Systematics of Paraleucopis Malloch with proposal of Paraleucopidae, a new family of acalyptrate Diptera

Zootaxa ◽  
2019 ◽  
Vol 4668 (3) ◽  
pp. 301-328
Author(s):  
TERRY A. WHEELER ◽  
BRADLEY J. SINCLAIR

Paraleucopidae Wheeler fam. nov. is proposed for the previously unplaced New World genera Paraleucopis Malloch, Mallochianamyia Santos-Neto and Schizostomyia Malloch and undescribed Australian species. A key to genera of Paraleucopidae is provided. Paraleucopis is revised and includes nine species: P. auripes Wheeler & Sinclair sp. nov. (type locality: Andalgala, Argentina); P. bispinosa Wheeler & Sinclair sp. nov. (type locality: Socos, Coquimbo, Chile); P. boharti Wheeler & Sinclair sp. nov. (type locality: Andalgala, Argentina); P. boydensis Steyskal (type locality: nr. Palm Desert, California, USA); P. corvina Malloch (type species of genus; type locality: New Mexico, USA); P. mexicana Steyskal (type locality: Kino Bay, Mexico); P. nigra Wheeler & Sinclair sp. nov. (type locality: Portal, Arizona, USA); P. paraboydensis Wheeler & Sinclair sp. nov. (type locality: Willis Palms Oasis, California, USA); P. saguaro Wheeler & Sinclair sp. nov. (type locality: Usery Mtn Park, Arizona, USA). A key to the species of Paraleucopis is provided. The distribution of Paraleucopis is disjunct, with six species in the western United States and northwestern Mexico and three species in northern Chile and northern Argentina.                The sister group and superfamilial assignment of the Paraleucopidae cannot be established based on current knowledge although the family has affinities to some families of the Asteioinea sensu J.F. McAlpine. A well-supported hypothesis on the relationships of the families of the Acalyptratae will be required before the sister group relationships of Paraleucopidae can be determined. 

2003 ◽  
Vol 17 (3) ◽  
pp. 387 ◽  
Author(s):  
Gonzalo Giribet

Examination of museum specimens belonging to the cyphophthalmid Gondwanan family Pettalidae has rendered a collection of specimens placed in the new genus Karripurcellia. The genus includes three species, K.�peckorum, sp. nov., K. sierwaldae, sp. nov. and K. harveyi, sp. nov., from the Pemberton area in Western Australia. These are the first cyphophthalmid species found outside Queensland in Australia. Karripurcellia, gen. nov. is distinguished from other Australian pettalids by the lack of modifications in the anal plate, as well as the lack of the typical male anal glands of pettalids, sironids and the stylocellid genus Fangensis Rambla, 1994. Two of the species, K. peckorum, sp. nov. and K. sierwaldae, sp. nov., live sympatrically and have been collected in the same litter samples in one locality. The proposition of the new genus is accompanied by a cladistic analysis of all pettalid genera and most species within each genus, with the exception of the species-rich genus Rakaia Hirst, 1925. The cladistic analysis supports the monophyly of the Karripurcellia, gen. nov. species, but their sister-group relationships are unclear. However, the remaining Australian species cluster with the pettalids from New Zealand and South Africa, but not with Karripurcellia, sp. nov.


1985 ◽  
Vol 16 (1) ◽  
pp. 27-67 ◽  
Author(s):  
Henrik Enghoff

AbstractThe family Nemasomatidae is redefined to include onty genera with all sterna secondarily free from pleurotergites. Comments are given on the included genera, viz., Antrokoreana, Basoncopus gen. n. (type-species B. filiformis sp. n.) (Kazakhstan), Dasynemasoma, Thalassisobates, Sinostemmiulus, Nemasoma, and Orinisobates. Isobates coiffaiti Demange, 1961 is synonymized with Thalassisobates littoralis (Silvestri, 1903). Orinisobates is revised and shown to include O. soror sp. n. (Kuril Islands), O. microthylax sp. n. (Kamchatka and Siberia), O. gracilis (Verhoeff, 1933) (NW China), O. sibiricus (Gulicka, 1963) (Altai region, Kazakhstan), O. kasakstanus (Lohmander, 1933) (Kazahkstan), O. nigrior (Chamberlin, 1943) (eastern United States), O. utus (Chamberlin, 1912) (northwestern United States), and O. expressus (Chamberlin, 1941) (northwestern United States and adjacent Canada). Mimolene oregona Chambertin, 1941 and M. sectile Loomis & Schmitt, 1971 are synonymized with O. expressus. A possible case of parthenogenesis in O. microthylax is recorded. Evidence is presented for the following sister-group relationships: Antrokoreana + (Basoncopus + (Dasynemasoma + (Thalassisobates + (Sinostemmiulus + (Orinisobates + Nemasoma))))). The position of Basoncopus is uncertain, and O. soror may belong in a separate genus and constitute the sister-group of Orinisohates + Nemasoma. If soror does belong in Orinisobates, it is the sister-group of all its congeners. The American species of Orinisobates are shown probably to constitute a monophyietic group. The family is suggested to have originated in the eastern Palearctic region, Orinisobates having invaded North America via the Bering Bridge. Doubtful species and species erroneously assoiciated with the Nemasomatidae are listed. The genera Okeanobates and Yosidaiulus are excluded from the family and referred to Okeanobatidae stat. n. in superfamily Blaniuloidea. The genera Trichonemasoma, Telsonemasoma, and Chelojulus are also excluded from the Nemasomatidae and relegated to Julida incertae sedis.


2021 ◽  
Author(s):  
Gonzalo Giribet ◽  
Kate Sheridan ◽  
Caitlin M. Baker ◽  
Christina J. Painting ◽  
Gregory I. Holwell ◽  
...  

The Opiliones family Neopilionidae is restricted to the terranes of the former temperate Gondwana: South America, Africa, Australia, New Caledonia and New Zealand. Despite decades of morphological study of this unique fauna, it has been difficult reconciling the classic species of the group (some described over a century ago) with recent cladistic morphological work and previous molecular work. Here we attempted to investigate the pattern and timing of diversification of Neopilionidae by sampling across the distribution range of the family and sequencing three markers commonly used in Sanger-based approaches (18S rRNA, 28S rRNA and cytochrome-c oxidase subunit I). We recovered a well-supported and stable clade including Ballarra (an Australian ballarrine) and the Enantiobuninae from South America, Australia, New Caledonia and New Zealand, but excluding Vibone (a ballarrine from South Africa). We further found a division between West and East Gondwana, with the South American Thrasychirus/Thrasychiroides always being sister group to an Australian–Zealandian (i.e. Australia + New Zealand + New Caledonia) clade. Resolution of the Australian–Zealandian taxa was analysis-dependent, but some analyses found Martensopsalis, from New Caledonia, as the sister group to an Australian–New Zealand clade. Likewise, the species from New Zealand formed a clade in some analyses, but Mangatangi often came out as a separate lineage from the remaining species. However, the Australian taxa never constituted a monophyletic group, with Ballarra always segregating from the remaining Australian species, which in turn constituted 1–3 clades, depending on the analysis. Our results identify several generic inconsistencies, including the possibility of Thrasychiroides nested within Thrasychirus, Forsteropsalis being paraphyletic with respect to Pantopsalis, and multiple lineages of Megalopsalis in Australia. In addition, the New Zealand Megalopsalis need generic reassignment: Megalopsalis triascuta will require its own genus and M. turneri is here transferred to Forsteropsalis, as Forsteropsalis turneri (Marples, 1944), comb. nov.


2010 ◽  
Vol 60 (2) ◽  
pp. 460-468 ◽  
Author(s):  
Miao Miao ◽  
Yangang Wang ◽  
Weibo Song ◽  
John C. Clamp ◽  
Khaled A. S. Al-Rasheid

Recently, an undescribed marine ciliate was isolated from China. Investigation of its morphology and infraciliature revealed it as an undescribed species representing a new genus, Eurystomatella n. gen., the type of the new family Eurystomatellidae n. fam. The new family is defined by close-set, apically positioned oral membranelles and a dominant buccal field that is surrounded by an almost completely circular paroral membrane. The new genus is defined by having a small oral membranelle 1 (M1), bipartite M2 and well-developed M3, a body surface faintly sculptured with a silverline system in a quadrangular, reticulate pattern and a cytostome located at the anterior third of a large buccal field. The type species of the new genus, Eurystomatella sinica n. sp., is a morphologically unique form that is defined mainly by the combination of a conspicuously flattened body, several caudal cilia, extremely long cilia associated with the buccal apparatus and a contractile vacuole located subcaudally. According to phylogenetic analyses of small-subunit (SSU) rRNA gene sequences, Eurystomatella clusters with the genus Cyclidium, as a sister group to the family Pleuronematidae. The great divergence in both buccal and somatic ciliature between Eurystomatella and all other known scuticociliates supports the establishment of a new family for Eurystomatella.


2018 ◽  
Vol 65 (5) ◽  
pp. 579-588 ◽  
Author(s):  
Xiaoping Wang ◽  
Burton K Lim ◽  
Nelson Ting ◽  
Jingyang Hu ◽  
Yunpeng Liang ◽  
...  

Abstract Among mammalian phylogenies, those characterized by rapid radiations are particularly problematic. The New World monkeys (NWMs, Platyrrhini) comprise 3 families and 7 subfamilies, which radiated within a relatively short time period. Accordingly, their phylogenetic relationships are still largely disputed. In the present study, 56 nuclear non-coding loci, including 33 introns (INs) and 23 intergenic regions (IGs), from 20 NWM individuals representing 18 species were used to investigate phylogenetic relationships among families and subfamilies. Of the 56 loci, 43 have not been used in previous NWM phylogenetics. We applied concatenation and coalescence tree-inference methods, and a recently proposed question-specific approach to address NWM phylogeny. Our results indicate incongruence between concatenation and coalescence methods for the IN and IG datasets. However, a consensus was reached with a single tree topology from all analyses of combined INs and IGs as well as all analyses of question-specific loci using both concatenation and coalescence methods, albeit with varying degrees of statistical support. In detail, our results indicated the sister-group relationships between the families Atelidae and Pitheciidae, and between the subfamilies Aotinae and Callithrichinae among Cebidae. Our study provides insights into the disputed phylogenetic relationships among NWM families and subfamilies from the perspective of multiple non-coding loci and various tree-inference approaches. However, the present phylogenetic framework needs further evaluation by adding more independent sequence data and a deeper taxonomic sampling. Overall, our work has important implications for phylogenetic studies dealing with rapid radiations.


2020 ◽  
Vol 3 (4) ◽  
pp. 382-389
Author(s):  
CORENTIN JOUAULT ◽  
JEAN-MARC POUILLON ◽  
ANDRÉ NEL

A new wood wasp, Cratosirex sennlaubi gen. et sp. nov., is described and figured from one specimen collected from the Lower Cretaceous Crato Formation in northeastern Brazil. This new genus is placed in the new siricid subfamily Cratosiricinae subfam. nov., based on a combination of plesiomorphic and autapomorphic characters. The presence of small and sub-equal forewing cells 1R1 and 2R1 is a synapomorphy with the extant subfamily Siricinae, absent in the other extinct subfamilies †Auliscinae and †Gigasiricinae, supporting a sister group relationships with the Siricinae. Our new discovery expands the distribution range of Siricidae fossil records, highlights the antiquity of the family, and emphasizes the need for more studies of this particular insect lineage in the Mesozoic deposits. Currently, all the representatives of the crown group of the extant Siricidae are Cenozoic.


2003 ◽  
Vol 60 (3) ◽  
pp. 425-447 ◽  
Author(s):  
M. MÖLLER ◽  
M. KIEHN

Our knowledge of cytological data published on members of the family Gesneriaceae is summarized and critically evaluated in the light of current taxonomic treatments and phylogenetic hypotheses. There are about 1000 published chromosome counts, covering 56% of the genera but only 18% of the species. In particular the New World tribes Beslerieae and Napeantheae and the Old World tribe Didymocarpeae are underexplored at generic level. In Gesneriaceae chromosome data are a valuable source of taxonomic characters. From our current knowledge of the phylogenetic relationships in the family we know that basic chromosome numbers in the New World subfamily Gesnerioideae appear to be rather conserved, but that a more complex pattern of genome evolution seems to be present among the Old World tribes. Both polyploidy and dysploid changes have played a significant role in the evolution of the family. However, the number of species for which both cytological and molecular data are available is at present too low to reach firm conclusions on ancestral basic chromosome numbers, particularly for the Old World group. To facilitate wider access to cytological data on the Gesneriaceae, a website has been developed (http://www.rbge.org.uk/rbge/web/search/index.jsp), which is introduced in this paper.


1998 ◽  
Vol 353 (1375) ◽  
pp. 1645-1671 ◽  
Author(s):  
S. A. Ridgway ◽  
D. G. Reid ◽  
J. D. Taylor ◽  
G. M. Branch ◽  
A. N. Hodgson

A phylogenetic hypothesis for the patellid limpets is reconstructed by cladistic analysis of morphological characters from 37 species, representing all but one of the living members of the family. Characters included in the analysis are derived from shell shape and microstructure, headfoot and pallial complex, radula and sperm. The species fall into four clades, providing the basis for a new phylogenetic classification into four monophyletic genera: Helcion (four species; southern Africa), Cymbula (eight species; southern Africa, eastern Atlantic, southern Indian Ocean), Scutellastra (17 species; southern and southwestern Africa, Australia, Indo–West Pacific, Eastern Pacific) and Patella (nine species; northeastern Atlantic and Mediterranean). The analysis suggests sister–group relationships between Helcion and Cymbula , and between Scutellastra and Patella . In combination with present–day patterns of geographical distribution, this phylogenetic hypothesis is used to discuss the historical biogeography of the Patellidae. Scutellastra may have originated in southern Africa and dispersed across the Pacific, or alternatively may be a primitively Tethyan group. Both Helcion and Cymbula appear to have originated in southern Africa, but three Cymbula species have dispersed respectively to northwest Africa, St Helena and the southern Indian Ocean. The patellids of the northeastern Atlantic form a single clade, Patella (including P. pellucida ), which may have arrived by northward dispersal of an ancestor from southern Africa, or possibly by vicariance of a widespread ancestral Tethyan distribution. The known fossil record of patellids is too fragmentary to permit choice between these alternatives.


2016 ◽  
Vol 149 (1) ◽  
pp. 8-25 ◽  
Author(s):  
Bjarte H. Jordal ◽  
Johanna Kaidel

AbstractMicracidini (Coleoptera: Curculionidae: Scolytinae) is an unusual tribe of mainly bigynous bark beetles found in dry forests and scrublands in Afrotropical and Neotropical regions. Their phylogenetic relationship to other bark beetle groups is poorly known with few clues from external morphology. Hence, a phylogenetic analysis of five genes (COI, EF-1a, 28S, CAD, ArgK) and morphological (internal and external) data was conducted to test potential sister group relationships, including 56 outgroup genera in 22 tribes, and 18 species in 10 genera of Micracidini.CactopinusSchwarz – a genus with many cactus feeding species – was nested within a clade of all Neotropical and Nearctic genera. The New World was colonised by an Afrotropical ancestor about 75–85 million years ago, where cactus feeding inCactopinusevolved much later. All analyses indicated a paraphyletic clade of Afrotropical micracidines, strongly supporting inclusion of the Ipini genusDendrochilusSchedl inAfromicracisSchedl. Hypoborini appear to be one of the more plausible sistergroup candidates to Micracidini, and revealed morphological similarity in protibial and proventricular characters. Most phylogenetic results were supported independently by morphological and molecular data and therefore document the power of thorough examination of morphological characters analysed properly in a phylogenetic context.


Bothalia ◽  
1967 ◽  
Vol 9 (2) ◽  
pp. 309-333 ◽  
Author(s):  
Peter H Raven

There are in Africa 10 native species of Epilobium, one of Circaea, and between 8 and 12 of Ludwigia (including  Jussiaea), depending upon how many species are considered native. These three genera include all native African species of Onagraceae. Four sections of Ludwigia, cornprising a total of five species, and one fairly distinctive group of Epilobium are the most characteristic African groups. Despite the fact that H. Perrier de la Bathie (in Humb., FI. Madagasc., Oenotheracs : 1-26, 1950) considered all three of the species of Epilobium that he recognized and four of the 11 species of Ludwigia (including  Jussiaea) to be endemic to Madagascar, there are in my opinion no Mascarene endemics in the family Onagraceae and nothing to suggest any great antiquity for the family on that island. This observation agrees with the general pattern for the family in Africa in suggesting that plants of this family are relatively recent arrivals on that continent. Indeed this generality could be extended to apply to the Old World as a whole, where there is no endemic genus of the family (as opposed to 16 endemic genera and 2 endemic tribes—Lopezieae and Onagreae—in the New World, principally in Mexico and the western United States) and relatively few endemic sections or subgenera. The ten African species and one subspecies of  Epilobium recognized are described, and a key provided.


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