Seed cones of Metasequoia milleri from the Middle Eocene of southern British Columbia

1984 ◽  
Vol 62 (2) ◽  
pp. 281-289 ◽  
Author(s):  
James F. Basinger
Keyword(s):  
British Columbia ◽  
Middle Eocene ◽  
Close Association ◽  
Single Species ◽  
Metasequoia Glyptostroboides ◽  
Adaxial Surface ◽  
Close Resemblance ◽  
Cone Axis ◽  
Living Species ◽  
Cone Scale

Permineralized ovulate cones of Metasequoia milleri Rothwell and Basinger (Taxodiaceae) are preserved in silicified peats of the Middle Eocene Allenby Formation near Princeton, B.C., Canada. Cones are about 17 mm wide and 25 mm long and are borne terminally on sparsely leaved stalks or peduncles. Twenty to 30 cone scales are arranged decussately on the cone axis. Externally, cone scales are broadly hexagonal and labiate in appearance owing to a medial cleft. Several ovules, with micropyles directed toward the cone axis, are attached to the adaxial surface of the cone scale. Seeds are 3–4 mm wide and about 5 mm long, with two lateral wings each approximately equal in size to the central seed body. Anatomical similarity to and close association with pollen cones and vegetative remains of M. milleri indicate that all organs belong to a single species. The vegetative body and pollen cones of M. milleri differ in few respects from those of living Metasequoia glyptostroboides Hu and Cheng. The seed cones of M. milleri cannot be distinguished from those of the living species. The close resemblance of the two species indicates evolutionary near stasis, at least structurally, throughout the Tertiary.

Metasequoia milleri n.sp., anatomically preserved pollen cones from the Middle Eocene (Allenby Formation) of British Columbia

1979 ◽  
Vol 57 (8) ◽  
pp. 958-970 ◽  
Author(s):  
Gar W. Rothwell ◽  
James F. Basinger
Keyword(s):  
British Columbia ◽  
Middle Eocene ◽  
Metasequoia Glyptostroboides ◽  
Plant Remains ◽  
Exine Ornamentation ◽  
Pollen Sacs ◽  
Pollen Cone ◽  
Fertile Region ◽  
A New Species ◽  
Subtending Leaves

The discovery of well preserved, permineralized plant remains in Eocene sediments near Princeton, British Columbia, provides an opportunity to describe Metasequoia milleri, a new species of taxodiaceous pollen cone. Individual specimens are up to 3.0 mm long and 2.9 mm in diameter and are subtended by a vegetative zone of scale-like leaves. Approximately 30 microsporophylls are attached to the axis, and each bears three ovoid pollen sacs. The distal-most subtending leaves imbricate and enclose the fertile region. Pollen is ovoid to subspheroidal with an erect, protruding leptoma. Grains measure 19–27 μn in diameter and have verrucate exine ornamentation with numerous orbicules. These fossils show that pollen cones anatomically similar to those of extant Metasequoia glyptostroboides were present as early as Middle Eocene time.

The vegetative body of Metasequoia milleri from the Middle Eocene of southern British Columbia

1981 ◽  
Vol 59 (12) ◽  
pp. 2379-2410 ◽  
Author(s):  
James F. Basinger
Keyword(s):  
British Columbia ◽  
Epidermal Cell ◽  
Cell Walls ◽  
Middle Eocene ◽  
Secondary Phloem ◽  
Metasequoia Glyptostroboides ◽  
Stem Anatomy ◽  
Marsh Soil ◽  
Anatomical Features ◽  
Resin Ducts

Anatomically preserved vegetative remains of Metasequoia milleri Rothwell and Basinger are common in the Princeton chert of the Allenby Formation in southern British Columbia. Deposition of the Allenby Formation and associated volcanics occurred during Middle Eocene time. The Princeton chert locality is in the upper strata of the Allenby Formation and is of late Middle Eocene age. The Princeton chert was formed by siliceous permineralization of marsh soil. Dissolved silicates were probably introduced by periodic influx of water from mineral springs or geysers.Anatomical features of stems, wood, and leaves are well preserved. Mature wood of the fossil resembles that of Metasequoia glyptostroboides in having traumatic resin cysts; opposite pitting on radial walls of tracheids; taxodioid cross-field pitting; tall, uniseriate rays; smooth-walled ray parenchyma; and diffuse, resinous, smooth-walled wood parenchyma. Leaves are linear, hypostomatic, and borne decussately, and have one or three resin ducts and slightly undulate to smooth epidermal cell walls. Leaves of living M. glyptostroboides differ in consistently having three resin ducts and much more pronounced undulations of epidermal cell walls. Metasequoia milleri has leaves of a generalized structure from which leaf types of many taxodiaceous genera could be derived. Roots associated with M. milleri are dimorphic. Primary tissues and secondary phloem are exceedingly well preserved. Cortex of both long and short roots contains mycorrhizal fungi.Compression remains of M. occidentalis are not distinguishable from M. glyptostroboides. Anatomical features of M. milleri, however, do reveal differences between Eocene and living Metasequoia. Wood rays of M. milleri are much higher than those of M. glyptostroboides. Other features of stem anatomy are similar in both species. Pollen cones differ in developmental and minor anatomical features. The close similarity of most organs of M. milleri to M. glyptostroboides and the dissimilarity of leaf structure indicate mosaic evolution within the genus.

Vegetative Growth of Eorhiza arnoldii Robison and Person from the Middle Eocene Princeton Chert Locality of British Columbia

1994 ◽  
Vol 155 (5) ◽  
pp. 606-616 ◽  
Author(s):  
Ruth A. Stockey ◽  
Kathleen B. Pigg
Keyword(s):  
British Columbia ◽  
Vegetative Growth ◽  
Middle Eocene

Further observations on Paleorosa similkameenensis (Rosaceae) from the Middle Eocene Princeton chert of British Columbia, Canada

1993 ◽  
Vol 78 (3-4) ◽  
pp. 277-291 ◽  
Author(s):  
Sergio R.S. Cevallos-Ferriz ◽  
Diane M. Erwin ◽  
Ruth A. Stockey
Keyword(s):  
British Columbia ◽  
Middle Eocene

Fossil Mosses from the Bisaccate Zone of the Mid-Eocene Allenby Formation, British Columbia

1974 ◽  
Vol 11 (3) ◽  
pp. 409-421 ◽  
Author(s):  
Marian Kuc
Keyword(s):  
British Columbia ◽  
Middle Eocene ◽  
Plant Tissues ◽  
Plant Remains ◽  
Lake Bottom ◽  
White Colour

New fossil taxa (Ditrichites fylesi, Muscites maycocki, M. ritchiei, Palaeohypnum jovet-asti and P. steerei); unnamed moss and moss-like fossils, detrital fragments of various plant tissues, and paleobotanical evidence of the bisaccate zone are described from the Middle Eocene Allenby Formation near Princeton, British Columbia. These remains occur in laminated, tuffaceous, silty and pyroclastic shale, deposited under lacustrine conditions.Detailed examination of the various laminae indicates that beds of white colour and composed of coarser silt grains are poor in fossils and could be related to periods of decreasing bioproduction; less silty and darker coloured beds are rich in macro- and microfossils and could be related to periods of extensive bioproduction. The rock features, lamination, and distribution of macrofossils indicate the slow and undisturbed accumulation of plant remains on a lake bottom.

Eocene age for Ag–Pb–Zn–Au vein and replacement deposits of the Kokanee Range, southeastern British Columbia

1992 ◽  
Vol 29 (1) ◽  
pp. 3-14 ◽  
Author(s):  
G. Beaudoin ◽  
J. C. Roddick ◽  
D. F. Sangster
Keyword(s):  
British Columbia ◽  
Middle Jurassic ◽  
Middle Eocene ◽  
Hanging Wall ◽  
Spatial Association ◽  
High Heat ◽  
Time Interval ◽  
Genetic Link ◽  
Metasedimentary Rocks ◽  
Magmatic Event

The Ag–Pb–Zn–Au vein and replacement deposits of the Kokanee Range, southeastern British Columbia, are hosted by the Middle Jurassic Nelson batholith and surrounding Cambrian to Triassic metasedimentary rocks in the hanging wall of the transcrustal Slocan Lake Fault, Field relations indicate that mineralization is younger than the Nelson batholith and a Middle Jurassic foliation in the Ainsworth area but coeval or older than Eocene unroofing of the Valhalla metamorphic core complex in the footwall of the Slocan Lake Fault. Lamprophyre and gabbro dykes are broadly coeval with mineralization and have biotite and hornblende K–Ar ages defining a short-lived Middle Eocene alkaline magmatic event between 52 and 40 Ma. An older, Early Cretaceous alkaline magmatic event (141 – 129 Ma) is possible but incompletely documented.K–Ar and step-heating 40Ar/39Ar analyses on hydrothermal vein and alteration muscovite indicate that hydrothermal fluids were precipitating vein and replacement deposits 58–59 Ma ago. Crosscutting relationships with lamprophyre dykes indicate the Kokanee Range hydrothermal system lasted for more than 15 Ma. Eocene crustal extension resulted in a high heat flow and structures which were probably responsible for hydrothermal fluid movement and flow paths.A 100 Ma time interval is documented between batholith emplacement and spatially associated mineralization, ruling out any genetic link between the two. Similar large age differences between granite intrusion and peripheral mineralization have recently been documented for two world-sea le Ag–Pb–Zn vein districts, which suggest that spatial association between granite and Ag–Pb–Zn mineralization is not sufficient to infer a genetic link.

Two new species of stalked crinoids from the northeastern Pacific in the genera Gephyrocrinus and Ptilocrinus (Echinodermata, Crinoidea, Hyocrinidae). Effects of ontogeny and variability on hyocrinid taxonomy

Zootaxa ◽  
2011 ◽  
Vol 2825 (1) ◽  
pp. 1 ◽  
Author(s):  
MICHEL ROUX ◽  
PHILIP LAMBERT
Keyword(s):  
British Columbia ◽  
New Species ◽  
Single Species ◽  
Depth Range ◽  
Evolutionary Trend ◽  
Exceptional Collection ◽  
The Family ◽  
Two New Species ◽  
Northeastern Pacific ◽  
Stalked Crinoids

Two new species of deep-sea stalked crinoids belonging to the family Hyocrinidae were collected in the northeastern Pacific. The descriptions contain detailed information on character variations and ontogeny. The five specimens of Gephyrocrinus messingi n. sp. lived at depths ranging from 1,777 m to 2,110 m off British Columbia and California. This new species is the first record of the genus Gephyrocrinus in the Pacific Ocean, which was previously known from only a single species, G. grimaldii, from the northeastern Atlantic at the same depth range. The two species illustrate opposing phenotypes within the same genus. Fifty-eight specimens of the second new species, Ptilocrinus clarki n. sp., were dredged off British Columbia close to the type-locality of P. pinnatus, the type species of the genus Ptilocrinus, but at shallower depths ranging from 1,178 to 1,986 m. This exceptional collection provides significant data on intraspecific variation in the main morphological characters, especially arm pattern. The ontogeny of stalk articulations and the main traits of adoral plate differentiation are described in detail. A complementary investigation on P. pinnatus was conducted using specimens collected by the “Albatross” expedition at a depth of 2,906 m. Despite similarities in external morphology, tegmen and cover plates, the two ptilocrinid species display significant differences in pinnule architecture, aboral cup and stalk articulations. From comparison with Gephyrocrinus messingi n. sp. and Ptilocrinus clarki n. sp., G. grimaldii and P. pinnatus are interpreted as the result of heterochronic development by paedomorphy after ecological or geographic isolation. Pinnule architecture in the two new species suggests first steps in an evolutionary trend toward a rigid box which protects gonad inflation in the proximal part of the pinnule. These new data on Ptilocrinus and Gephyrocrinus create problems in the current taxonomy of the family Hyocrinidae. The main derived characters, especially in pinnule and arm pattern, are used to propose new hypotheses for hyocrinid phylogeny.

scholarly journals Additional records and stratigraphic distribution of the middle Eocene carettochelyid turtle Anosteira pulchra from the Uinta Formation of Utah, North America

PeerJ ◽  
2020 ◽  
Vol 8 ◽  
pp. e9775
Author(s):  
Brent Adrian ◽  
Patricia A. Holroyd ◽  
J. Howard Hutchison ◽  
KE Beth Townsend
Keyword(s):  
North America ◽  
New Records ◽  
Middle Eocene ◽  
Single Species ◽  
Uinta Basin ◽  
Stratigraphic Framework ◽  
Multiple Levels ◽  
Land Mammal ◽  
Anatomical Information ◽  
Insight Into

Background Anosteira pulchra is one of two species of the obligately-aquatic freshwater clade Carettochelyidae (pig-nosed turtles) from the Eocene of North America. Anosteira pulchra is typically rare in collections, and their distribution is poorly documented. The Uinta Formation [Fm.] contains a diverse assemblage of turtles from the Uintan North American Land Mammal Age. Whereas turtles are abundantly preserved in the Uinta Fm., A. pulchra has been reported only from a few specimens in the Uinta C Member. Methods We describe new records of Anosteira pulchra from the Uinta Basin and analyze the distribution of 95 specimens from multiple repositories in the previously published stratigraphic framework of the middle and upper Uinta Fm. Results Here we report the first records of the species from the Uinta B interval, document it from multiple levels within the stratigraphic section and examine its uncommon appearance in only approximately 5% of localities where turtles have been systematically collected. This study details and extends the range of A. pulchra in the Uinta Fm. and demonstrates the presence of the taxon in significantly lower stratigraphic layers. These newly described fossils include previously unknown elements and associated trace fossils, with new anatomical information presented. This study provides insight into the taxonomy of Anosteira spp. in the middle Eocene, and suggests the presence of a single species, though no synonymy is defined here due to limits in Bridger material.

Eocene paleo-physiography and drainage directions, southern Interior Plateau, British Columbia

2005 ◽  
Vol 42 (2) ◽  
pp. 215-230 ◽  
Author(s):  
Selina Tribe
Keyword(s):  
British Columbia ◽  
Middle Eocene ◽  
Regional Scale ◽  
River Valley ◽  
Coast Mountains ◽  
Base Level ◽  
The North ◽  
Bedrock Geology ◽  
River Valleys ◽  
Early Cenozoic

A map of reconstructed Eocene physiography and drainage directions is presented for the southern Interior Plateau region, British Columbia south of 53°N. Eocene landforms are inferred from the distribution and depositional paleoenvironment of Eocene rocks and from crosscutting relationships between regional-scale geomorphology and bedrock geology of known age. Eocene drainage directions are inferred from physiography, relief, and base level elevations of the sub-Eocene unconformity and the documented distribution, provenance, and paleocurrents of early Cenozoic fluvial sediments. The Eocene landscape of the southern Interior Plateau resembled its modern counterpart, with highlands, plains, and deeply incised drainages, except regional drainage was to the north. An anabranching valley system trending west and northwest from Quesnel and Shuswap Highlands, across the Cariboo Plateau to the Fraser River valley, contained north-flowing streams from Eocene to early Quaternary time. Other valleys dating back at least to Middle Eocene time include the North Thompson valley south of Clearwater, Thompson valley from Kamloops to Spences Bridge, the valley containing Nicola Lake, Bridge River valley, and Okanagan Lake valley. During the early Cenozoic, highlands existed where the Coast Mountains are today. Southward drainage along the modern Fraser, Chilcotin, and Thompson River valleys was established after the Late Miocene.

Taxonomy, phylogeny, and paleoecology of Eoseira wilsonii gen. et sp. nov., a Middle Eocene diatom (Bacillariophyceae: Aulacoseiraceae) from lake sediments at Horsefly, British Columbia, Canada

2005 ◽  
Vol 42 (2) ◽  
pp. 243-257 ◽  
Author(s):  
Alexander P Wolfe ◽  
Mark B Edlund
Keyword(s):  
British Columbia ◽  
Middle Eocene ◽  
Life Strategy ◽  
Centric Diatom ◽  
Evolutionary Trajectory ◽  
The Family ◽  
Freshwater Diatoms ◽  
Marine Realm ◽  
Diatom Genus ◽  
Early Cenozoic

A new centric diatom genus is described from laminated freshwater sediments of Middle Eocene age near Horsefly, British Columbia, Canada. This diatom, Eoseira wilsonii gen. et sp. nov., grew in filaments that constitute dense monospecific sub-horizons within the white couplets that represent summer deposition in the varved sequence. Although Eoseira clearly belongs within the Family Aulacoseiraceae, several features distinguish its valve structure from Aulacoseira. Gigantism of spines and the lack of geometric relationships between spine position and mantle areolae are the most conspicuous features of the genus. Eoseira is among the oldest freshwater diatoms known and one of relatively few extinct freshwater genera. In addition to serving as a potential biostratigraphic marker, Eoseira is a cornerstone to undertanding the evolutionary trajectory of the Aulacoseiraceae, likely the oldest family of freshwater diatoms. As such, it refines our understanding of early radiations from the marine realm in western North America. Furthermore, paleoecological inferences based on Eoseira life strategy pertain directly to limnological conditions during early Cenozoic hot-house conditions.

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