Floral development of Rosa setigera

1993 ◽  
Vol 71 (1) ◽  
pp. 74-86 ◽  
Author(s):  
James R. Kemp ◽  
Usher Posluszny ◽  
Jean M. Gerrath ◽  
Peter G. Kevan

The development of the flower of Rosa setigera from initiation to the onset of anthesis is described. Rosa setigera is the only known member of the genus Rosa to exhibit dioecy. Flowers of functionally staminate (male) and functionally carpellate (female) plants appear identical, a condition referred to as cryptic dioecy. Discrete sepals and petals are formed on the floral meristem. As the hypanthium forms, stamens are initiated in alternating whorls on the wall of the hypanthium and continue to develop as the hypanthium extends. Carpel primordia arise individually on the remainder of the floral meristem and show neither adnation to the hypanthial wall nor coalescence to one another as they give rise to the styles and stigmas that are exserted above the hypanthium lip. The only observable fusion in this species appears to be the postgenital fusion of the margins of the carpel primordia to form the enclosed locule. Although historically the hypanthium has been variously interpreted as either axial and (or) appendicular in nature, resulting from congenital fusion of sepals, petals, and stamens, this paper uses a more realistic, testable and functional approach to the development of the hypanthium that is in keeping with current concepts such as process morphology. Key words: Rosa setigera, dioecy, floral development, fusion, hypanthium.


1991 ◽  
Vol 69 (4) ◽  
pp. 844-857 ◽  
Author(s):  
Taylor A. Steeves ◽  
Margaret W. Steeves ◽  
A. Randall Olson

The development of the flower of Amelanchier alnifolia from initiation to the onset of anthesis is described. Sepals are formed sequentially, but interprimordial zonal growth results in the initiation of the hypanthium. Petals and stamens arise in whorls around the floral meristem as the hypanthium extends. They show neither coalescence nor adnation and do not appear to contribute to the development of the hypanthium. Gynoecial primordia arise individually, give rise to the styles and stigmas, and are joined basally by zonal growth to produce the roof of the ovary. The wall of the inferior ovary is interpreted as a gynoecial hypanthium. It is difficult to determine the extent to which the gynoecial primordia contribute to the development of the ovary. They do not give rise to most of its structure but may be responsible for the initiation of the ovules. There is evidence of postgenital fusion of the septal margins as they converge in the centre of the ovary. The timing of events in floral development is recorded for the locality of the study. The observations are discussed in relation to current theories concerning the nature of the inferior ovary. Key words: Amelanchier, flower, development, inferior ovary, hypanthium.



Author(s):  
Svitlana Gruschko

The article investigates the role of communicative-functional approach in the translation of scientific and technical literature, when the translation itself is seen as a tool to accomplish purposeful activity of communicators, representatives of different interlingual groups. In practical terms, the difficulties of translating scientific and technical texts are due to the peculiarities of scientific style, insufficient understanding of the terminology of a particular field of knowledge, which complicates the choice of adequate translation solutions. In this context, an important role is played by the translation strategy, where the communicativefunctional approach dominates. The translation is implicitly included in the communication structure between the author and the recipient. The relevance of the article is determined by the necessity of studying the problems of scientific and technical literature translation, taking into account the current realities of interlanguage information exchange. Key words: communicative-functional approach, interlingual communication, translation, structure



2020 ◽  
pp. 125-127
Author(s):  
Мээрим Рахматалиева

Аннотация: Коммуникативдик окуу китептери окутуунун алдыңкы принциби катары жарыяланган активдүү коммуникативдүүлүктүн негизинде түзүлөт. Бул принципке ылайык тилге үйрөтүү кептик коммуникациянын жүрүшүнө шайкеш болуусу абзел. Мында окутуунун тажрыйбалык багыты, тилдик материалдын тандалышы жана берилишине карата функционалдык мамиле, окуу материалынын кырдаалдык-тематикалык багыты, окуучулардын жекече-психологиялык өзгөчөлүгү жана алардын өз эне тилинин таасири эске алынат. Коммуникативдик окутуунун маселелерин Е.И. Пассов илимий изилдөөлөрүндө карап чыгып, «коммуникативдик метод» деген терминди сунуштаган. Азыркы учурда коммуникативдик окуу китептери кеп ишмердүүлүгүнүн бардык түрүн окутууда колдонулат. Коммуникативдик окуу китептеринде окутуунун негизги акценти тилдин формалдуу белгилерин терең үйрөтүүдөн чегинип, кептик материал менен аракет жүргүзүүгө басым кылынган. Алар тилдин эрежелерине караганда кептик көнүгүүлөрдүн көп болушу менен, атайын грамматикалык материалдын дээрлик жоктугу менен, ошондой эле текстке жараша көргөзмө куралдын кеңири берилиши менен мүнөздөлөт. Түйүндүү сөздөр: тил, коммуникативдүүлүк, компетенция, окуу китеби, принцип, кеп, коммуникация, өзгөчөлүктөр, окуу куралы, илим, окуу комплекси, кеп ишмердүүлүгү Аннотация: В основе коммуникативных учебников лежит принцип активной коммуникативности, провозглашенный в качестве ведущего принципа обучения. Согласно этому принципу обучение языку строится адекватно процессу речевой коммуникации, отмечается практическая направленность обучения, функциональный подход к отбору и подаче языкового материала, ситуативно-тематическое представление учебного материала, учет индивидуально-психологических особенностей и родного языка учащихся. Проблемы коммуникативного обучения стали объектом научных исследований Е.И. Пассова, который предложил термин «коммуникативный метод». На данный момент коммуникативные учебники используются для обучения всем видам речевой деятельности. В коммуникативных учебниках основной акцент обучения направлен на работу с речевым материалом, в то время как развернутый грамматический материал отсутствует. Такие учебники характеризуются широко используемой наглядностью, иллюстрирующей тексты коммуникативного направления. Ключевые слова: язык, коммуникативность, компетенция, учебник, принцип, речь, коммуникация, особенности, учебниковедение, учебный комплекс, речевая деятельность. Abstract: The basis of communicative textbooks is the principle of active communication, proclaimed as the leading principle of learning. According to this principle, language teaching is built adequately to the process of speech communication, there is a practical orientation of training, a functional approach to the selection and presentation of language material, situational and thematic presentation of educational material, taking into account individual psychological characteristics and the native language of students. The problems of communicative training became the object of scientific research E.I. Passov, who proposed the term "communicative method". At the moment, communicative textbooks are used to teach all types of speech activity. In communicative text-books, the main emphasis of training is aimed at working with speech material, while the expanded grammatical material is almost absent. Such textbooks are characterized by widely used visibility illustrating texts of a communicative direction. Key words: language, communicativeness, competence, textbook, principle, speech, communication, features, textbook studies, educational complex, speech activity.



1973 ◽  
Vol 51 (3) ◽  
pp. 647-656 ◽  
Author(s):  
U. Posluszny ◽  
R. Sattler

The floral appendages of Potamogeton densus are initiated in an acropetal sequence. The first primordia to be seen externally are those of the lateral tepals, though sectioning young floral buds (longitudinally, parallel to the inflorescence axis) reveals initial activity in the region of the lower median (abaxial) tepal and stamen at a time when the floral meristem is not yet clearly demarcated. The lateral (transversal) stamens are initiated simultaneously and unlike the median stamens each arises as two separate primordia. The upper median (adaxial) tepal and stamen develop late in relation to the other floral appendages, and in some specimens are completely absent. Rates of growth of the primordia vary greatly. Though the lower median tepal and stamen are initiated first, they grow slowly up to gynoecial inception, while the upper median tepal appears late in the developmental sequence but grows rapidly, soon overtaking the other tepal primordia. The four gynoecial primordia arise almost simultaneously, although variation in their sequence of inception occurs. The two-layered tunica of the floral apices gives rise to all floral appendages through periclinal divisions in the second layer. The third layer (corpus) is involved as well in the initiation of the stamen primordia. Procambial strands develop acropetally, lagging behind primordial initiation. The lateral stamens though initiating as two primordia each form a single, central procambial strand, which differentiates after growth between the two primordia of the thecae has occurred. A great amount of deviation from the normal tetramerous flower is found, including completely trimerous flowers, trimerous gynoecia with tetramerous perianth and androecium, and organs differentiating partially as tepals and partially as stamens.



2020 ◽  
Author(s):  
Liling Yang ◽  
Shilian Qi ◽  
Arfa touqeer ◽  
Haiyang Li ◽  
Xiaolan Zhang ◽  
...  

Abstract Background: Flower development directly affects fruit production in tomato. Despite the framework mediated by ABC genes have been established in Arabidopsis, the spatiotemporal precision of floral development in tomato has not been well examined.Results: Here, we analyzed a novel tomato stamenless like flower (slf) mutant in which the development of stamens and carpels is disturbed, with carpelloid structure formed in the third whorl and ectopic formation of floral and shoot apical meristem in the fourth whorl. Using bulked segregant analysis (BSA), we assigned the causal mutation to the gene Solanum lycopersicum GT11 (SlGT11) that encodes a transcription factor belonging to Trihelix gene family. SlGT11 is expressed in the early stages of the flower and the expression becomes more specific to the primordium position corresponding to stamens and carpels in later stages of the floral development. Further RNAi silencing of SlGT11 verifies the defective phenotypes of the slf mutant. The carpelloid stamen in slf mutant indicates that SlGT11 is required for B-function activity in the third whorl. The failed termination of floral meristem and the occurrence of floral reversion in slf indicate that part of the C-function requires SlGT11 activity in the fourth whorl. Furthermore, we find that at higher temperature, the defects of slf mutant are substantially enhanced, with petals transformed into sepals, all stamens disappeared, and the frequency of ectopic shoot/floral meristem in fourth whorl increased, indicating that SlGT11 functions in the development of the three inner floral whorls. Consistent with the observed phenotypes, it was found that B, C and an E-type MADS-box genes were in part down regulated in slf mutants.Conclusions: Together with the spatiotemporal expression pattern, we suggest that SlGT11 functions in floral organ patterning and maintenance of floral determinacy in tomato.



1992 ◽  
Vol 70 (2) ◽  
pp. 258-271 ◽  
Author(s):  
Claudia Erbar

The early floral development of Stylidium adnatum and Stylidium graminifolium is characterized by an initial circular primordium whose areas in the transversal plane of the floral primordium show enhanced growth. The spiral inception of the five sepals starts before the differentiation of the initial circular primordium into two stamen primordia in transversal position (in relation to the floral diagram) and the corolla ring primordium below the stamen primordia. Then five petal primordia, which alternate with the sepals, arise on the corolla ring primordium (early sympetaly). Peculiar to the flowers of Stylidiaceae is the column that bears at its top both stigma and anthers. Probably this column should be interpreted as a receptacular tube. No distinct carpel primordia have been observed. The inferior ovary results from intercalary growth in the peripheral parts of the receptacle below the calyx, corolla, and stamen primordia. The residual floral apex gives rise to a transversal septum, by which the ovary becomes bilocular. None of the morphological, palynological, and embryological characters discussed contradicts a position of the Stylidiaceae near the Campanulales, and several of these characters support this position. Key words: Stylidiaceae, Campanulales, floral development, systematic position, floral biology.



2007 ◽  
Vol 55 (1) ◽  
pp. 30 ◽  
Author(s):  
Sandra Luz Gómez-Acevedo ◽  
Susana Magallón ◽  
Lourdes Rico-Arce

The complete sequence of floral development in three species of Acacia was analysed. These species were sampled from each of the three Acacia subgenera. The species were Acacia berlandieri Benth. (subg. Aculeiferum), A. pennatula (Schltdl. & Cham.) Benth. (subg. Acacia) and A. saligna (Labill.) H.L.Wendl. (subg. Phyllodineae). The aim of the study was to determine whether the different subgenera share developmental pathways during flower formation. This study showed that development in the genus Acacia is heterogeneous. Each species studied showed different inception patterns of the calyx and androecium, whereas the inception patterns of the corolla and gynoecium were similar. These differences of inception in the calyx are not necessarily constant within each subgenus. Nevertheless, each subgenus was differentiated on the basis of inception patterns of the androecium, and other features such as the presence or absence of congenital or postgenital fusion in the calyx and corolla, and the time of differentiation of calyx and corolla tubes and the style.



1992 ◽  
Vol 70 (9) ◽  
pp. 1765-1776 ◽  
Author(s):  
L. P. Ronse Decraene ◽  
E. F. Smets

A study of the floral development of Dicentra formosa, Corydalis lutea, and Hypecoum procumbens was carried out to better understand the nature of the androecium in Fumariaceae. Sepals emerge successively in a median position and are followed by two alternating pairs of petals. Four stamen primordia are formed in a diagonal position. They are promptly followed by two lateral, slightly externally inserted primordia. In Dicentra and Corydalis the stamens arise on two crescent-shaped protuberances. In Hypecoum, four diagonal androecial primordia fuse into two median staminal complexes. The gynoecium emerges as a girdling primordium with four growth centers. Different interpretations of the androecium are discussed. It is demonstrated that the androecium in the Fumariaceae consists basically of two whorls: an outer whorl of four alternipetalous stamens and an inner whorl of two lateral stamens superposed to the outer petals. The monothecal nature of the alternipetalous stamens and the fusion of the stamens in two triplets is probably caused by a spatial median compression of the flower bud. The androecium of Hypecoum is the result of interprimordial growth between the pairs of monothecal stamens, and the androecium of Pteridophyllum arises through the loss of the two lateral stamens superposed to the outer petals. Key words: Fumariaceae, floral development, androecium, stamen whorls.



2020 ◽  
Vol 20 (1) ◽  
Author(s):  
Liling Yang ◽  
Shilian Qi ◽  
Arfa Touqeer ◽  
Haiyang Li ◽  
Xiaolan Zhang ◽  
...  

Abstract Background Flower development directly affects fruit production in tomato. Despite the framework mediated by ABC genes have been established in Arabidopsis, the spatiotemporal precision of floral development in tomato has not been well examined. Results Here, we analyzed a novel tomato stamenless like flower (slf) mutant in which the development of stamens and carpels is disturbed, with carpelloid structure formed in the third whorl and ectopic formation of floral and shoot apical meristem in the fourth whorl. Using bulked segregant analysis (BSA), we assigned the causal mutation to the gene Solanum lycopersicum GT11 (SlGT11) that encodes a transcription factor belonging to Trihelix gene family. SlGT11 is expressed in the early stages of the flower and the expression becomes more specific to the primordium position corresponding to stamens and carpels in later stages of the floral development. Further RNAi silencing of SlGT11 verifies the defective phenotypes of the slf mutant. The carpelloid stamen in slf mutant indicates that SlGT11 is required for B-function activity in the third whorl. The failed termination of floral meristem and the occurrence of floral reversion in slf indicate that part of the C-function requires SlGT11 activity in the fourth whorl. Furthermore, we find that at higher temperature, the defects of slf mutant are substantially enhanced, with petals transformed into sepals, all stamens disappeared, and the frequency of ectopic shoot/floral meristem in fourth whorl increased, indicating that SlGT11 functions in the development of the three inner floral whorls. Consistent with the observed phenotypes, it was found that B, C and an E-type MADS-box genes were in part down regulated in slf mutants. Conclusions Together with the spatiotemporal expression pattern, we suggest that SlGT11 functions in floral organ patterning and maintenance of floral determinacy in tomato.



Author(s):  
Daria Chevdar

The article is devoted to the specificity of translating scientific and technical discourse from English into Ukrainian in terms of communicative-functional and linguistic approaches. The translator’s goal is to produce a target text that would convey the same information as conveyed by the ST; the amount of the communicatively relevant information in the TT must be the same as in the ST. The results of the paper discover that the implementation of the strategy of communicatively equivalent translation determines the use of certain tactics that come down to a set of translation operations. Key words: scientific and technical discourse, communicative-functional approach, strategy of translation, tactic, operation.



Sign in / Sign up

Export Citation Format

Share Document