Triassic roots of the amphiastraeid scleractinian corals

2001 ◽  
Vol 75 (1) ◽  
pp. 34-45 ◽  
Author(s):  
Ewa Roniewicz ◽  
Jarosław Stolarski

The Early Carnian (Upper Triassic) phaceloid coral originally described by Volz (1896) asHexastraea fritschi, type species ofQuenstedtiphylliaMelnikova, 1975, reproduced asexually by “Taschenknospung” (pocket-budding), a process documented herein for the first time. This type of budding is recognized only in the Amphiastraeidae, a family thus far recorded only from Jurassic-Cretaceous strata. Similar to amphiastraeids,Quenstedtiphyllia fritschi(Volz, 1896) has separate septal calcification centers and a mid-septal zone built of serially arranged trabeculae. The most important discriminating characters of the new amphiastraeid subfamily Quenstedtiphylliinae are one-zonalendotheca and radial symmetry of the corallite in the adult stage (in contrast to two-zonal and bilateral symmetry in the adult stage in Amphiastraeinae).Quenstedtiphyllia fritschishares several primitive skeletal characters (plesiomorphies) with representatives of Triassic Zardinophyllidae and, possibly, Paleozoic plerophylline rugosans: e.g., thick epithecal wall and strongly bilateral early blastogenetic stages with the earliest corallite having one axial initial septum. To interpret the phylogenetic status of amphiastraeid corals, we performed two analyses using plerophylline rugosans and the solitary scleractinianProtoheterastraea, respectively, as the outgroups. The resulting phylogenetic hypotheses support grouping the Zardinophyllidae with the Amphiastraeidae in the clade Pachythecaliina (synapomorphy: presence of pachytheca). Taschenknospung is considered an autapomorphy for the Amphiastraeidae. This study is the first attempt to analyze the relationships of the Triassic corals cladistically.

Zootaxa ◽  
2012 ◽  
Vol 3271 (1) ◽  
pp. 43 ◽  
Author(s):  
REYES PEÑA-SANTIAGO ◽  
JOAQUÍN ABOLAFIA ◽  
MAJID PEDRAM

Labronemella labiata, the type species of its genus, is described after studying an Iranian population. An emendeddiagnosis of the species is provided, with new morphological and morphometrical features. The lip region, the mostdistinctive character of Labronemella is observed under SEM by the first time and its detailed morphology elucidated.Anterior surface (oral field) appears distinctly sunken and with its inner (perioral) area differentiated in six large, separatedliplets protruding on oral field. The lateral, perioral liplets are trapezoidal and visibly larger than subdorsal and subventralones, which are triangular, consequently the lip region shows a bi-radial symmetry. Inner labial papillae migrate to themargin of labial disc, being located close to outer labial and cephalic papillae. The systematics of Labronemella is brieflydiscussed, including diagnosis, updated list of species, key to their identification as well as a compendium of their main measurements and morphometrics.


Zootaxa ◽  
2019 ◽  
Vol 4706 (1) ◽  
pp. 48-70
Author(s):  
ANDREY V. MATALIN

Within the Asiatic tiger beetle fauna, Parmecus Motschulsky, 1864 stat. rest., stat. nov., is reestablished as a subgenus of Cylindera Westwood, 1831 with Cylindera (Parmecus) dromicoides (Chaudoir, 1852), as its type species, and the lectotype and paralectotypes of Cicindela dromicoides Chaudoir, 1852 are designated as well. Two other species are included, Cylindera (Parmecus) armandi (Fairmaire, 1886), from the Himalayan Region, and Cylindera (Parmecus) mosuoa, sp. nov., from Yunnan, China. Cylindera (Parmecus) as a subgenus is characterized, a key to identify its species is provided, and its species composition is discussed. Cylindera (Parmecus) dromicoides (Chaudoir, 1852) is newly recorded from Pakistan and the Indian state of Jammu and Kashmir, while Cylindera (Parmecus) armandi (Fairmaire, 1886) is recorded for the first time from the Chinese province of Sichuan. The records of C. armandi from Bhutan, as well as C. dromicoides from Yunnan Province (China) are rejected due to erroneous identifications. 


2014 ◽  
Vol 369 (1648) ◽  
pp. 20130348 ◽  
Author(s):  
Lena C. Hileman

A striking aspect of flowering plant (angiosperm) diversity is variation in flower symmetry. From an ancestral form of radial symmetry (polysymmetry, actinomorphy), multiple evolutionary transitions have contributed to instances of non-radial forms, including bilateral symmetry (monosymmetry, zygomorphy) and asymmetry. Advances in flowering plant molecular phylogenetic research and studies of character evolution as well as detailed flower developmental genetic studies in a few model species (e.g. Antirrhinum majus , snapdragon) have provided a foundation for deep insights into flower symmetry evolution. From phylogenetic studies, we have a better understanding of where during flowering plant diversification transitions from radial to bilateral flower symmetry (and back to radial symmetry) have occurred. From developmental studies, we know that a genetic programme largely dependent on the functional action of the CYCLOIDEA gene is necessary for differentiation along the snapdragon dorsoventral flower axis. Bringing these two lines of inquiry together has provided surprising insights into both the parallel recruitment of a CYC -dependent developmental programme during independent transitions to bilateral flower symmetry, and the modifications to this programme in transitions back to radial flower symmetry, during flowering plant evolution.


Development ◽  
1958 ◽  
Vol 6 (3) ◽  
pp. 486-490
Author(s):  
S. Løvtrup ◽  
A. Pigon

According to the hypothesis advanced by Løvtrup (1958) the supply of oxygen is one of the factors responsible for the determination of bilateral symmetry in amphibian embryos. The protein coat covering the outside of the egg is known to have a very low permeability (Holtfreter, 1943), and it was suggested in the hypothesis that the formation of the grey crescent consists in a stretching of this coat by which the permeability is increased (cf. the work of Dalcq & Dollander (1948) and of Dollander & Melnotte (1952) on permeability of Nile blue), in this way the radial symmetry of the egg is changed to a bilateral symmetry from a metabolic point of view. As a consequence of the increase in permeability those oxidative, energy-supplying processes which are associated with gastrulation are enabled to proceed at a higher rate at one side of the egg.


Phytotaxa ◽  
2021 ◽  
Vol 497 (2) ◽  
pp. 127-137
Author(s):  
XIAN-LIN GUO ◽  
MEGAN PRICE ◽  
WEI GOU ◽  
SONG-DONG ZHOU ◽  
XIN-FEN GAO ◽  
...  

The genus Similisinocarum Cauwet & Farille (Apiaceae, Apioideae), was first described by Cauwet & Farille in 1984, but it has been considered as a synonym of Sinocarum H.Wolff ex R.H. Shan & F.T. Pu. In this study, Simlisinocarum normanianum, the type species of Similisinocarum was found in China for the first time, and the molecular phylogenetic evidence based on ITS sequences reveals that Similisinocarum normanianum occupies an individual clade, which is sister to the clade comprising seven species of Acronema, Sinocarum, and Oreocomopsis in Acronema clade, the values of genetic distances (ITS) between Similisinocarum and Acronema, Similisinocarum and Oreocomopsis, Similisinocarum and Pternopetalum, Similisinocarum and Sinocarum are close (≥0.0916), while deviated from the values within Acronema, Sinocarum and Pternopetalum (≤0.0580). Morphologically, we found Similisinocarum normanianum develops reflex bracteoles with white membranous margin, petals with an entire margin and incurved apex, and mericarps with many vittae in the furrow (≥ 3) and commissure (6), which are distinguishable from other species of the Acronema clade. Consequently, results manifested in molecular and morphological analyses indicate that Similisinocarum should be treated as an independent genus distinct from Sinocarum.


Zootaxa ◽  
2010 ◽  
Vol 2481 (1) ◽  
Author(s):  
CARLOS MOLINERI

The 12 species previously placed in Tortopus together with 3 species newly described here, are revised and included in a phylogenetic analysis. Based on synapomorphic characters on the nymphs and adults of both sexes, Tortopus is restricted to T. igaranus Needham & Murphy, T. circumfluus Ulmer, T. harrisi Traver, T. zottai (Navás), T. bellus Lugo-Ortiz & McCafferty, and T. arenales sp. nov., and the genus is defined by: female parastyli receptors with long furrows anterior to sockets; penes entirely flattened; male ninth abdominal sternum almost separated in two portions by a median notch; mesosternum with furcasternal protuberances contiguous only on basal corner; and nymphs with two subapical tubercles on mandibular tusks. Tortopsis is newly described for T. bruchianus (Navás), T. limoncocha sp. nov., T. obscuripennis (Domínguez), T. parishi (Banks), T. primus (McDunnough), T. puella (Pictet), T. sarae (Domínguez), T. spatula sp. nov., and T. unguiculatus (Ulmer). Tortopsis is characterized by: R sector of female fore wing without additional veins between R 2 and IR; female parastyli receptors C or V-shaped, with sockets opening towards median line; male gonopore associated with a claw-like structure; penes separated from the base; parastyli more than 5 times length of pedestals; parastyli curved in lateral view; nymphs with a single subapical tubercle on mandibular tusks. The study of available type material permitted inclusion of comparative diagnoses, with figures and redescriptions as needed. The male imago of the type species of Tortopus (T. igaranus Needham & Murphy) is described for the first time, as are the female adults of Tortopus bellus Lugo-Ortiz & McCafferty and T. harrisi Traver. Three new Neotropical species based on male and female adults are described: Tortopus arenales and Tortopsis limoncocha from Ecuador, and Tortopsis spatula from Colombia. Keys to separate the adults and nymphs of the genera of Polymitarcyidae, and for male and female adults of all the species of Tortopus and Tortopsis are presented, as well as line drawings, pictures and SEM photographs of important structures.


2021 ◽  
Vol 63 (3-4) ◽  
pp. 351-390
Author(s):  
S. Y. Kondratyuk ◽  
L. Lőkös ◽  
I. Kärnefelt ◽  
A. Thell ◽  
M.-H. Jeong ◽  
...  

Seven genera new to science, i.e.: Helmutiopsis, Huriopsis, Johnsheardia, Klauskalbia, Kudratovia, Kurokawia and Poeltonia of the Physciaceae are proposed for the ‘Rinodina’ atrocinerea, the ‘Rinodina’ xanthophaea, the ‘Rinodina’ cinnamomea, the ‘Heterodermia’ obscurata, the ‘Rinodina’ straussii, the ‘Anaptychia’ isidiata and the ‘Physconia’ grisea groups consequently that all form strongly supported monophyletic branches in a phylogeny analysis based on a combined matrix of nrITS and mtSSU sequences. Phylogenetic positions of species belonging to the genera Kashiwadia s. l., Leucodermia, Mischoblastia,Oxnerella, Phaeorrhiza s. l., Polyblastidium and Rinodinella s. l. are discussed. Oxnerella afghanica which for the first time recorded as parasitic lichen species from both epiphytic and saxicolous crustose lichens is designated as type species for the genus Oxnerella. Sequences of the recently described Physcia orientostellaris as well as Huriopsis xanthophaea and additional sequences of Kashiwadia aff. orientalis and Mischoblastia aff. oxydata are submitted to the GenBank. The positions of Polyblastidium casaterrinum from Costa Rica, ‘Rinodina’ efflorescens from Białowieża, Poland, and ‘Mischoblastia’ confragosula from Cambodia in the Physciaceae are confirmed in a phylogeny analysis based on the nrITS sequences. The presence of ‘extraneous mycobiont DNA’ in lichen associations is exemplified with earlier incorrect identifications of Heterodermia, Kashiwadia, Kurokawia,Oxnerella and Poeltonia specimens. Fifty-six new combinations are presented: Helmutiopsis alba (for Rinodina alba Metzler ex Arn.), Helmutiopsis aspersa (for Lecanora aspersa Borrer), Helmutiopsis atrocinerea (for Parmelia atrocinerea Fr.), Huriopsis chrysidiata (for Rinodina chrysidiata Sheard), Huriopsis chrysomelaena (for Rinodina chrysomelaena Tuck.), Huriopsis lepida (for Lecanora lepida Nyl.), Huriopsis luteonigra (for Rinodina luteonigra Zahlbr.), Huriopsis plana (for Rinodina plana H. Magn.), Huriopsis thiomela (for Lecanora thiomela Nyl.), Huriopsis xanthomelana (for Rinodina xanthomelana Müll. Arg.), Huriopsis xanthophaea (for Lecanora xanthophaea Nyl.), Johnsheardia cinnamomea (for Rinodina mniaroea var. cinnamomea Th. Fr.), Johnsheardia herteliana (for Rinodina herteliana Kaschik), Johnsheardia jamesii (for Rinodina jamesii H. Mayrhofer), Johnsheardia reagens (for Rinodina reagens Matzer et H. Mayrhofer), Johnsheardia zwackhiana (for Lecanora zwackhiana Kremp.), Kashiwadia austrostellaris (for Physcia austrostellaris Elix), Kashiwadia jackii (for Physcia jackii Moberg), Kashiwadia littoralis for Physcia littoralis Elix), Kashiwadia nubila (for Physcia nubila Moberg), and Kashiwadia tropica (for Physcia tropica Elix), Klauskalbia crocea (for Heterodermia crocea R. C. Harris), Klauskalbia flabellata (for Parmelia flabellata Fée), Klauskalbia obscurata (for Physcia speciosa (Wulfen) Nyl. *obscurata Nyl.), Klauskalbia paradoxa (for Heterodermia paradoxa Schumm et Schäfer-Verwimp), Kudratovia bohlinii (for Rinodina bohlinii H. Magn.), Kudratovia candidogrisea (for Rinodina candidogrisea Hafellner, Muggia et Obermayer), Kudratovia luridata (for Buellia luridata Körb.), Kudratovia metaboliza (for Rinodina metaboliza Vain.), Kudratovia pycnocarpa (for Rinodina pycnocarpa H. Magn.), Kudratovia roscida (for Lecanora roscida Sommerf.), Kudratovia straussii (for Rinodina straussii J. Steiner), Kudratovia terrestris (for Rinodina terrestris Tomin), Kurokawia bryorum (for Anaptychia bryorum Poelt), Kurokawia isidiata (for Anaptychia isidiata Tomin), Kurokawia mereschkowskii (for Physcia mereschkowskii Tomin), Kurokawia palmulata (for Psoroma palmulatum Michx.), Kurokawia runcinata (for Lichen runcinatus With.), Kurokawia stippea (for Parmelia aquila var. stippea Ach.), Lecania safavidiorum (for Oxnerella safavidiorum S. Y. Kondr., Zarei-Darki, Lőkös et Hur), Leucodermia erinacea (for Lichen erinaceus Ach.), Mischoblastia confragosula (for Lecanora confragosula Nyl.), Mischoblastia destituta (for Lecidea destituta Nyl.), Mischoblastia moziana (for Lecanora moziana Nyl.), Mischoblastia moziana subsp. parasitica (comb. et stat. nova for Rinodina moziana var. parasitica Kaschik et H. Mayrhofer), Mischoblastia ramboldii (for Rinodina ramboldii Kaschik), Mischoblastia vezdae (for Rinodina vezdae H. Mayrhofer), Oxnerella afghanica (for Rinodina afghanica M. Steiner et Poelt), Oxnerella castanomelodes (for Rinodina castanomelodes H. Mayrhofer et Poelt), Physciella nigricans (for Lecanora nigricans Flörke), Poeltonia elegantula (for Physconia elegantula Essl.), Poeltonia grisea (for Lichen griseus Lam.), Poeltonia isidiomuscigena (for Physconia isidiomuscigena Essl.), Poeltonia perisidiosa (for Physcia perisidiosa Erichsen), Poeltonia venusta (for Parmelia venusta Ach.), and Polyblastidium albicans (for Parmelia albicans Pers.) are proposed.


1987 ◽  
Vol 119 (3) ◽  
pp. 215-230 ◽  
Author(s):  
Alasdair J. Ritchie ◽  
Joseph D. Shorthouse

AbstractThe species of Synophromorpha Ashmead (Hymenoptera: Cynipidae) are reviewed. One new species is described (Synophromorpha kaulbarsi; type locality: Naupan, Puebla, Mexico). The previously described species (S. rubi Weld, S. sylvestris (O.S.), and S. terricola Weld) are redescribed, and a key to species is presented. All species are illustrated for the first time. Synophromorpha sylvestris is designated type-species for the genus and a lectotype is chosen. Hypotheses on the phylogenetic relationships between the species of Synophromorpha are presented.


1977 ◽  
Vol 114 (3) ◽  
pp. 209-214 ◽  
Author(s):  
Harrell L. Strimple

SummaryCryphiocrinus Kirk is a genus which may possess a delicate stem, or may lose the stem as an adult, making it essentially eleutherozoic. C. girtyi Kirk, type species, is documented with certainty from Chesterian rocks of Oklahoma and Arkansas for the first time. C. rotundus Kirk is reported from the Renault Formation of Illinois and a possible occurrence in the Pride Mountain Formation of Alabama is noted. The genus has previously been recognized in Chesterian, Upper Mississippian, (Lower Carboniferous), strata exposed in West Virginia, Kentucky and Oklahoma, U.S.A. Possible affinity with Hosieocrinus Wright from the Lower Limestone Group (Visean) in Scotland is discussed.


2018 ◽  
Vol 93 (1) ◽  
pp. 115-125
Author(s):  
John S. Peel

AbstractPhosphatic sclerites of the problematicTarimspiraYue and Gao, 1992 (Cambrian Series 2) recovered by weak acid maceration of limestones display a unique range of mainly strongly coiled morphologies. They were likely organized into multielement scleritomes, but the nature of these is poorly known; some sclerites may have had a grasping function.Tarimspirasclerites grew by basal accretion in an analogous fashion to younger paraconodonts (Cambrian Series 3–4) but lack a basal cavity. Based on proposed homologies,Tarimspiramay provide an extension of the early vertebrate paraconodont–euconodont clade back into the early Cambrian.Tarimspirais described for the first time from Laurentia (North Greenland), extending its known range from China and Siberia in Cambrian Series 2. In addition to the type species,Tarimspira planaYue and Gao, 1992, the Greenland record ofTarimspiraincludes two morphotypes of a new species,Tarimspira artemi.UUID:http://zoobank.org/c7c536c8-cdaf-49a9-ae1d-77c392f553fc.


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