Evolutionary tempo and mode in a sequence of the Upper Cretaceous bivalve Pleuriocardia

Paleobiology ◽  
1987 ◽  
Vol 13 (2) ◽  
pp. 140-151 ◽  
Author(s):  
Dana H. Geary

This study addresses the question of evolutionary tempo and mode in a sequence of Upper Cretaceous bivalves in the genus Pleuriocardia from the Western Interior Basin of North America. Change between species was probably phyletic (without persistence of ancestors). There is some evidence for weak gradual change within the lineage, but most important change is concentrated in short intervals of time. Detailed examination of the differences among samples reveals pronounced geographic variation, whereas temporal variation within localities is generally minor. The relatively rapid episodes of change fit the model of punctuated equilibrium, but the phyletic nature of species-level change does not.The value of the debate about the punctuated and gradualistic models has been to force a more criticial examination of fossil sequences, but these sequences should not be forced into narrowly defined categories. A variety of evolutionary patterns may exist in the fossil record, and it is this variation in pattern that will inform us of the underlying processes.

1987 ◽  
Vol 65 (5) ◽  
pp. 1053-1060 ◽  
Author(s):  
Philip D. Gingerich

Mammals have an unusually good Cenozoic fossil record providing evidence of their evolutionary diversification. We view this record in hindsight, which biases our perception in many ways. Overall worldwide diversity appears to increase exponentially through time, while intensive sampling in local areas indicates that modern levels of diversity were achieved early in the Cenozoic. The evident significance of Pleistocene extinctions depends critically on how extinction rates are quantified. Our taxonomic hierarchy probably reflects the number of major faunal turnovers a group has survived rather than declining intensity of successive turnovers. Morphological innovation and taxonomic diversification appear following intervals of climatic cooling, suggesting that major features of evolution are extrinsically controlled. Favorable stratigraphic settings yield detailed records of gradual anagenesis and cladogenesis in mammals, with intermediates present as evidence of transition. The apparent dichotomy between high evolutionary rates measured by neontologists over short intervals of time and low evolutionary rates measured by paleontologists over long intervals of time disappears when rates are measured on intermediate scales of time. Microevolution and macroevolution are manifestations of common underlying processes expressed on different time scales.


Author(s):  
Max Wisshak

Dendritic and/or rosetted microborings in calcareous and osteic skeletal substrates have a diverse trace fossil record, spanning most of the Phanerozoic, whereas the ichnodiversity of comparable bioerosion traces produced in modern seas is rather limited. The most prominent occurrences are known from Devonian brachiopods and from Upper Cretaceous belemnite rostra. Ichnotaxonomically, they are comprised within one of the few ichnofamilies established to date, the Dendrinidae Bromley et al., 2007. As an outcome of the present revision of this ichnofamily, the plethora of 84 ichnospecies established within 25 ichnogenera since the erection of the type ichnogenus Dendrina Quenstedt, 1849 was considerably condensed to 22 ichnospecies included in 7 ichnogenera, based on a coherent morphological categorisation and ichnotaxobasis assessment. The suite of ichnogenera now subsumed within the Dendrinidae includes Dendrina Quenstedt, 1849; Clionolithes Clarke, 1908; Calcideletrix Mägdefrau, 1937; Dictyoporus Mägdefrau, 1937; Abeliella Mägdefrau, 1937; Nododendrina Vogel et al., 1987; and Pyrodendrina Tapanila, 2008. New combinations thereby concern Dendrina dendrina (Morris, 1851) comb. nov., Clionolithes pannosus (Solle, 1938) comb. nov., C. alcicornis (Vogel et al., 1987) comb. nov., C. convexus (Hofmann, 1996) comb. nov., Calcideletrix anomala (Mägdefrau, 1937) comb. nov., C. fastigata (Radtke, 1991) comb. nov., Dictyoporus balani (Tavernier et al., 1992) comb. nov., Nododendrina europaea (Fischer, 1875) comb. nov., N. incomposita (Mägdefrau, 1937) comb. nov. and N. paleodendrica (Elias, 1957) comb. nov. Investigation of new material and a reassessment of 63 dendrinid microborings previously addressed in informal nomenclature allowed the establishment of two complementing ichnogenera, Rhopalondendrina igen. nov. and Antodendrina igen. nov., and eight new ichnospecies, comprising Pyrodendrina arctica isp. nov., P. belua isp. nov., P. villosa isp. nov., Rhopalondendrina avis igen. et isp. nov., R. acanthina igen. et isp. nov., R. contra igen. et isp. nov., R. tigris igen. et isp. nov. and Antodendrina ligula igen. et isp. nov. In densely bioeroded calcareous substrates, different dendrinids and other bioerosion traces may be found in direct contact with each other, forming composite trace fossils, but some of these associations appear rather systematic in nature and could be the work of the same tracemaker under different behavioural modes, thus forming compound trace fossils. In these cases, however, the distinction between the two concepts remains largely equivocal. Dendrinid microborings are primarily found in living and dead calcareous skeletal substrates of bivalves, brachiopods, belemnites and corals, with complementing records from six other substrate types. Facing considerable sampling artefacts, evidence for true substrate specificity or symbiotic relationships is inconclusive as yet, whereas there is direct evidence for post-mortem infestation in several cases, such as the diverse dendrinid associations in Upper Cretaceous belemnite guards. Despite a wealth of available interpretations, the actual biological identity of the dendrinids’ tracemakers remains largely speculative. The most convincing evidence has been put forward in support of foraminiferans as the producers of Nododendrina, and excavating micro-sponges producing Clionolithes and some Calcideletrix. Since most of the dendrinids are found in aphotic (palaeo-)environments, these two principal types of organotrophic tracemakers are also potential candidates for the other ichnogenera. With regards to evolutionary patterns through geologic time, strong adaptive radiations are evident from the ichnodiversity of dendrinid ichnospecies in the Early to Mid-Palaeozoic, reflecting the “Ordovician Bioerosion Revolution” (sensu Wilson & Palmer 2006) and the “Mid-Palaeozoic Precursor of the Mesozoic Marine Revolution” (sensu Signor & Brett 1984), respectively, and in the Mesozoic, coinciding with the prominent “Marine Mesozoic Revolution” (sensu Vermeij 1977). This pattern mimics that of other micro- and macro-bioerosion trace fossils and is interpreted as a reflection of increased predation pressure and consequent infaunalisation. For extinction events, in turn, a differential effect is recorded in that the first four of the “Big Five” mass extinctions appear not to have had any noticeable effect on dendrinid ichnodiversity, whereas the end-Cretaceous mass-extinction resulted in a 77% drop following the Cretaceous peak ichnodiversity of 13 dendrinid ichnospecies.


2019 ◽  
Vol 498 (1) ◽  
pp. 129-146 ◽  
Author(s):  
Erik Wolfgring ◽  
Michael Wagreich ◽  
Ismail O. Yilmaz ◽  
Liu Shasha ◽  
Katharina Boehm

AbstractUpper Cretaceous strata at Göynük, northwestern Anatolia, Turkey, provide a geological record of the Campanian–Maastrichtian from the Sakarya Terrane along the active Neotethys margin. Shales and shaly marls with siliciclastic and volcaniclastic intercalations indicate a pelagic palaeoenvironment rich in planktonic and benthonic foraminifera and calcareous nannofossil assemblages. A composite record from the Campanian to the Maastrichtian records nannofossil standard zones UC15c (CC21) to UC20a (CC26) as well as the Globotrunanella havanensis planktonic foraminifera Zone to the Racemiguembelina fructicosa planktonic foraminifera Zone. The complete ‘mid’-Campanian to early Maastrichtian composite section can be correlated to other western Tethyan sections. The Campanian–Maastrichtian boundary is positioned between the first occurrence of the planktonic foraminifera Gansserina gansseri and the last occurrence of the nannofossil Uniplanarius trifidus. Clastic input and higher sedimentation rates constrain regional sea-level lowstands around the late Campanian calcarata Zone and the Campanian–Maastrichtian boundary, corresponding to the Campanian–Maastrichtian boundary event.


2020 ◽  
Vol 57 (3) ◽  
pp. 177-198
Author(s):  
Richard H. Groshong ◽  
Ken Kittleson

The Boulder-Weld fault zone, located southeast of Boulder, Colorado, is about 10 km (6 mi) wide, 34 km (21 mi) long, and involves at least 335 m (1100 ft) of upper Cretaceous sedimentary rocks. It affects the Cretaceous upper Pierre Shale, Fox Hills Sandstone, and the coal–bearing lower Laramie Formation. This study is a detailed examination of the eastern portion of the fault zone which consists of undisturbed areas separated by three long, narrow, fault-bounded uplifts that have received a variety of interpretations over the years. The fault zone geometry is determined from 21 closely spaced cross sections that use more subsurface data than previous studies, incorporate the elevations of the major economic coal seam derived from a published composite structure-contour map, and are area balanced using area-depth-strain (ADS) analysis. The most common structural style is a pop-up structure in which the uplifts are bounded on both sides by reverse faults. At larger-displacement the pop-ups are at the tip of the ramp and a second fault has formed close to the base of the ramp. A few sections show simple ramp anticlines developed above listric thrusts. The lower detachment for all structures is the distinctive Kp2 marker in the upper Pierre Shale. ADS analysis of the best-controlled uplifts shows that the uplifts are area balanced and confirms the lower detachment to be near Kp2. The structures are interpreted to have formed as a gravity slide because they formed in a break-back sequence, a characteristic of gravity gliding, and because the transport direction is approximately down the current southeast dip of the Kp2 detachment.


1997 ◽  
Vol 2 (4) ◽  
pp. 451-463 ◽  
Author(s):  
WILLIAM R. MOOMAW ◽  
GREGORY C. UNRUH

Environmental Kuznets curve (EKC) analysis links changes in environmental quality to national economic growth. The reduced form models, however, do not provide insight into the underlying processes that generate these changes. We compare EKC models to structural transition models of per capita CO2 emissions and per capita GDP, and find that, for the 16 countries which have undergone such a transition, the initiation of the transition correlates not with income levels but with historic events related to the oil price shocks of the 1970s and the policies that followed them. In contrast to previous EKC studies of CO2 the transition away from positive emissions elasticities for these 16 countries is found to occur as a sudden, discontinuous transition rather than as a gradual change. We also demonstrate that the third order polynomial 'N' dependence of emissions on income is the result of data aggregation. We conclude that neither the 'U'- nor the 'N'-shaped relationship between CO2 emissions and income provide a reliable indication of future behaviour.


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