Growth rate effects on correlations among ring width, wood density, and mean tracheid length in Norway spruce (Picea abies)

1998 ◽  
Vol 28 (1) ◽  
pp. 56-68 ◽  
Author(s):  
Pierre Dutilleul ◽  
Marc Herman ◽  
Tomas Avella-Shaw
Keyword(s):  
Growth Rate ◽  
Picea Abies ◽  
Wood Density ◽  
Negative Correlation ◽  
Ring Width ◽  
Main Hypothesis ◽  
Growing Period ◽  
Tracheid Length ◽  
Rate Effects ◽  
Tree Approach

The main hypothesis tested in this paper is whether heavy thinnings affect the correlations among ring width, wood density, and mean tracheid length. Within-tree correlations were calculated between time series of yearly measurements. Among-tree correlations were computed (1) between averages over a growing period and (2) year by year. Correlations were analyzed on 20 fast-grown and 20 slow-grown Norway spruces (Picea abies (L.) Karst.) from an even-aged, plantation-grown stand near Rendeux, Belgian Ardennes. In the within-tree approach, fast-grown spruces showed a stronger negative correlation between ring width and fiber length. In among-tree approach 1, the widely held negative correlation between ring width and wood density vanished when the spruce growth rate was above 2.2 cm/year in circumference. Among-tree approach 2 demonstrated that the magnitude and sign of the correlations also depended on the year; a few years showed a significant correlation between ring width and wood density for the fast-grown Norway spruces, whereas the correlation was systematically negative and significant on many years for the slow-grown spruces. This study may explain part of the contradictory results reported in the literature concerning hypotheses similar to ours.

Intra-Ring and Inter-Ring Variations of Tracheid Length in Fast-Grown Versus Slow-Grown Norway Spruces (Picea Abies)

IAWA Journal ◽  
1998 ◽  
Vol 19 (1) ◽  
pp. 3-23 ◽  
Author(s):  
Marc Herman ◽  
Pierre Dutilleul ◽  
Tomas Avella-Shaw
Keyword(s):  
Growth Rate ◽  
Picea Abies ◽  
Frequency Distribution ◽  
Tree Growth ◽  
Slow Growth ◽  
Ring Width ◽  
Fast Growth ◽  
Tracheid Length ◽  
Tree Growth Rate ◽  
The Mean

Our study was conducted on 40 Norway spruces [Picea abies (L.) Karst.] from a stand located in the Belgian Ardennes. Twenty trees were randomly sampled from a slow-growth category, and twenty others from a fast -growth category. The hypothesis under testing is fourfold: increased tree growth rate may affect 1) the intra-ring weighted frequency distribution of tracheid length, 2) the inter-ring variation (from pith to bark) of the parameters describing this frequency distribution, 3) the interring variation of the mean tracheid length, and 4) the correlation between yearly mean tracheid length and yearly ring width.

scholarly journals Wood density and tracheid length of Aleppo pine (Pinus halepensis Mill.) grafts in relation to cambium age and growth rate

2018 ◽  
Vol 64 (No. 3) ◽  
pp. 101-107
Author(s):  
Foti Dafni ◽  
Passialis Costas ◽  
Voulgaridis Elias ◽  
Skaltsoyiannes Apostolos ◽  
Tsaktsira Maria
Keyword(s):  
Growth Rate ◽  
Wood Density ◽  
Juvenile Wood ◽  
Ring Width ◽  
Aleppo Pine ◽  
Age And Growth ◽  
Dry Density ◽  
Tracheid Length ◽  
Cambial Age ◽  
The Relationship

Wood density, tracheid length and growth rate were measured in Aleppo pine scions, 21–23 years old, and in Brutia pine rootstocks. In regard to the relationship between cambial age and dry density the results showed that the density increased with cambial age in both scions and rootstocks while the differences between Aleppo pine and Brutia pine were small. The relationship between cambial age and tracheid length showed an increase of tracheid length with cambial age. Differences between scions and rootstocks were small. From the last relationship it can be extracted that juvenile wood is produced in both scions and rootstocks although the Aleppo pine branches which were used for grafting were genetically matured. Between ring width and dry density and between ring width and tracheid length no statistical correlations were found either in scions or in rootstocks. The tracheid length in mature wood was higher than in juvenile wood. An increase of tracheid length with ring width was observed only in the case of juvenile wood.

Influence of Growth Rate and Cambial Age on Density of Rauli (Nothofagus nervosa) in Relation to Vessel Lumen Areas and Numbers

Holzforschung ◽  
1999 ◽  
Vol 53 (2) ◽  
pp. 199-203 ◽  
Author(s):  
M. P. Denne ◽  
C. M. Cahalan ◽  
D. P. Aebischer
Keyword(s):  
Growth Rate ◽  
Wood Density ◽  
Wood Quality ◽  
Ring Width ◽  
Substantial Effect ◽  
Present Sample ◽  
Vessel Lumen ◽  
Cambial Age ◽  
Old Trees ◽  
Nothofagus Nervosa

Summary To assess the likely effects of silvicultural treatment on the wood quality of Nothofagus nervosa grown in the UK, and the possibilities of independent selection within seed origins for density and growth rate, ring width and wood density were analysed from pith to bark of 19 trees. Variations in vessel lumen size, vessel number mm−2, and total lumen area mm−2 were analysed in ten trees. Since density increased by only 0.005g cm−3 per mm increase in ring width, silvicultural practices such as initial spacing and thinning are not likely to have a substantial effect on the wood density of rauli. Similarly, variation in density with cambial age was significant but relatively minor in the 40- and 60-year old trees of the present sample, suggesting that the rotation length is unlikely to have any practical influence on the density of rauli. Considerable between-tree differences in density were found which did not correspond to differences in ring width, suggesting it should be feasible to select independently for density and growth rate. These differences in density were associated with differences in both vessel lumen size and vessel number mm−2

Wood density of western hemlock: effect of ring width

1994 ◽  
Vol 24 (3) ◽  
pp. 638-641 ◽  
Author(s):  
Jeffrey D. DeBell ◽  
John C. Tappeiner II ◽  
Robert L. Krahmer
Keyword(s):  
Growth Rate ◽  
Wood Density ◽  
Growth Rates ◽  
Ring Width ◽  
Western Hemlock ◽  
X Ray ◽  
Breast Height ◽  
Latewood Density ◽  
Earlywood Width ◽  
Average Wood

Wood density of western hemlock (Tsugaheterophylla (Raf.) Sarg.) was determined by X-ray densitometry of strips from breast-height samples consisting of rings 20–24 from the pith. Ring parameters were averaged over the 5 years for each strip. Wood density was negatively correlated with radial growth rate. Average wood density dropped from 0.47 to 0.37 g/cm3 as average ring width increased from 2 to 8 mm. Wood density decreased at higher growth rates primarily because earlywood width increased while latewood width remained the same; as a result, percentage of latewood decreased. Earlywood density decreased slightly at higher growth rates, but latewood density was not significantly related to growth rate.

Effect of ring width, cambial age, and climatic variables on the within-ring wood density profile of Norway spruce Picea abies (L.) Karst.

Trees ◽  
2013 ◽  
Vol 27 (4) ◽  
pp. 913-925 ◽  
Author(s):  
Tony Franceschini ◽  
Fleur Longuetaud ◽  
Jean-Daniel Bontemps ◽  
Olivier Bouriaud ◽  
Benoît-Damien Caritey ◽  
...  
Keyword(s):  
Picea Abies ◽  
Norway Spruce ◽  
Wood Density ◽  
Density Profile ◽  
Ring Width ◽  
Climatic Variables ◽  
Cambial Age

Growth Rate Effects on Intra-Ring and Inter-Ring Trajectories of Microfibril Angle in Norway Spruce (Picea Abies)

IAWA Journal ◽  
1999 ◽  
Vol 20 (1) ◽  
pp. 3-21 ◽  
Author(s):  
Marc Herman ◽  
Pierre Dutilleul ◽  
Tomas Avella-Shaw
Keyword(s):  
Growth Rate ◽  
Picea Abies ◽  
Repeated Measures ◽  
Microfibril Angle ◽  
Solid Wood ◽  
Growth Rings ◽  
Repeated Measures Design ◽  
Rate Effects ◽  
The Subject ◽  
Tree Growth Rate

Fourteen Norway spruces [Picea abies (L.) Karst.], randomly sampled as 7 fast-grown and 7 slow-grown trees, were used to test whether an increased tree growth rate in circumference affects the intra-ring and inter-ring trajectories of the microfibril angle in the S2 layer of the tracheid wall. Those trajectories describe the fluctuations of the S2 microfibril angle, respectively, from earlywood to latewood within rings and from pith to bark among rings. Using the cross-field pit apertures, intra-ring measures of microfibril angle were made at 11 equally-spaced sampling sites over each of 8 growth rings, following an 11 × 8 doubly repeated measures design with the tree as the 'subject' on which repeated measures were made. All the intra-ring trajectories of microfibril angle decreased linearly from earlywood to latewood, whereas the inter-ring trajectories showed significant year effects. Both types of trajectories are significantly affected by the growth rate after first thinning, as the fast-grown spruces showed a systematically larger microfibril angle and a mean microfibril angle of 290 compared to 21 0 for the slow-grown spruces. Thus, lower tensile and tear strengths of tracheids as well as some modifications of the mechanical properties of solid wood and paper can be expected from Norway spruces growing faster than 2.2 cm/year in circumference.

Effects of timing and intensity of thinning on wood structure and chemistry in Norway spruce

Holzforschung ◽  
2010 ◽  
Vol 64 (1) ◽  
Author(s):  
Tuula Jyske ◽  
Seija Kaakinen ◽  
Urban Nilsson ◽  
Pekka Saranpää ◽  
Elina Vapaavuori
Keyword(s):  
Growth Rate ◽  
Norway Spruce ◽  
Wood Density ◽  
Ring Width ◽  
Basal Area ◽  
Wood Structure ◽  
Wood Chemistry ◽  
Thinning Intensity ◽  
Thinning Operations ◽  
Treatment Onset

AbstractThe effects of thinning intensity on wood structure and chemistry of Norway spruce [Picea abies(L.) Karst.] have been studied in a long-term thinning experiment established in the 1960s in south-western Sweden. The experiment comprised datasets of unthinned control trees and trees of four thinning treatments, in which thinning operations of different intensities were carried out. These thinning operations were applied either once, three, or five times over the course of the stand rotation. Thinning intensities varied from 0% to 60% based on the stand basal area (BA) before thinning: (A) 5×20% (i.e., 20% of the BA removed five times); (B) 3×40% (i.e., 40% of the BA removed three times); (C) 1×60% (i.e., 60% of the BA removed once); and (D) 5×40% (i.e., 40% of the BA removed five times). Altogether 60 trees that were 67 years old were sampled 38 years after the treatment onset. Radial growth rate, wood density, tracheid properties, and the chemical composition of wood were studied in detail. All four thinning treatments increased growth rate and decreased wood density; however, treatment D showed the greatest change. Single, heavy thinning in treatment C resulted in unfavourable intra-stem variation in ring width and wood density. No significant differences in tracheid properties and wood chemistry were found between the treatments. We conclude that only a remarkable increase in growth rate would induce detrimental changes in tracheid properties and wood chemistry in the context of thinning.

The Genetic Control of Ring Width, Wood Density and Tracheid Length in the Juvenile Core ofPinus patula

1994 ◽  
Vol 169 (1) ◽  
pp. 15-20
Author(s):  
R. D. Barnes ◽  
J. S. Birks ◽  
G. Battle ◽  
L. J. Mullin
Keyword(s):  
Genetic Control ◽  
Wood Density ◽  
Ring Width ◽  
Tracheid Length

Wood density of young-growth western hemlock: relation to ring age, radial growth, stand density, and site quality

2004 ◽  
Vol 34 (12) ◽  
pp. 2433-2442 ◽  
Author(s):  
Dean S DeBell ◽  
Ryan Singleton ◽  
Barbara L Gartner ◽  
David D Marshall
Keyword(s):  
Growth Rate ◽  
Wood Density ◽  
Ring Width ◽  
Stand Density ◽  
Age And Growth ◽  
Western Hemlock ◽  
Site Class ◽  
Ring Density ◽  
Wide Range ◽  
Young Growth

Breast-high stem sections were sampled from 56 western hemlock (Tsuga heterophylla (Raf.) Sarg.) trees growing in 15 plots representing a wide range of tree and site conditions in northwestern Oregon. Growth and wood density traits of individual rings were measured via X-ray densitometry, and relationships of ring density and its components to age and growth rate were analyzed. Ring density was highest (0.49 g/cm3) near the pith, declined to 0.40 g/cm3 at age 10, remained stable to about age 25, and then increased gradually and remained between 0.43 and 0.44 g/cm3 from age 38 to 45 and beyond. A negative influence of rapid growth on whole ring density was greatest at young ages and diminished with time, becoming nonsignificant beyond age 30. Earlywood density, latewood density, and latewood proportion were all negatively related to ring width at young ages, but by age 21-25, latewood proportion was the only component of ring density that remained significantly diminished by increased growth rate. Residual differences in wood density (after age and growth rate were considered) did not appear to be related to either stand density or site class. Overall, young-growth hemlock trees are relatively uniform in wood density and likely to be more so if grown in intensively managed stands.

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