Llandovery secuicollactinae and rotasphaeridae (radiolaria) from the Cape Phillips Formation, Cornwallis Island, Arctic Canada
Radiolaria extracted from two bedded-limestone and two limestone-concretion samples from the upper Llandovery of the Cape Phillips Formation, Cornwallis Island, allow the distinction between the spicular haplentactinid subfamily Secuicollactinae and the nonspicular Rotasphaeridae. This restores Secuicollactinae and returns Secuicollacta to the subfamily from the Rotasphaeridae. Secuicollactinae represent 36 percent of the fauna and are assigned to three genera, Secuicollacta, Diparvapila new genus, and Parvalanapila new genus; these form 27, five, and three percent of the fauna, respectively. The remaining one percent of the subfamily is unassigned. Rotasphaeridae form four percent of the fauna and are represented by Rotasphaera. Secuicollacta is diagnosed on the basis of the presence of an ectopic spicule in addition to several rotasphaeridlike primary units. The salient characters of Diparvapila include two lattice shells with an ectopic spicule on the medullary shell. Parvalanapila is diagnosed on the basis of a lattice sphere overlain by a labyrinthine layer and an ectopic spicule on the lattice sphere. New species described here are Secuicollacta malevola, S. sceptri, S. gliris, S. herrimani, S. segari, S. vallipuellae, S. resodiosae, S. glaebosa, S. teli, Diparvapila hicocki, D. saintrochae, D. larseni, Parvalanapila fleischerorum, and Rotasphaera severa. Four species of Secuicollacta are reported in open nomenclature, as are two of Diparvapila and two of Rotasphaera.