Intra- and inter-specific competition between Barrow's goldeneye (Bucephala islandica) and bufflehead (Bucephala albeola)
Barrow's goldeneye and bufflehead pairs defend well-delineated territories from which they exclude conspecifics. Only the male defends the territory and does so even when the female is absent. However, after the young have hatched, females of both species defend brood territories from which they exclude conspecifics. Both species are also interspecifically territorial. Barrow's goldeneye and bufflehead drakes threatened and (or) attacked 15 and 11 species, respectively. However, 55% of Barrow's goldeneye interspecific encounters were with bufflehead and 52% of bufflehead encounters were with Barrow's goldeneye. Barrow's goldeneye drakes always excluded bufflehead from their territory whereas bufflehead drakes successfully excluded yearlings, females, and unpaired Barrow's goldeneye drakes but were usually dominated by paired males. Female Barrow's goldeneye excluded all bufflehead from the brood territory and were particularly aggressive toward bufflehead broods. Food and mates are the most likely defended resources in intraspecific encounters. In interspecific encounters diving ducks were more violently attacked than dabbling ducks indicating that food may be the defended resource. Interspecific aggression of Barrow's goldeneye toward bufflehead does not appear to be misdirected intraspecific aggression but rather a directed aggression toward a food competitor and may therefore be adaptive.