Review of Gonatocerus (Hymenoptera: Mymaridae) in the Neotropical region, with description of eleven new species

Zootaxa ◽  
2010 ◽  
Vol 2456 (1) ◽  
pp. 1 ◽  
Author(s):  
SERGUEI V. TRIAPITSYN ◽  
JOHN T. HUBER ◽  
GUILLERMO A. LOGARZO ◽  
VLADIMIR V. BEREZOVSKIY ◽  
DANIEL A. AQUINO
Keyword(s):  
New Species ◽  
Costa Rica ◽  
Dominican Republic ◽  
Neotropical Region ◽  
Species Group ◽  
The Other ◽  
Valid Species ◽  
Neotropical Species ◽  
Distribution Ranges ◽  
Species Groups

The described Neotropical species of the common and speciose fairyfly genus Gonatocerus Nees ab Esenbeck (Hymenoptera: Mymaridae) are reviewed and re-diagnosed. Eighty-four valid species are recognized including 11 newly described ones. Illustrated identification keys are provided to identify the 5 subgenera recognized in Gonatocerus and to females of each subgenus. The keys include 79 of the treated species, the other five species are known from males only. The known distribution ranges of the species, including new records of extralimital specimens examined, are given. Host associations of the treated species are also given, with emphasis on those species that parasitize eggs of proconiine sharpshooters (Hemiptera: Cicadellidae: Cicadellinae: Proconiini), all of which belong to the ater and morrilli subgroups of the ater species group of G. (Cosmocomoidea Howard), stat. rev. The other four recognized subgenera within Gonatocerus are the nominate subgenus, G. (Gonatocerus) (= sulphuripes species group of authors), G. (Lymaenon Walker), stat. rev. (= litoralis and straeleni species groups of authors), G. (Gastrogonatocerus Ogloblin), stat. rev. (= membraciphagus species group of authors), and G. (Gahanopsis Ogloblin), syn. n. & stat. n. (= deficiens species group of authors). Newly treated as synonyms of G. (Lymaenon) are Rachistus Foerster, Oophilus Enock, Agonatocerus Girault, Gonatoceroides Girault, and Decarthrius Debauche, all syn. n. Two species groups are recognized within G. (Cosmocomoidea) in the New World: the ater group, to which most Neotropical species belong (with the ater, bucculentus, morrilli, and the newly defined chusqueicolus subgroups) and the masneri group, with two described species from the Dominican Republic. The 11 new species described are G. (Gahanopsis) arkadak Triapitsyn sp. n. (Colombia), G. (Cosmocomoidea) barbos Triapitsyn sp. n. (Costa Rica and Mexico), G. (Cosmocomoidea) blefuscu Triapitsyn sp. n. (Costa Rica), G. (Cosmocomoidea) cuscus Triapitsyn sp. n. (Peru), G. (Cosmocomoidea) garchamp Triapitsyn sp. n. (Argentina), G. (Cos- mocomoidea) gerasim Triapitsyn sp. n. (Mexico), G. (Cosmocomoidea) hispaniolus Triapitsyn & Huber sp. n. (Dominican Republic), G. (Cosmocomoidea) kiskis Triapitsyn sp. n. (Argentina), G. (Cosmocomoidea) logarzoi Triapitsyn sp. n. (Argentina), G. (Cosmocomoidea) mumu Triapitsyn sp. n. (Argentina), and G. (Cosmocomoidea) rakitovi Triapitsyn sp. n. (Costa Rica). Seven new synonymies are proposed: Gonatocerus h-luteum (Ogloblin) syn. n. under G. (Cosmoco- moidea) nigriflagellum (Girault); G. enicmophilus Huber syn. n. and G. necator (Ogloblin) syn. n. under G. (Cosmoco- moidea) bonariensis (Brèthes); G. dimorphus (Ogloblin) syn. n. and G. monrosi (Ogloblin) syn. n. under G. (Gastrogonatocerus) margiscutum Girault; and G. setulosus (Ogloblin) syn. n. and G. dorsiniger (Ogloblin) syn. n. under G. (Gastrogonatocerus) membraciphagus Ogloblin. Litus maculipennis Ashmead, Gahanopsis deficiens (Ogloblin), and Gahanopsis straeleni (Debauche) are transferred to Gonatocerus as, respectively, G. (Cosmocomoidea) maculipennis (Ashmead) comb. n., G. (Gahanopsis) deficiens (Ogloblin) comb. n., and G. (Lymaenon) straeleni (Debauche) comb. n. Lectotypes are designated for the following 23 species: G. (Gahanopsis) acanophorae (Ogloblin), G. (Gahanopsis) aethalionis (Ogloblin), G. (Cosmocomoidea) annulicornis (Ogloblin), G. (Gastrogonatocerus) anomocerus Crawford, G. (Gonatocerus) appendiculatus (Ogloblin), G. (Gonatocerus) bonaerensis (Ogloblin), G. (Cosmocomoidea) caudatus (Ogloblin), G. (Cosmocomoidea) concinnus (Ogloblin), Lymaenon (Gastrogonatocerus) dimorphus Ogloblin, G. (Gona- tocerus) excisus (Ogloblin), G. (Cosmocomoidea) gracilicornis (Ogloblin), G. (Cosmocomoidea) grandis (Ogloblin), Lymaenon h-luteum Ogloblin, G. (Cosmocomoidea) inauditus (Ogloblin), G. (Gastrogonatocerus) juvator Perkins, G. (Gastrogonatocerus) margiscutum Girault, G. (Cosmocomoidea) metanotalis (Ogloblin), G. (Cosmocomoidea) nasutus (Ogloblin), Lymaenon necator Ogloblin, G. (Cosmocomoidea) nigrithorax (Ogloblin), G. (Lymaenon) pratensis (Ogloblin), G. (Gonatocerus) stenopterus (Ogloblin), and G. (Gonatocerus) urocerus (Ogloblin).

scholarly journals Characterization of the types of the Neotropical Pseudisobrachium (Hymenoptera: Bethylidae), with a key to species

2008 ◽  
Vol 25 (4) ◽  
pp. 737-801 ◽  
Author(s):  
Celso O. Azevedo
Keyword(s):  
New Species ◽  
New Genus ◽  
Neotropical Region ◽  
Species Group ◽  
New Combination ◽  
Valid Species ◽  
Key To Species ◽  
Neotropical Species

The identity of many Neotropical species of Pseudisobrachium Kieffer, 1904 is currently unreachable, mostly because the available descriptions and illustrations are not sufficiently accurate to allow precise identifications. To help solve the problem, 115 valid species and their junior synonyms were examined. Twelve nomenclatural acts are made, and 110 valid species are recognized for the Neotropical region. Two lectotypes are designated: Pristocera crassicornis Westwood and Pristocera haemorrhoidalis Westwood. Seven new species-group synonyms are proposed: Pseudisobrachium retusum Evans syn. nov. for P. pauxillum Evans; P. cunco Perez syn. nov. for P. erythrocephalum Evans; P. navajo Evans, P. rectangulatum Evans, P. emarginatum Evans and P. foutsi Evans syn. nov. for P. flavinervis Fouts; P. acuminatum Waichert & Azevedo syn. nov. for P. latum Waichert & Azevedo. The following new genus-group synonym is established: Parisobrachium Kieffer syn. nov. for Dissomphalus Ashmead. The following new combination and reinstated name are proposed: Dissomphalus albipes (Kieffer) comb. nov. and nom. rev. from Pseudisobrachium paraguayense Kieffer.

Additions to the knowledge of Manota Williston (Diptera: Mycetophilidae) from the Neotropical region, with description of four new species

Zootaxa ◽  
2021 ◽  
Vol 4938 (1) ◽  
pp. 85-100
Author(s):  
OLAVI KURINA ◽  
HEIKKI HIPPA
Keyword(s):  
New Species ◽  
Dominican Republic ◽  
French Guiana ◽  
New Records ◽  
First Record ◽  
Neotropical Region ◽  
Caribbean Island ◽  
Additional Species ◽  
Neotropical Species ◽  
Southern Caribbean

The Neotropical species of the genus Manota Williston are studied, based on material of 146 specimens from French Guiana, Ecuador, Nicaragua, Dominica and the Dominican Republic. Four new species are described, viz. M. corniculata sp. n. (French Guiana), M. pseudocavata sp. n. (French Guiana), M. truuverki sp. n. (French Guiana) and M. vladi sp. n. (Dominican Republic). Manota defecta Williston, 1896, the type species of the genus, is listed from Dominica, representing the first record since its description more than a century ago from a Southern Caribbean Island, St. Vincent. New records of 13 additional species are provided: M. acutistylus Jaschhof & Hippa, 2005 (Dominica), M. aligera Hippa, Kurina & Sääksjärvi, 2017 (French Guiana), M. digitata Hippa, Kurina & Sääksjärvi, 2017 (French Guiana), M. iota Hippa & Kurina, 2013 (French Guiana), M. micula Hippa & Kurina, 2013 (French Guiana), M. nordestina Kurina, Hippa & Amorim, 2018 (French Guiana), M. parva Jaschhof & Hippa, 2005 (Nicaragua), M. pauloides Hippa, Kurina & Sääksjärvi, 2017 (Ecuador), M. perplexa Kurina, Hippa & Amorim, 2017 (Nicaragua), M. rotundistylus Jaschhof & Hippa, 2005 (Ecuador), M. serrulata Hippa, Kurina & Sääksjärvi, 2017 (French Guiana), M. spinosa Jaschhof & Hippa, 2005 (French Guiana), M. subaristata Kurina, Hippa & Amorim, 2017 (Ecuador). The number of Neotropical Manota species has risen to 96. 

Review of some species groups of the genus Oospila Warren, with descriptions of nine new species (Lepidoptera: Geometridae: Geometrinae)

Zootaxa ◽  
2018 ◽  
Vol 4497 (2) ◽  
pp. 151 ◽  
Author(s):  
AARE LINDT ◽  
AXEL HAUSMANN ◽  
JAAN VIIDALEPP
Keyword(s):  
New Species ◽  
Costa Rica ◽  
Species Complex ◽  
French Guiana ◽  
Species Group ◽  
New Subspecies ◽  
Wing Patterns ◽  
Species Groups ◽  
Males And Females

The Neotropical geometrine genus Oospila Warren, 1897 includes seventy-nine species and was revised by Cook & Scoble (1995). The genus is distinctive in having a row of raised abdominal crests, which are composed of specialized, erect, metallic shining scales. This paper focuses on the integrative morphological and molecular delimitation of the smallest Oospila species. The wing patterns and genitalia structures of males and females are illustrated. Cook & Scoble (1995) distinguished 13 species groups within Oospila. We discuss the species of the Oospila flavilimes species group, the O. stigma species group and O. miccularia species group below, and separate the O. arpata species complex into a group of its own. Nine new species and two new subspecies are described in this paper: O. cristae sp. n. from Ecuador, O. falcata sp. n. from French Guiana, O. pallidaria boliviensis subsp. n. from Bolivia, and O. loreenae sp. n. from Bolivia (flavilimes species group), O. ehakernae sp. n. from Costa Rica, O. similiplaga bolarpata subsp. n. from Bolivia (arpata species group), O. brehmi sp. n. and O. bifida sp. n. both from Bolivia, O. moseri sp. n. from Brazil, O. absaloni sp. n. and O. pipa sp. n. both from Ecuador (miccularia species group). Oospila similiplaga (Warren) (stat. nov.) is raised here from synonymy with O. arpata (Schaus) and O. imula (Dognin) from synonymy with O. miccularia (Guenée), respectively. Oospila agnetaforslundae nom. nov. is proposed as a replacement name for Oospila marginata Schaus, 1912 (nec Oospila marginata Warren, 1897), raising it to species rank from synonymy of Oospila permagna (Warren, 1909). With this paper, the number of Neotropical Oospila species is raised to 88. 

scholarly journals Fourteen new species ofOecetisMcLachlan, 1877 (Trichoptera: Leptoceridae) from the Neotropical region

PeerJ ◽  
2017 ◽  
Vol 5 ◽  
pp. e3753 ◽  
Author(s):  
Fabio B. Quinteiro ◽  
Ralph W. Holzenthal
Keyword(s):  
New Species ◽  
Neotropical Region ◽  
Valid Species ◽  
Light Traps ◽  
Larval Stages ◽  
The Past ◽  
Biogeographical Region ◽  
Species Groups ◽  
Malaise Traps ◽  
Maxillary Palps

BackgroundThe caddisfly genusOecetiscurrently contains 534 valid species. Its larval stages are found in freshwaters around the world. The adults can be distinguished from other Leptoceridae by the unbranched forewing M vein and the exceptionally long maxillary palps. In the Neotropical region, 55 species ofOecetishave been recorded and most of them can be placed in one of the six species groups known from this biogeographical region: theavara-,falicia-,inconspicua-,punctata-,punctipennis-, andtestacea-groups. More than 50% of the known diversity of NeotropicalOecetishas been described in the past 40 years. Here, we describe an additional 14 new species ofOecetisto further document the diversity of this genus in the Neotropical region.MethodsThe descriptions and illustrations presented here are based on male specimens. Specimens were collected with Malaise traps or ultraviolet light traps. They were preserved in alcohol or pinned as stated in material examined section. Specimens had their genitalia prepared in 85% lactic acid to better observe internal characters and illustrations were aided by the use of a microscope with drawing tube attached.Results and DiscussionThis study raises the number of species ofOecetisin the Neotropics from 55 to 69. Eight of the new species presented here could not be reliably placed in one of the known species groups (Oecetis acuticlaspern. sp.,Oecetis flintin. sp.,Oecetis carinatan. sp.,Oecetis cassicoleatan. sp.,Oecetis blahnikin. sp.,Oecetis gibbosan. sp.,Oecetis licinan. sp., andOecetis pertican. sp.). The others are placed in thepunctata-group (Oecetis bidigitatan. sp.,Oecetis quasipunctatan. sp.),testacea-group (Oecetis plenuspinosan. sp.), andfalicia-group (Oecetis calorin. sp.,Oecetis hastapullan. sp.,Oecetis machaeran. sp.). Most of the diagnostic characters rely on structures of the inferior appendages and phallic apparatus, and the shape of tergum X.

scholarly journals Taxonomic revision of the Pheidole sikorae species group (Hymenoptera, Formicidae) from Madagascar

ZooKeys ◽  
2020 ◽  
Vol 949 ◽  
pp. 1-185
Author(s):  
Sebastian Salata ◽  
Brian L. Fisher
Keyword(s):  
New Species ◽  
Taxonomic Revision ◽  
Species Group ◽  
Species Level ◽  
Junior Synonym ◽  
Identification Key ◽  
Valid Species ◽  
Species Groups ◽  
Worker Castes

The present study represents a taxonomic revision of the Pheidole sikorae species group from Madagascar. Forty-four members of this group are recognised and described, and an illustrated identification key to this group is also presented. One species is raised to species level: P. litigiosa Forel, 1892 stat. nov.Pheidole veteratrix angustinoda Forel, 1892 syn. nov. is proposed as a junior synonym of Pheidole veteratrix Forel, 1891. Worker castes are also described and lectotypes designated for P. litigiosa Forel, 1892, P. sikorae Forel, 1891, and P. veteratrix Forel, 1891. The following 41 new species are described: P. alinasp. nov., P. ambohimangasp. nov., P. analavelonasp. nov., P. andohahelasp. nov., P. anomalasp. nov., P. anosyennesp. nov., P. antranohofasp. nov., P. beankasp. nov., P. befotakasp. nov., P. dasossp. nov., P. flavominutasp. nov., P. gracilissp. nov., P. habokasp. nov., P. havoanasp. nov., P. hazosp. nov., P. itremosp. nov., P. joffrevillesp. nov., P. kelysp. nov., P. lavasoasp. nov., P. mahamavosp. nov., P. maintysp. nov., P. mamiratrasp. nov., P. mananteninasp. nov., P. masoandrosp. nov., P. mavohavoanasp. nov., P. midongysp. nov., P. mikrossp. nov., P. mivorysp. nov., P. nitidobrunasp. nov., P. parvulasp. nov., P. parvulogibbasp. nov., P. reniranosp. nov., P. savasp. nov., P. sofiasp. nov., P. sparsasp. nov., P. tamponysp. nov., P. trichotossp. nov., P. tsaravonianasp. nov., P. vadumsp. nov., P. volontanysp. nov., and P. vonysp. nov. At present, there are 109 valid species and subspecies of Pheidole known from Madagascar, but this number is expected to increase with upcoming taxonomic revisions of the species groups not revised in this study.

Description of two new species of Prosapia (Hemiptera: Cercopidae) from the Nearctic and Neotropics, with a key to species of the P. inferens species group

2020 ◽  
Vol 152 (3) ◽  
pp. 288-297
Author(s):  
Ulises Castro-Valderrama ◽  
Daniel C. Peck ◽  
Gervasio S. Carvalho ◽  
Jorge Manuel Valdez-Carrasco ◽  
Jesús Romero Nápoles
Keyword(s):  
New Species ◽  
Costa Rica ◽  
Neotropical Region ◽  
Species Group ◽  
Pest Species ◽  
Key To Species ◽  
Two New Species

AbstractInsects in the subfamily Ischnorhininae (Hemiptera: Auchenorrhyncha: Cercopidae), known as spittlebugs or froghoppers, are mainly distributed in the Neotropical region. A few genera include pest species of economic relevance, including Prosapia Fennah, 1949. Two new species from Mexico and Costa Rica are now described for this genus, and a key to species is proposed for the P. inferens (Walker, 1858) species group.

scholarly journals Taxonomy and phylogeny of New World Polyplectropus Ulmer, 1905 (Trichoptera: Psychomyioidea: Polycentropodidae) with the description of 39 new species

Zootaxa ◽  
2010 ◽  
Vol 2582 (1) ◽  
pp. 1 ◽  
Author(s):  
MARIA LOURDES CHAMORRO ◽  
RALPH W. HOLZENTHAL
Keyword(s):  
New Species ◽  
Bayesian Methods ◽  
New World ◽  
Morphological Characters ◽  
Neotropical Region ◽  
Species Group ◽  
Biogeographic Region ◽  
Parsimony Tree ◽  
Group A ◽  
Species Groups

The taxonomy of the New World species of Polyplectropus (Ulmer, 1905a) is revised to include detailed male and female diagnoses, descriptions, illustrations, distribution records, and keys to males of all species and species groups. A phylogenetic analysis based on 59 morphological characters, 89 of 92 (97%) New World Polyplectropus species, and 2 outgroup taxa was inferred using parsimony and Bayesian methods, which resulted in minor topological differences. Conflicting estimates of relationship among and within most species groups led to a less resolved Bayesian tree (vs. parsimony tree) due to high variation in rates of change among characters and an overall low number of characters. A new classification for New World Polyplectropus is proposed with revised characterization of 10 recognized species groups, 6 newly established. Four species remain unassigned to species group. A key to genera of New World Polycentropodidae, including a redescription of Polyplectropus is provided. The homology of the male genitalia of Polyplectropus is discussed. Ninety-two species are treated. The following 39 new species are described: Polyplectropus adamsae (Peru), P. alatespinus (Brazil), P. amazonicus (Brazil), P. andinensis (Argentina, Bolivia), P. blahniki (Venezuela), P. bolivianus (Bolivia), P. brasilensis (Brazil), P. brborichorum (Ecuador), P. cressae (Venezuela), P. colombianus (Colombia), P. corniculatus (Peru), P. cuzcoensis (Peru), P. ecuadoriensis (Ecuador), P. flintorum (Venezuela), P. gaesum (Brazil), P. guyanae (Guyana, Venezuela), P. hollyae (Brazil), P. hystricosus (Brazil), P. insularis (Panama), P. juliae (Brazil), P. kanukarum (Guyana), P. maculatus (Venezuela), P. manuensis (Peru), P. matatlanticus (Brazil), P. minensium (Brazil), P. novafriburgensis (Brazil), P. peruvianus (Peru), P. petrae (Brazil), P. pratherae (Brazil), P. puyoensis (Ecuador), P. robertsonae (Bolivia), P. rodmani (Brazil), P. rondoniensis (Brazil), P. tragularius (Brazil), P. tripunctatum (Peru), P. venezolanus (Venezuela), P. woldai (Panama), P. zamoranoensis (Honduras), and P. zuliae (Venezuela). Polyplectropus buchwaldi (Ulmer, 1911) is designated as a nomen dubium.Key words: phylogeny, taxonomy, Neotropics, Nearctric, Bayesian methods, parsimony, classification, cladistics, morphology, Trichoptera, Polycentropodidae, Polyplectropus, new speciesThe polycentropodid caddisfly genus Polyplectropus Ulmer contains 219 species found primarily in the tropical and subtropical regions of the world, with 125 species recorded from the Oriental biogeographic region, 53 from the Neotropical region (6 species extending their ranges into the Nearctic region), 37 from the

Four new Neotropical species of Episcepsis Butler, 1877 (Lepidoptera: Erebidae: Arctiinae)

Zootaxa ◽  
2021 ◽  
Vol 5020 (2) ◽  
pp. 352-366
Author(s):  
KARLA MANTILLA ◽  
JUAN GRADOS
Keyword(s):  
New Species ◽  
Male Genitalia ◽  
Neotropical Region ◽  
External Morphology ◽  
Valid Species ◽  
Similar Species ◽  
Genital Morphology ◽  
Neotropical Species ◽  
Species Complexes

The genus Episcepsis Butler, 1877 is widely distributed in the Neotropical region and currently includes 43 valid species many of them described solely based on external morphology. The existence of species complexes makes their identification difficult without examination of genital morphology. Four new species from Brazil, Cuba and Peru are described herein; Episcepsis andina sp. nov., E. diversa sp. nov., E. insularis sp. nov. and E. travassosi sp. nov. Comparisons with other similar species as well as photographs of external morphology and male genitalia are provided.  

Taxonomic revision of New World species of the genus Oosternum Sharp (Coleoptera: Hydrophilidae: Sphaeridiinae) I. Definition of species groups and revision of the Oosternum aequinoctiale group

Zootaxa ◽  
2009 ◽  
Vol 2054 (1) ◽  
pp. 1-37 ◽  
Author(s):  
MARTIN FIKÁČEK ◽  
FRANZ HEBAUER ◽  
MICHAEL HANSEN
Keyword(s):  
Costa Rica ◽  
New World ◽  
Taxonomic Revision ◽  
Neotropical Region ◽  
Species Group ◽  
Lesser Antilles ◽  
World Species ◽  
Diagnostic Characters ◽  
Species Groups ◽  
Definition Of

The genus Oosternum Sharp, 1882 is divided into ten species groups based on external adult characters. An identification key to the species groups and a table of diagnostic characters as well as many charater state illustrations for each group are provided. Representatives of the newly defined O. aequinoctiale species group are revised. Six species of this group are recognized, all occurring in the Neotropical region: Oosternum acutheca sp.n. (Honduras, Costa Rica, Nicaragua), O. aequinoctiale (Motschulsky, 1855) (Bolivia, Colombia, Costa Rica, Ecuador, Mexico, Panama, Peru, Venezuela), O. attenuatum sp. n. (Panama, Colombia, Ecuador), O. gibbicolle sp. n. (Ecuador, Panama), O. holosericeum sp. n. (Argentina), and O. latum sp. n. (Lesser Antilles: St. Vincent Island). A key to the species and drawings or SEM photographs of diagnostic characters are provided.

Revision of the genus Bradyporus Charpentier, 1825 (Orthoptera: Tettigoniidae: Bradyporinae)

Zootaxa ◽  
2017 ◽  
Vol 4272 (4) ◽  
pp. 491 ◽  
Author(s):  
MUSTAFA ÜNAL
Keyword(s):  
New Species ◽  
Phylogenetic Relationships ◽  
Species Group ◽  
Iucn Red List ◽  
Red List ◽  
Valid Species ◽  
Check List ◽  
Species Groups ◽  
First Time ◽  
A New Species

The genus Bradyporus Charpentier is taxonomically revised using information obtained from numerous recently collected specimens. Bradyporus is divided into two subgenera Bradyporus Charpentier s.str. and Callimenus Fischer von Waldheim. Five species groups are distinguished in the latter subgenus. Sixteen species and subspecies are listed. Bradyporus (Callimenus) multituberculatus multituberculatus (Fischer von Waldheim, 1833) and Bradyporus (Callimenus) multituberculatus montandoni (Burr, 1898) are designated subspecies for the first time. Previously synonymized Bradyporus (Callimenus) macrogaster skopjensis Karaman, 1961 is reinstated as a valid species. A new species, Bradyporus (Callimenus) gocmeni Ünal, sp. nov. from Eastern Turkey and the previously unknown female of Bradyporus (Callimenus) skopjensis Karaman, 1961 are described. A Neotype for Bradyporus (Callimenus) macrogaster longicollis (Fieber, 1853) is designated. Nomenclatural and taxonomical problems are discussed under the relevant taxa. Phylogenetic relationships based on morphology of the subgenera, species groups, and the species and subspecies of each species group are proposed. Morphological derived and ancestral characters are discussed. A key to the lower taxa of the genus is provided with 207 original figures including 3 distributional maps. A check-list of the lower taxa of the genus is given. Additionally the IUCN Red List statuses of the species and subspecies are included. 

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