scholarly journals Desmotheca (Orthotrichaceae): Gondwanan fragmentation and the origin of a Southeast Asian genus

1990 ◽  
Vol 3 (1) ◽  
pp. 79-88 ◽  
Author(s):  
Dale H. Vitt

Desmotheca Lindberg was proposed in 1872 as a replacement name for the illegitimate Cryptocarpon Dozy & Molk. (1844) and Cryptocarpus Dozy & Molk. (1846). Seven names have been placed in these genera, from which two species appear to be taxonomically valid. Desmotheca apiculata (Dozy & Molk.) Card. occurs from New Caledonia and New Guinea west to Indonesia and the Andaman Islands, north to Burma, Thailand, Vietnam and Mindanao in the Philippines, while D. brachiata (Hook. & Wils.) Vitt comb. nova is restricted to the island of Luzon in the Philippines. The evolution and dispersal of this genus can be related to the northward movement of Gondwanan island blocks during the Cretaceous.

REINWARDTIA ◽  
2020 ◽  
Vol 19 (1) ◽  
pp. 1-25
Author(s):  
Ruth Kiew

KIEW, R. 2020. Towards a Flora of New Guinea: Oleaceae. Part 1. Jasminum, Ligustrum, Myxopyrum and Olea. Reinwardtia 19(1): 1‒25. ‒‒ Oleaceae in New Guinea is represented by five genera and about 32 species, namely Chionanthus (about 16 species), Jasminum (10 species), Ligustrum (3 species), Myxopyrum (2 species) and Olea (1 species). A key to genera as well as descriptions of and keys to species of Jasminum, Ligustrum, Myxopyrum and Olea are provided. Of the three Ligustrum species, L. glomeratum is widespread throughout Malesia, L. novoguineense is endemic and L. parvifolium Kiew is a new endemic species. Six species of Jasminum are endemic (J. domatiigerum, J. gilgianum, J. magnificum, J. papuasicum, J. pipolyi and J. rupestre). Jasminum turneri just reaches the northern tip of Australia; of the two species from the Pacific Islands J. simplicifolium subsp. australiense just reaches SE Papua New Guinea and J. didymum, a coastal species, reaches into Malesia as far north as E Java; J. elongatum is widespread from Asia to Australia. Neither Myxopyrum species is endemic: M. nervosum subsp. nervosum extends from Peninsular Malaysia to Indonesian New Guinea, and M. ovatum from the Philippines to the Admiralty Islands. The sole species of Olea, O. paniculata, stretches from Java to Australia and New Caledonia. 


1991 ◽  
Vol 4 (1) ◽  
pp. 151 ◽  
Author(s):  
Andersen N Moller

More than 120 species of marine water striders (Hemiptera, Gerromorpha), representing three families and eight genera, are distributed throughout the lndo-Pacific region. They live in marine habitats such as mangroves, intertidal coral reef flats and the sea surface near coral and rocky coasts. Five species of sea skaters, Halobates (Gerridae), have colonised the surface of the open ocean. Adult marine water striders are wingless but may disperse along coasts, chains of islands and possibly across wider stretches of open sea. Although some species of coral bugs, Halovelia (Veliidae) and Halobates are widespread, most species of marine water striders have rather restricted distributions. Cladistic hypotheses are now available for the genera Halovelia, Xenobates (Veliidae) and Halobates. Based upon distributional data for about 110 species, a number of areas of endemism can be delimited within the Indo-Pacific region. The results of component analyses of taxon-area cladograms for several monophyletic species-groups of marine water striders are presented. The faunas of northern New Guinea, the Bismarck and Solomon Islands (Papuasia) are closely related and show much greater affinity with Maluku, Sulawesi and the Philippines than with the fauna of northern Australia. Relationships between the faunas of Papuasia + Sulawesi + the Philippines and those of Borneo + Jawa + Malaya are relatively weak. Marine water striders endemic to islands of the western Pacific show relationships among themselves and with Australia. Most marine water striders from the Indian Ocean (East Africa, Madagascar, Mauritius, Seychelles and Maldives) can be derived from the Indian-South-east Asian fauna. Composite faunas of marine water striders (either of different age or origin) are found in New Guinea, New Caledonia, Fiji Islands, the Philippines, tropical Australia and East Africa. The biogeography of marine water striders does not support the traditional division of the Indo- Pacific into the Ethiopian, Oriental and Australian regions. The distributional patterns are more compatible with a set of hierarchical relationships between more restricted areas of endemism.


2012 ◽  
Vol 25 (6) ◽  
pp. 390 ◽  
Author(s):  
Gillian K. Brown ◽  
Daniel J. Murphy ◽  
James Kidman ◽  
Pauline Y. Ladiges

Acacia sensu stricto is found predominantly in Australia; however, there are 18 phyllodinous taxa that occur naturally outside Australia, north from New Guinea to Indonesia, Taiwan, the Philippines, south-western Pacific (New Caledonia to Samoa), northern Pacific (Hawaii) and Indian Ocean (Mascarene Islands). Our aim was to determine the phylogenetic position of these species within Acacia, to infer their biogeographic history. To an existing molecular dataset of 109 taxa of Acacia, we added 51 new accessions sequenced for the ITS and ETS regions of nuclear rDNA, including samples from 15 extra-Australian taxa. Data were analysed using both maximum parsimony and Bayesian methods. The phylogenetic positions of the extra-Australian taxa sampled revealed four geographic connections. Connection A, i.e. northern Australia?South-east Asia?south-western Pacific, is shown by an early diverging clade in section Plurinerves, which relates A. confusa from Taiwan and the Philippines (possibly Fiji) to A. simplex from Fiji and Samoa. That clade is related to A. simsii from southern New Guinea and northern Australia and other northern Australian species. Two related clades in section Juliflorae show a repeated connection (B), i.e. northern Australia?southern New Guinea?south-western Pacific. One of these is the ?A. auriculiformis clade', which includes A. spirorbis subsp. spirorbis from New Caledonia and the Loyalty Islands as sister to the Queensland species A. auriculiformis; related taxa include A. mangium, A. leptocarpa and A. spirorbis subsp. solandri. The ?A. aulacocarpa clade' includes A. aulacocarpa, A. peregrinalis endemic to New Guinea, A. crassicarpa from New Guinea and Australia, and other Australian species. Acacia spirorbis (syn. A. solandri subsp. kajewskii) from Vanuatu (Melanesia) is related to these two clades but its exact position is equivocal. The third biogeographic connection (C) is Australia?Timor?Flores, represented independently by the widespread taxon A. oraria (section Plurinerves) found on Flores and Timor and in north-eastern Queensland, and the Wetar island endemic A. wetarensis (Juliflorae). The fourth biogeographic connection (D), i.e. Hawaii?Mascarene?eastern Australia, reveals an extreme disjunct distribution, consisting of the Hawaiian koa (A. koa, A. koaia and A. kaoaiensis), sister to the Mascarene (R�union Island) species A. heterophylla; this clade is sister to the eastern Australian A. melanoxylon and A. implexa (all section Plurinerves), and sequence divergence between taxa is very low. Historical range expansion of acacias is inferred to have occurred several times from an Australian?southern New Guinean source. Dispersal would have been possible as the Australian land mass approached South-east Asia, and during times when sea levels were low, from the Late Miocene or Early Pliocene. The close genetic relationship of species separated by vast distances, from the Indian Ocean to the Pacific, is best explained by dispersal by Austronesians, early Homo sapiens migrants from Asia.


Author(s):  
David R. Smith

Die Schwertwespen (Xiphydriidae) der Philippinen, der Malaysischen Inseln, Indonesiens, Papua-Neuguineas, Neukaledoniens und der Fiji-Inseln werden revidiert. Sechsundzwanzig Arten aus sechs Gattungen werden beschrieben, abgebildet, und für diese ein Bestimmungsschlüssel erstellt. Zwei Arten werden von den Philippinen nachgewiesen, drei von den Malaysischen Inseln, acht aus Indonesien, neun von Papua-Neuguinea, vier aus Neukaledonien und eine von den Fiji-Inseln. Die behandelten Taxa sind: Lissoxiphyda, n. gen., L. kiunga, n. sp. (Papua-Neuguinea), L. lucida, n. sp. (Papua-Neuguinea), L. morobe, n. sp. (Papua-Neuguinea), L. rufipes (F. Smith, 1859), n. comb. (Indonesien: Aru), L. shinoharai, n. sp. (Indonesien: Sulawesi), L. simbai, n. sp. (Papua-Neuguinea), L. tripotini, n. sp. (Neukaledonien); Calexiphyda, n. gen., C. blanki, n. sp. (Neukaledonien), C. caledonia, n. sp. (Neukaledonien), C. crocea, n. sp. (Neukaledonien); Indoxiphia Maa, 1949, I. darlingi, n. sp. (Indonesien: Kalimantan), I. falcata, n. sp. (Papua-Neuguinea: Neuirland), I. fijiensis, n. sp. (Fiji), I. laeviceps (F. Smith, 1861) (Indonesien: Ambon, Buru; Papua-Neuguinea), I. maai, n. sp. (Papua-Neuguinea), I. papuaensis, n. sp. (Papua-Neuguinea), I. quadricincta (Benson, 1935) (Indonesien: Java), I. schiffi, n. sp. (Papua-Neuguinea), I. sumatra, n. sp. (Indonesien: Sumatra); Obesaxiphyda, n. gen., O. banahao, n. sp. (Philippinen), O. borneensis (Rohwer, 1921), n. comb. (Malaysia: Sabah); Hyperxiphia Maa, 1949, H. cyanea (Mocsáry, 1891) (Indonesien: Java), H. flavicornis (Rohwer, 1921), n. comb. (Malaysia: Sabah), H. sulawesi, n. sp. (Indonesien: Sulawesi); Lataxiphyda, n. gen., L. erythropus (Cameron, 1903), n. comb. (Malaysia: Sarawak), L. paraerythropus, n. sp. (Malaysia: Selangor), L. pyrura (Rohwer, 1921), n. comb. (Philippinen). Zwei nicht genauer deutbare Männchen, für die bereits Namen existieren, werden den folgenden Gattungen zugeordnet: Calexiphyda novacaledonica (Jennings und Austin, 2007), n. comb., und Indoxiphia testacea (Mocsáry, 1900), n. comb. Cingalixiphia Maa, 1949, ist ein neues Synonym von Indoxiphia Maa, 1949, und Palpixiphia Maa, 1949, ein neues Synonym von Hyperxiphia Maa, 1949. Hyperxiphia formosana (Enslin, 1911) und Obesaxiphyda heritierae (Rohwer, 1921) sind neue Kombinationen.StichwörterSymphyta, Xiphydriidae, woodwasps, southeastern Asia, Indo-Australia, Philippines, insular Malaysia, Indonesia, Papua New Guinea, New Caledonia, Fiji.Nomenklatorische Handlungenblanki Smith, 2008 (Calexiphyda), spec. n.caledonica Smith, 2008 (Calexiphyda), spec. n.crocea Smith, 2008 (Calexiphyda), spec. n.novacaledonica (Jennings & Austin, 2007) (Calexiphyda), comb. n. hitherto ??flavicornis (Rohwer, 1921) (Hyperxiphia), comb. n. hitherto Xiphydria flavicornissulawesi Smith, 2008 (Hyperxiphia), spec. n.Cingalixiphia Maa, 1949 (Indoxiphia), syn. n. of Indoxiphia Maa, 1949; hitherto a subgenus of this genusdarlingi Smith, 2008 (Indoxiphia), spec. n.falcata Smith, 2008 (Indoxiphia), spec. n.fijiensis Smith, 2008 (Indoxiphia), spec. n.laeviceps F. Smith, 1861 (Indoxiphia), Lectotype; comb. n. hitherto Xiphidria laevicepsmaai Smith, 2008 (Indoxiphia), spec. n.papuaensis Smith, 2008 (Indoxiphia), spec. n.schiffi Smith, 2008 (Indoxiphia), spec. n.sumatra Smith, 2008 (Indoxiphia), spec. n.testacea (Mocsáry, 1900) (Indoxiphia), comb. n. hitherto Xiphydria testaceaerythropus (Cameron, 1903) (Lataxiphyda), Lectotype; comb. n. hitherto Xiphydria eryrthropusparaerythropus Smith, 2008 (Lataxiphyda), spec. n.pyrura (Rohwer, 1921) (Lataxiphyda), comb. n. hitherto Xiphydria pyrurakiunga Smith, 2008 (Lissoxiphyda), spec. n.lucida Smith, 2008 (Lissoxiphyda), spec. n.morobe Smith, 2008 (Lissoxiphyda), spec. n.rufipes (F. Smith, 1859) (Lissoxiphyda), comb. n. hitherto Xiphidria rufipesshinoharai Smith, 2008 (Lissoxiphyda), spec. n.simbai Smith, 2008 (Lissoxiphyda), spec. n.tripotini Smith, 2008 (Lissoxiphyda), spec. n.banahao Smith, 2008 (Obesaxiphyda), spec. n.borneensis (Rohwer, 1921) (Obesaxiphyda), comb. n. hitherto Xiphydria heritierae borneensisheritierae (Rohwer, 1921) (Obesaxiphyda), comb. n. hitherto Xiphydria heritieraemelanopus Cameron, 1903 (Xiphydria), Lectotype now a synonym of Lataxiphyda erythropus (Cameron, 1903)Calexiphyda Smith, 2008 (Xiphydriidae), gen. n.Lataxiphyda Smith, 2008 (Xiphydriidae), gen. n.Lissoxiphyda Smith, 2008 (Xiphydriidae), gen. n.Obesaxiphyda Smith, 2008 (Xiphydriidae), gen. n.


Zootaxa ◽  
2012 ◽  
Vol 3294 (1) ◽  
pp. 1 ◽  
Author(s):  
OFER GON ◽  
GERALD R. ALLEN

The Indo-Pacific apogonid genus Siphamia Weber 1909 is unique among cardinalfishes in having a bacterial biolumines-cent system and spinoid scales. Light is produced by luminous bacteria found in a small pocket connected to the gut in theabdominal cavity and in a sac on each side of the tip of the tongue. Siphamia consists of 23 small species many of whichare associated with invertebrates such as sea urchins, crown-of-thorns starfish and coral. Species of this genus fall intotwo main groups with different dark pigment pattern of the longitudinal translucent muscle acting as a light organ thatdiffuses light along the ventral edge of the body. The S. tubifer group, with a striated light organ, includes S. arabica, newspecies, from the Gulf of Oman; S. argentea from the Philippines and northern Western Australia; S. fraseri, new species,from New Caledonia, Tonga and Fiji; S. fuscolineata from the Marshall and Line islands; S. goreni, new species, from thesouthern Red Sea; S. guttulata from Darnley Island, Queensland; S. jebbi from the western Pacific, ranging from the Phil-ippines to Western Australia and east to the Caroline Islands, Fiji, and Tonga; S. majimai from the Ryukyu and Ogasawaraislands to northwestern Australia, ranging eastward to New Caledonia and Tonga; S. mossambica from the western IndianOcean; S. randalli, new species, from the Society and Cook islands; S. spinicola, new species, from Biak in eastern Indo-nesia, Papua New Guinea, Woleai Atoll, Vanuatu, New Caledonia and the Chesterfield Islands; S. stenotes, new species,from the Triton Bay area of Irian Jaya Barat Province of Indonesia; and S. tubifer ranging widely in the Indo-West Pacificfrom the Red Sea to Madagascar and east to Vanuatu. The S. tubulata group, with a dark-dotted light organ, includes S.brevilux, new species, from Papua New Guinea; S. cephalotes from southern Australia; S. corallicola from Indonesia, Sa-bah, and Timor Sea; S. cuneiceps from Western Australia and the east coast of Queensland; S. cyanophthalma, new species,from the Philippines, Palau, Indonesia, and Papua New Guinea; S. elongata from the Philippines and Brunei; S. fistulosafrom Java, Sumbawa and Komodo, Indonesia, and Brunei; S. roseigaster from Western Australia, ranging along the north-ern and eastern coast of Australia south to Sydney Harbour, New South Wales; S. senoui, new species, from the RyukyuIslands, Japan; and S. tubulata from the Papua Barat Province, Indonesia, south coast of Papua New Guinea, northern Western Australia and Queensland.


Zootaxa ◽  
2011 ◽  
Vol 3046 (1) ◽  
pp. 1 ◽  
Author(s):  
PETER JÄGER

The genera Nilus O. Pickard-Cambridge 1876, Sphedanus Thorell 1877 and Dendrolycosa Doleschall 1859 are revised. Nilus is recognised as junior synonym of Thalassius Simon 1885; accordingly all species previously placed in Thalassius including nomina dubia are transferred to Nilus. Nilus curtus O. Pickard-Cambridge 1876 is recognised as senior synonym of Thalassius spinosissimus Karsch 1879. Nilus decoratus (Patel & Reddy) comb. nov. from India is transferred from Pisaura Simon 1885. Nilus amazonicus Simon 1898 from Brazil is transferred to the genus Architis Simon 1898: Architis amazonica (Simon 1898) comb. nov. Nilus oblongus Pavesi 1897 from Ethiopia and N. marginatus (Simon 1888) from the Andaman Islands are considered nomina dubia. Sphedanus Thorell 1877 is removed from the synonymy with Nilus and recognised as a senior synonym of Eurychoera Thorell 1897. The type species, S. undatus Thorell 1877, is redecribed and its copulatory organs are illustrated for the first time. Two species previously placed in the genus Eurychoera are transferred: Sphedanus banna (Zhang, Zhu & Song 2004) comb. nov. and Sphedanus quadrimaculatus (Thorell 1897) comb. nov. Campostichommides Strand 1911 and Dianpisaura Zhang, Zhu & Song 2004 are recognised as junior synonyms of Dendrolycosa. Nilus lanceolatus Simon 1898 and Pisaura lizhii Zhang 2000 are proposed as junior synonyms of Dendrolycosa robusta Thorell 1895. Four species are described as new: D. bairdi spec. nov. from Laos (male, female), D. duckitti spec. nov. from Laos (male, female), D. sierwaldae spec. nov. from New Guinea (female) and D. yuka spec. nov. from Australia (female). Eight species previously placed in the genera Nilus, Campostichommides Strand 1911, Pisaura Simon 1885 and Dianpisaura Zhang, Zhu & Song 2004, respectively are transferred to Dendrolycosa: D. ornata (Berland 1924) comb. nov. from New Caledonia, D. spadicaria (Simon 1897) comb. nov. from India (nomen dubium), D. bobbiliensis (Reddy & Patel 1993) comb. nov. from India, D. gitae (Tikader 1970) comb. nov. from India, D. inquirenda (Strand 1911) comb. nov. nomen dubium from Indonesia, D. parangbusta (Barrion & Litsinger 1995) comb. nov. from Philippines, D. putiana (Barrion & Litsinger 1995) comb. nov. from Philippines, D. songi (Zhang 2000) comb. nov. from Yunnan, China. Dendrolycosa stauntoni Pocock 1900 is considered a nomen dubium. One species originally described in Dendrolycosa is transferred to the genus Hygropoda: H. gracilis (Thorell 1891) comb. nov. Two species previously placed in the genus Tinus F. O. Pickard-Cambridge 1901 are transferred to Hygropoda: H. sikkimus (Tikader 1970) comb. nov., H. chandrakantii (Reddy & Patel 1993) comb. nov.


2016 ◽  
Vol 48 (1) ◽  
pp. 13-52 ◽  
Author(s):  
Arve ELVEBAKK ◽  
Soon Gyu HONG ◽  
Chae Haeng PARK ◽  
Eli Helene ROBERTSEN ◽  
Per Magnus JØRGENSEN

AbstractReports of ‘Psoroma sphinctrinum’ from Palaeotropical areas are shown to represent instead species of the genus Gibbosporina, which is described here as new to science. This genus is superficially similar to tripartite, austral Pannaria species, such as the species now referred to as Pannaria sphinctrina (Mont.) Tuck. ex Hue. A phylogram based on an analysis of the nuclear large subunit rDNA (LSU) locus shows that Gibbosporina is instead a clade in a Pannariaceae branch referred to as the ‘Physma group’, a most unexpected addition to Pannariaceae dealt with by several previous studies. Genera assigned to this group have very contrasting general appearances. However, this diverse group shares distinctly ring-like thalline excipular margins; strongly amyloid internal ascus structures; well-developed perispores which have irregular gibbae and/or nodulose or acuminate apical extensions, but not verrucae; lacks TLC-detectable secondary compounds and have tropical distributions. Gibbosporina is the only tripartite genus in the group, with distinct, nodulose, placodioid, mini-fruticose to mini-foliose cephalodia with a high diversity of Nostoc cyanobionts. The cyanomorphs can apparently exist independently in some cases, although the apothecia on such cephalodia on a specimen from Réunion were unexpectedly found to belong to the chloromorph. The genus and related genera forming the ‘Physma group’ are probably evolutionarily old, and their weak affinity to the remaining part of Pannariaceae, concentrated in the Southern Hemisphere, is discussed. The genus includes 13 known species, and the generitype is Gibbosporina boninensis from the Japanese Ogasawara Islands, originally described as Psoroma boninense and recombined here. The following 12 species are described here as new to science, seven of them with molecular support in an LSU and ITS-based phylogram: Gibbosporina acuminata (Australia, the Philippines), G. amphorella (New Caledonia), G. bifrons (Malaysia, New Caledonia, the Philippines, Solomon Islands), G. didyma (Mauritius, Réunion), G. elixii (Australia), G. leptospora (Australia, Papua New Guinea), G. nitida (Australia, Papua New Guinea, the Philippines), G. mascarena (Mauritius, Réunion, Sri Lanka), G. papillospora (the Philippines), G. phyllidiata (Solomon Islands), G. sphaerospora (Australia, Indonesia, Malaysia, the Philippines, Samoa, and with Psoroma sphinctrinum var. endoxanthellum as a new synonym), and G. thamnophora (Australia and the Philippines). Except for the phyllidiate G. phyllidiata and for G. thamnophora which has cephalodia adapted for vegetative propagation, the species are all primarily fertile. A key for determining the species is provided.


Zootaxa ◽  
2021 ◽  
Vol 5081 (4) ◽  
pp. 505-523
Author(s):  
ALICE WELLS ◽  
DAVID CARTWRIGHT

Several collections of adults of the caddisfly order Trichoptera were studied from Timor-Leste, the nation-state comprising the eastern region of the island of Timor. The specimens represent ten families: Hydrobiosidae (2 species), Glossosomatidae (1 species), Hydroptilidae (3 species), Philopotamidae (5 species), Hydropsychidae (3 species), Polycentropodidae (1 species), Psychomyiidae (3 species), Xiphocentronidae (1 species), Lepidostomatidae (1 species), Leptoceridae (3 species). Among the 24 species listed, 16 were identified as established Southeast Asian species. Among these are two very widespread species, one extending further east to New Guinea, northern Australia, and New Caledonia and another that was described from Fiji. An additional seven species are newly described here: Ulmerochorema hatubuilico sp. nov., Hydroptila bellisi sp. nov. and H. aileuensis sp. nov., Chimarra lawaliu sp. nov., C. multidentata sp. nov., C. sameana sp. nov. and C. timorensis sp. nov. Hitherto, the genus Ulmerochorema Mosely was believed to be an Australian endemic. A xiphocentronid specimen could be identified to genus Drepanocentron only.  


Zootaxa ◽  
2020 ◽  
Vol 4747 (2) ◽  
pp. 275-301
Author(s):  
WOLFGANG DENZER ◽  
PATRICK D. CAMPBELL ◽  
ULRICH MANTHEY ◽  
ANDREA GLÄSSER-TROBISCH ◽  
ANDRÉ KOCH

Currently three different species are recognized within the Southeast Asian agamid genus Hydrosaurus: H. amboinensis (Schlosser, 1768) from Ambon, Seram, Sulawesi and New Guinea, H. pustulatus (Eschscholtz, 1829) from the Philippines and H. weberi Barbour, 1911 from Halmahera and adjacent islands. Historically, two additional species were described from the island of Sulawesi, but were synonymized with H. amboinensis more than a century ago and have been treated as such in most subsequent publications. In order to revise the taxonomy and diversity of these enigmatic agamid lizards, we examined the corresponding type specimens and additional material originating from Sulawesi and compared them to photographs of live specimens from field trips. Due to differences in colour pattern and scalation characters, we resurrect the taxa celebensis Peters, 1872 and microlophus Bleeker, 1860 from the synonymy of H. amboinensis, which in turn is restricted to the central Moluccas and New Guinea. Hence, Sulawesi is currently the only known island within the genus’ range to be inhabited by two different species of sailfin lizards. Our systematic investigation brings the number of recognized species within the genus Hydrosaurus to five. 


1990 ◽  
Vol 4 (5) ◽  
pp. 973 ◽  
Author(s):  
DA Duckhouse

Brunettia Annandale (sensu Duckhouse 1966) has previously been known in Australopapua from thirty-three species, comprising twenty-eight from Papua-New Guinea, but only five from Australia, all in the southern states. This anomaly is now removed with the description of seventeen new species from Queensland and the Northern Territory, showing that the major evolutionary centre extends from Papua-New Guinea far into northern Australia, and that the southern species are not in reality separated by a disjunction. Three new species are also described from southern Australia, two from New Guinea, and two from New Caledonia (the first from this island). The phylogenetic importance of Brunettia is especially due to the inclusion in it of taxa that are cladistically basal and annectant, nearly all Australopapuan. The mix of tribal, generic, subgeneric and species characters found in earlier descriptions is ordered into a strict hierarchical sequence, and Brunettia is divided into seven subgenera — Brunettia, s. str., Plesiobrunettia, subg. nov., Atrichobrunettia Satchell, Maurobrunettia, subg. nov., Campanulobrunettia, subg. nov., Horobrunettia, subg. nov., and Mrrousiella Vaillant, stat. nov., this last resurrected from synonymy with Atrichobrunettia. Of these, Maurobrunettia occurs in northern Australia, Plesiobrunettia is New Guinean, Campanulobrunettia and Atrrchobrunettia are Australopapuan, and Horobrunettia is mainly Australopapuan but has one species in the Philippines. Brunettia s. str. is more widely distributed, but extensively diversified in Papua-New Guinea, and Mirousiella is European. The ten Papua-New Guinean species placed by Quate & Quate (1967) in Atrichobrunettia are transferred into the various subgenera of Brunettia (combs. nov.), and their Brunettia species are also assorted into these subgenera. New keys are provided covering all Australopapuan Brunettia species. The genealogical status of Mormiini and Maruinini are discussed. It is concluded that because Mormiini are an offshoot of the Maruinini, the Maruinini are paraphyletic, but that this defect cannot be overcome until more is known of maruinine phylogeny.


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