Climatic Aridity Shapes Post-Fire Interactions between Ceanothus spp. and Douglas-Fir (Pseudotsuga menziesii) across the Klamath Mountains

Climate change is leading to increased drought intensity and fire frequency, creating early-successional landscapes with novel disturbance–recovery dynamics. In the Klamath Mountains of northwestern California and southwestern Oregon, early-successional interactions between nitrogen (N)-fixing shrubs (Ceanothus spp.) and long-lived conifers (Douglas-fir) are especially important determinants of forest development. We sampled post-fire vegetation and soil biogeochemistry in 57 plots along gradients of time since fire (7–28 years) and climatic water deficit (aridity). We found that Ceanothus biomass increased, and Douglas-fir biomass decreased with increasing aridity. High aridity and Ceanothus biomass interacted with lower soil C:N more than either factor alone. Ceanothus biomass was initially high after fire and declined with time, suggesting a large initial pulse of N-fixation that could enhance N availability for establishing Douglas-fir. We conclude that future increases in aridity and wildfire frequency will likely limit post-fire Douglas-fir establishment, though Ceanothus may ameliorate some of these impacts through benefits to microclimate and soils. Results from this study contribute to our understanding of the effects of climate change and wildfires on interspecific interactions and forest dynamics. Management seeking to accelerate forest recovery after high-severity fire should emphasize early-successional conifer establishment while maintaining N-fixing shrubs to enhance soil fertility.

Relocation of habituated black bears in the Klamath Mountains of California: an adaptive management case study

Black bear (Ursus americanus) populations in California have increased in abundance and distribution despite rising trends in the urban encroachment of wildlands. As human-black bear conflicts increase, opportunities to study the relocation of black bears in an adaptive management setting are important for improving the management of this highprofile species. Habituated black bears, some tamed and made tractable through human-controlled food conditioning, were relocated to a remote region of the Klamath Mountains to analyze home range use, survival, return rates, and mortality. Relocated black bears with known outcomes demonstrated an 80% return rate, with 55% not surviving beyond five months. Female bears established home ranges significantly larger than males, and may suggest an enhanced maternal instinct in search of similar nutritional conditions prior to relocation. This study showed that the relocation of food-conditioned black bears resulted in high return rates, poor survival, and risk to public safety.

Middle Jurassic to Early Cretaceous tectonic evolution of the western Klamath Mountains and outboard Franciscan assemblages, northern California–southern Oregon, USA

ABSTRACT The Klamath Mountains province and adjacent Franciscan subduction complex (northern California–southern Oregon) together contain a world-class archive of subduction-related growth and stabilization of continental lithosphere. These key elements of the North American Cordillera expanded significantly from Middle Jurassic to Early Cretaceous time, apparently by a combination of tectonic accretion and continental arc– plus rift-related magmatic additions. The purpose of this field trip is twofold: to showcase the rock record of continental growth in this region and to discuss unresolved regional geologic problems. The latter include: (1) the extent to which Mesozoic orogenesis (e.g., Siskiyou and Nevadan events plus the onset of Franciscan accretion) was driven by collision of continental or oceanic fragments versus changes in plate motion, (2) whether growth involved “accordion tectonics” whereby marginal basins (and associated fringing arcs) repeatedly opened and closed or was driven by the accretion of significant volumes of material exotic to North America, and (3) the origin of the Condrey Mountain schist, a composite low-grade unit occupying an enigmatic structural window in the central Klamaths—at odds with the east-dipping thrust sheet regional structural “rule.” Respectively, we assert that (1) if collision drove orogenesis, the requisite exotic materials are missing (we cannot rule out the possibility that such materials were removed via subduction and/or strike slip faulting); (2) opening and closure of the Josephine ophiolite-floored and Galice Formation–filled basin demonstrably occurred adjacent to North America; and (3) the inner Condrey Mountain schist domain is equivalent to the oldest clastic Franciscan subunit (the South Fork Mountain schist) and therefore represents trench assemblages underplated >100 km inboard of the subduction margin, presumably during a previously unrecognized phase of shallow-angle subduction. In aggregate, these relations suggest that the Klamath Mountains and adjacent Franciscan complex represent telescoped arc and forearc upper plate domains of a dynamic Mesozoic subduction zone, wherein the downgoing oceanic plate took a variety of trajectories into the mantle. We speculate that the downgoing plate contained alternating tracts of smooth and dense versus rough and buoyant lithosphere—the former gliding into the mantle (facilitating slab rollback and upper plate extension) and the latter enhancing basal traction (driving upper plate compression and slab-shallowing). Modern snapshots of similarly complex convergent settings are abundant in the western Pacific Ocean, with subduction of the Australian plate beneath New Guinea and adjacent island groups providing perhaps the best analog.

A crucial geologic test of Late Jurassic exotic collision versus endemic re-accretion in the Klamath Mountains Province, western United States, with implications for the assembly of western North America

Differing interpretations of geophysical and geologic data have led to debate regarding continent-scale plate configuration, subduction polarity, and timing of collisional events on the western North American plate margin in pre–mid-Cretaceous time. One set of models involves collision and accretion of far-traveled “exotic” terranes against the continental margin along a west-dipping subduction zone, whereas a second set of models involves long-lived, east-dipping subduction under the continental margin and a fringing or “endemic” origin for many Mesozoic terranes on the western North American plate margin. Here, we present new detrital zircon U-Pb ages from clastic rocks of the Rattlesnake Creek and Western Klamath terranes in the Klamath Mountains of northern California and southern Oregon that provide a test of these contrasting models. Our data show that portions of the Rattlesnake Creek terrane cover sequence (Salt Creek assemblage) are no older than ca. 170–161 Ma (Middle–early Late Jurassic) and contain 62–83% Precambrian detrital zircon grains. Turbidite sandstone samples of the Galice Formation are no older than ca. 158–153 Ma (middle Late Jurassic) and contain 15–55% Precambrian detrital zircon grains. Based on a comparison of our data to published magmatic and detrital ages representing provenance scenarios predicted by the exotic and endemic models (a crucial geologic test), we show that our samples were likely sourced from the previously accreted, older terranes of the Klamath Mountains and Sierra Nevada, as well as active-arc sources, with some degree of contribution from recycled sources in the continental interior. Our observations are inconsistent with paleogeographic reconstructions that are based on exotic, intra-oceanic arcs formed far offshore of North America. In contrast, the incorporation of recycled detritus from older terranes of the Klamath Mountains and Sierra Nevada, as well as North America, into the Rattlesnake Creek and Western Klamath terranes prior to Late Jurassic deformation adds substantial support to endemic models. Our results suggest that during long-lived, east-dipping subduction, the opening and subsequent closing of the marginal Galice/Josephine basin occurred as a result of in situ extension and subsequent contraction. Our results show that tectonic models invoking exotic, intra-oceanic archipelagos composed of Cordilleran arc terranes fail a crucial geologic test of the terranes’ proposed exotic origin and support the occurrence of east-dipping, pre–mid-Cretaceous subduction beneath the North American continental margin.

Supplemental Material: A Crucial geologic test of Late Jurassic exotic collision versus endemic re-accretion in the Klamath Mountains Province, western United States, with implications for the assembly of western North America

Analytical methods, sample collection locations in the southern Klamath Mountains, and Data tables.

Supplemental Material: A Crucial geologic test of Late Jurassic exotic collision versus endemic re-accretion in the Klamath Mountains Province, western United States, with implications for the assembly of western North America

Analytical methods, sample collection locations in the southern Klamath Mountains, and Data tables.

Linking modern pollen accumulation rates to biomass: Quantitative vegetation reconstruction in the western Klamath Mountains, NW California, USA

Quantitative reconstructions of vegetation abundance from sediment-derived pollen systems provide unique insights into past ecological conditions. Recently, the use of pollen accumulation rates (PAR, grains cm−2 year−1) has shown promise as a bioproxy for plant abundance. However, successfully reconstructing region-specific vegetation dynamics using PAR requires that accurate assessments of pollen deposition processes be quantitatively linked to spatially-explicit measures of plant abundance. Our study addressed these methodological challenges. Modern PAR and vegetation data were obtained from seven lakes in the western Klamath Mountains, California. To determine how to best calibrate our PAR-biomass model, we first calculated the spatial area of vegetation where vegetation composition and patterning is recorded by changes in the pollen signal using two metrics. These metrics were an assemblage-level relevant source area of pollen (aRSAP) derived from extended R-value analysis ( sensu Sugita, 1993) and a taxon-specific relevant source area of pollen (tRSAP) derived from PAR regression ( sensu Jackson, 1990). To the best of our knowledge, aRSAP and tRSAP have not been directly compared. We found that the tRSAP estimated a smaller area for some taxa (e.g. a circular area with a 225 m radius for Pinus) than the aRSAP (a circular area with a 625 m radius). We fit linear models to relate PAR values from modern lake sediments with empirical, distance-weighted estimates of aboveground live biomass (AGLdw) for both the aRSAP and tRSAP distances. In both cases, we found that the PARs of major tree taxa – Pseudotsuga, Pinus, Notholithocarpus, and TCT (Taxodiaceae, Cupressaceae, and Taxaceae families) – were statistically significant and reasonably precise estimators of contemporary AGLdw. However, predictions weighted by the distance defined by aRSAP tended to be more precise. The relative root-mean squared error for the aRSAP biomass estimates was 9% compared to 12% for tRSAP. Our results demonstrate that calibrated PAR-biomass relationships provide a robust method to infer changes in past plant biomass.