington). This species is closely related to Lecidea tornoensis Nyl. but is distinguished in being sorediate and by the production of lobaric acid (accessory) and acetone-soluble pigments. It grows on bark of deciduous and coniferous trees. Lecidea carrollii Coppins & P. James and L. tornoensis Nyl. are transferred to Japewia.

The Lower Jurassic phosphorites of southeastern British Columbia and terrane accretion to western North America

1989 ◽  
Vol 26 (8) ◽  
pp. 1612-1616 ◽  
Author(s):  
T. P. Poulton ◽  
J. D. Aitken
Keyword(s):  
British Columbia ◽  
North America ◽  
Oceanic Crust ◽  
West Coast ◽  
Cold Water ◽  
Lower Jurassic ◽  
Depositional Model ◽  
Western North America ◽  
The West ◽  
Terrane Accretion

Sinemurian phosphorites in southeastern British Columbia and southwestern Alberta conform with the "West Coast type" phosphorite depositional model. The model indicates that they were deposited on or near the Early Jurassic western cratonic margin, next to a sea or trough from which cold water upwelled. This suggests that the allochthonous terrane Quesnellia lay well offshore in Sinemurian time. The sea separating Quesnellia from North America was partly floored by oceanic crust ("Eastern Terrane") and partly by a thick sequence of rifted, continental terrace wedge rocks comprising the Purcell Supergroup and overlying Paleozoic sequence. This sequence must have been depressed sufficiently that access of upwelling deep currents to the phosphorite depositional area was not impeded.

NEW NEARCTIC CRANE-FLIES (TIPULIDAE, DIPTERA), PART XIX

1943 ◽  
Vol 75 (8) ◽  
pp. 139-145 ◽  
Author(s):  
Charles P. Alexander
Keyword(s):  
British Columbia ◽  
North America ◽  
Type Material ◽  
Western North America ◽  
Crane Flies ◽  
The Individual

The crane-flies considered herewith are all from Western North America, from British Columbia to California. The names of the collectors and the location of the type material are indicated at the end of the individual specific accounts; where not stated to the contrary, such types are preserved in my own collection of these flies.

A New Gunnel, Pholis clemensi, from the Coast of Western North America

1964 ◽  
Vol 21 (5) ◽  
pp. 933-939 ◽  
Author(s):  
Richard H. Rosenblatt
Keyword(s):  
British Columbia ◽  
New Species ◽  
North America ◽  
North American ◽  
North American Species ◽  
Western North America ◽  
The North ◽  
The Family ◽  
Juan De Fuca ◽  
A New Species

A new species, Pholis clemensi, referred to the family Pholidae, is named and described from 12 specimens taken in southern British Columbia waters and the Strait of Juan de Fuca. Pholis clemensi is compared with other members of the genus, and a key is given to the North American species.

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