New insights into the origins and radiation of the mid-Palaeozoic Gondwanan stem tetrapods

Author(s):  
John A. LONG ◽  
Alice M. CLEMENT ◽  
Brian CHOO

ABSTRACTThe earliest tetrapodomorph fishes appear in Chinese deposits of Early Devonian age, and by the Middle Devonian they were widespread globally. Evidence for the earliest digitated tetrapods comes from largely uncontested Middle Devonian trackways and Late Devonian body fossils. The East Gondwana Provence (Australasia, Antarctica) fills vital gaps in the phylogenetic and biogeographic history of the tetrapods, with the Gondwanan clade Canowindididae exhibiting a high degree of endemism within the early part of the stem tetrapod radiation. New anatomical details of Koharalepis, from the Middle Devonian Aztec Siltstone of Antarctica, are elucidated from synchrotron scan data. These include the position of the orbit, the condition of the hyomandibular, the shape of the palate and arrangement of the vomerine fangs. Biogeographical and phylogenetic models of stem tetrapod origins and radiations are discussed.

Author(s):  
A. Munkhjargal ◽  
P. Königshof ◽  
J. A. Waters ◽  
S. K. Carmichael ◽  
S. Gonchigdorj ◽  
...  

AbstractThe Bayankhoshuu Ruins section in southern Mongolia is characterized by strongly thrusted and folded sequences. Overall, three sections ranging from Ordovician to Carboniferous rocks were studied. Facies analysis combined with stratigraphic data provide improved lithostratigraphic descriptions of Palaeozoic successions in the Mushgai region. The overall marine sedimentary sequence is punctuated by volcanic rocks–basaltic lava of Silurian and Middle Devonian age and volcaniclastic bentonite and tuff in the Middle to Late Devonian and Mississippian suggesting an island arc setting. The Minjin Member of the Botuulkhudag Formation (Middle Devonian to Late Devonian) is primarily composed of thick basaltic and subaerial volcanic rocks with minor silicified siltstone and chert inclusions. Thicker successions of limestone occur in the Ordovician/Silurian, Early Devonian, and the Mississippian. The macrofauna is scarce, except distinct limestone horizons where different fossil groups were recognized. Microfossils, such as radiolarians and conodonts, are scarce and generally poorly preserved. However, based on the re-study of collections from earlier publications and new conodont data, a more detailed biostratigraphic record of the Khoyormod, Botuulkhudag, and Arynshand formations of the Bayankhoshuu Ruins section can be developed. For instance, the Arynshand Formation likely ranges from the late Bispathodus ultimus conodont biozone to the Scaliognathus anchoralis–Doliognathus latus conodont biozone. A tectonic breccia occurs in the early Mississippian and is overlain by a red shale of remarkable thickness at the top of this formation which points to subaerial exposure in the early Mississippian (near the Tournaisian/Visean transition). Due to strong tectonic overprint and/or facies, some unconformities/hiatuses occur. Most strata are intensively folded and faulted, ranging from centimeter to meter scale. Overall, deposition likely occurred on either the Mandalovoo or Gurvansayhan Terrane.


2002 ◽  
Vol 76 (2) ◽  
pp. 229-238
Author(s):  
Zhong-Qiang Chen ◽  
Neil W. Archbold

Two new genera of the Chonostrophiidae are proposed herein to accommodate the resupinate shells from the Famennian sediments of the Late Devonian in the Santanghu Basin of the Balikun area, Xinjiang Province, northwestern China. Santanghuia santanghuensis new genus and species is distinguishable from other chonostrophiids by the possession of a pair of long dorsal anderidia and absence of a dorsal median septum. Balikunochonetes liaoi new genus and species is distinct because of the presence of a pair of anderidia with secondary anderidia, and a dorsal median septum. Santanghuia new genus is considered to be phylogenetically related to Chonostrophia of late Early to Middle Devonian age, while Balikunochonetes has possibly given rise to Chonostrophiella of Early Devonian age and is a likely ancestor of Tulcumbella of Early Carboniferous age.


Author(s):  
Alexander J.P. Houben ◽  
Geert-Jan Vis

Abstract Knowledge of the stratigraphic development of pre-Carboniferous strata in the subsurface of the Netherlands is very limited, leaving the lithostratigraphic nomenclature for this time interval informal. In two wells from the southwestern Netherlands, Silurian strata have repeatedly been reported, suggesting that these are the oldest ever recovered in the Netherlands. The hypothesised presence of Silurian-aged strata has not been tested by biostratigraphic analysis. A similar lack of biostratigraphic control applies to the overlying Devonian succession. We present the results of a palynological study of core material from wells KTG-01 and S05-01. Relatively low-diversity and poorly preserved miospore associations were recorded. These, nonetheless, provide new insights into the regional stratigraphic development of the pre-Carboniferous of the SW Netherlands. The lower two cores from well KTG-01 are of a late Silurian (Ludlow–Pridoli Epoch) to earliest Devonian (Lochkovian) age, confirming that these are the oldest sedimentary strata ever recovered in the Netherlands. The results from the upper cored section from the pre-Carboniferous succession in well KTG-01 and the cored sections from the pre-Carboniferous succession in well S05-01 are more ambiguous. This inferred Devonian succession is, in the current informal lithostratigraphy of the Netherlands, assigned to the Banjaard group and its subordinate Bollen Claystone formation, of presumed Frasnian (i.e. early Late Devonian) age. Age-indicative Middle to Late Devonian palynomorphs were, however, not recorded, and the overall character of the poorly preserved palynological associations in wells KTG-01 and S05-01 may also suggest an Early Devonian age. In terms of lithofacies, however, the cores in well S05-01 can be correlated to the upper Frasnian – lower Famennian Falisolle Formation in the Campine Basin in Belgium. Hence, it remains plausible that an unconformity separates Silurian to Lower Devonian strata from Upper Devonian (Frasnian–Famennian) strata in the SW Netherlands. In general, the abundance of miospore associations points to the presence of a vegetated hinterland and a relatively proximal yet relatively deep marine setting during late Silurian and Early Devonian times. This differs markedly from the open marine depositional settings reported from the Brabant Massif area to the south in present-day Belgium, suggesting a sediment source to the north. The episodic presence of reworked (marine) acritarchs of Ordovician age suggests the influx of sedimentary material from uplifted elements on the present-day Brabant Massif to the south, possibly in relation to the activation of a Brabant Arch system.


2014 ◽  
Vol 62 (1) ◽  
pp. 44 ◽  
Author(s):  
Gavin C. Young ◽  
John A. Long

A small collection of arthrodire remains is described from the Devonian Aztec Siltstone of southern Victoria Land, Antarctica. Barwickosteus antarcticus, gen. et sp. nov., is a small phlyctaeniid arthrodire probably closely related to Barrydalaspis from the Bokkeveld Group of South Africa. Grifftaylor antarcticus, gen. et sp. nov., is a generalised phlyctaeniid resembling Phlyctaenius and Neophlyctaenius. New specimens of Boomeraspis show that it had a high-spired trunk-armour with a median dorsal plate of similar proportions to Tiaraspis, Mithakaspis, Turrisaspis or Africanaspis. Other fragmentary median dorsal plates are provisionally referred to Turrisaspis and Mulgaspis. With these new taxa the vertebrate assemblage from the Aztec Siltstone comprises at least 37 genera and 50 species, making it one of the most diverse of Middle–Late Devonian age.


2017 ◽  
Author(s):  
Christine D. Bacon ◽  
Francisco Velásquez-Puentes ◽  
Luis Felipe Hinojosa ◽  
Thomas Schwartz ◽  
Bengt Oxelman ◽  
...  

Several studies have demonstrated the contribution of northern immigrants to the flora of the tropical Andes – the world’s “hottest” biodiversity hotspot. However, much less is known about the biogeographic history and diversification of Andean groups with southern origins, although it has been suggested that northern and southern groups have contributed roughly equally to the high Andean (i.e. páramo) flora. Here we infer the evolutionary history of the southern hemisphere plant genus Gunnera, a lineage with a rich fossil history and an important ecological role as an early colonising species characteristic of wet, montane environments. Our results show striking contrasts in species diversification, where some species may have persisted for some 90 million years, whereas others date to less than 2 Ma since origination. The outstanding longevity of the group is likely linked to a high degree of niche conservatism across its highly disjunct range, whereby Gunnera tracks damp and boggy soils in cool habitats. Colonisation of the northern Andes is related to Quaternary climate change, with subsequent rapid diversification appearing to be driven by their ability to take advantage of environmental opportunities. This study demonstrates the composite origin of a mega-diverse biota.


Author(s):  
John Graham ◽  
Nancy Riggs

The Silurian Croagh Patrick succession, which crops out just south of a fundamental Caledonian structural zone near Clew Bay, western Ireland, is a series of psammites and pelites with a strong penetrative cleavage. These rocks are intruded by the Corvock granite. A suite of minor intrusions associated with the granite contains the regional cleavage whereas the Corvock granite is undeformed. New U-Pb dates are 413 + 7 / -4 Ma for a strongly cleaved sill and 410 ± 4 Ma for the main granite and closely constrain the age of crystallization of the granite and coeval cleavage formation as Lower Devonian (Lochkovian or Pragian), implying syn- to late-kinematic granite emplacement. These data are consistent with evidence for strong sinistral shear shown by the Ox Mountains granodiorite just to the north-east dated at 412.3 ± 0.8 Ma. This Devonian cleavage is superimposed on Ordovician rocks of the South Mayo Trough. The localisation of the strong deformation is interpreted as being due to its position at a restraining bend during regional sinistral motion on a segment of the Fair Head-Clew Bay Line to the north. Contemporaneous deformation in the syn-kinematic Donegal batholith suggests a transfer of sinistral motion to this intra-Grampian structure rather than simple along-strike linkage to the Highland Boundary Fault in Scotland. Our new data indicate diachronous deformation during the late Silurian and early Devonian history of the Irish and Scottish Caledonides and also support previous interpretations of diachronous deformation between these areas and the Appalachian orogens.


2004 ◽  
Vol 141 (3) ◽  
pp. 329-344 ◽  
Author(s):  
M. A. KENDRICK ◽  
E. A. EIDE ◽  
D. ROBERTS ◽  
P. T. OSMUNDSEN

The regionally significant 0.5–2 km thick Høybakken detachment in central Norway bounds the southern margin of the Central Norway Basement Window and exhibits a well-developed top-to-the-WSW fabric characteristic of late Scandian, Devonian ductile extension. 40Ar–39Ar data obtained from hornblende, mica and K-feldspar mineral separates of rocks collected in a transect through the Høybakken detachment yield well-defined plateau and isochron mineral ages. Early Devonian exhumation and cooling of the Høybakken detachment footwall is recorded by hornblende ages of ∼ 400 Ma and mica ages of ∼ 390 Ma. The mylonitic fabric overlying the footwall records younger Middle Devonian mica crystallization ages of 384–381 Ma that are among the youngest extensional ductile fabrics dated in the Caledonides and suggest prolonged extensional activity on the Høybakken detachment. After inferred cessation of ductile extension at 381 Ma, the rate of uplift and cooling was reduced, and the footwall records Late Devonian to Early Carboniferous K-feldspar ages of 371–356 Ma. Prolonged extensional activity at Høybakken is compatible with recent U–Pb ages of deformed titanite crystals and established Rb–Sr ages of white mica in shear-related pegmatites, both from the southwestern part of the Fosen Peninsula, and 40Ar–39Ar ages of syn-tectonic mica overgrowth from the adjacent Hitra–Snåsa Fault. Together, these ages suggest the onset of ductile extension soon after ∼ 401 Ma, and with the Middle Devonian crystallization ages determined here, suggest that ductile extension on the Høybakken detachment had a duration of 11–20 Ma. The youngest age of 320 Ma was obtained from a K-feldspar in a cataclastic granite of the Høybakken detachment's hangingwall and is considered to date a phase of post-Scandian brittle deformation that overprinted the mylonitic shear fabric.


During the Silurian and Devonian, the sequence of continental collisions that were ultimately to result in the formation of the supercontinent of Pangaea had begun. By the Early to Middle Devonian North America (Laurentia), Acadia, Great Britain, and Northern Europe (Baltica) had collided to form the ‘Old Red Sandstone’ continent (Laurussia). Palaeomagnetic data, however, indicate that the configuration of the continents that made up Laurussia did not resemble the pre-breakup, Mesozoic reassembly. Rather, Britain, Baltica, and Acadia were displaced 10—20° to the south with respect to Laurentia. New palaeomagnetic data for Laurentia and Gondwana, suggests that the ocean separating the northern and southern continents was relatively narrow during the early Devonian, and may have been nearly closed by the late Devonian.


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