COMBINING ABILITY FOR KERNEL PLUMPNESS IN A DIALLEL CROSS OF FIVE CANADIAN BARLEY CULTIVARS
A complete diallel cross of Bonanza, Conquest, Galt, Jubilee, and Olli barley was made and the parents, F1 and F2 bulks, were subjected to a kernel-sizing procedure, using five different standard sieves. Mean values for the P, F1, and F2 were obtained and heritabilities and estimates of general (GCA) and specific (SCA) combining ability were calculated. Reciprocal effects were not significant in either F1 or F2. For the F1 data significant differences could only be found on the 19 × 2.4-mm (3/4 × 6/64-inch) sieve. Only GCA was significant for the F1, suggesting additive gene action. By contrast, in the F2 GCA was highly significant for all sieve sizes from 19 × 2.2 mm (3/4 × 11/128 inch) upwards, and SCA was significant on those of size 19 × 2.4 mm (3/4 × 6/64 inch) upwards. For the F2 on the 2.4-mm (6/64-inch) screen, the SCA effect exceeded the GCA effect in magnitude, and was in a negative direction, contributing to the two cases of negative heterosis for kernel plumpness. No significant positive heterosis was found in F2, and F2 bulks were normally intermediate in value to their parents. The amount of SCA effect was unrelated to the degree of similarity of the parents by pedigree, or to whether they were malting or feed types. Bonanza, Conquest, and Jubilee were significantly plumper than Galt and Olli, and Olli transmitted its less plump kernel type to the F2 generation, as indicated by its large negative GCA effect. None of the other cultivars exhibited significant GCA effects. Little progress in increasing kernel plumpness can be expected from crosses between these cultivars, as heritabilities were also generally low.