Hardseededness and the pattern of softening in Biserrula pelecinus L., Ornithopus compressus L., and Trifolium subterraneum L. seeds

1999 ◽  
Vol 50 (6) ◽  
pp. 1073 ◽  
Author(s):  
A. Loi ◽  
P. S. Cocks ◽  
J. G. Howieson ◽  
S. J. Carr
Keyword(s):  
Seed Bank ◽  
Subterranean Clover ◽  
Soil Surface ◽  
Trifolium Subterraneum ◽  
Farming System ◽  
First Year ◽  
Ley Farming ◽  
Sufficient Variation ◽  
Depth Of Burial ◽  
Selection Of

Experiments measuring seed bank size, hardseededness, and seed softening of biserrula (Biserrula pelecinus L.) were conducted at Merredin and Perth in Western Australia. At Merredin, a mixture of 2 biserrula accessions was grazed, shallow cultivated, or left uncultivated and ungrazed. Seed bank size, seedling regeneration, and seed softening were measured over 2 years. At Perth, softening of biserrula, yellow serradella, and subterranean clover seeds grown at 2 sites (Binnu and Northam) was compared on the soil surface and after burial at 2 and 10 cm over a period of 2 years. Seed bank size of biserrula at Merredin ranged from 14000 to 17500 seeds/m2. Regeneration was greater in the second year (800–1700 seedlings/m2) than in the first year (40–600 seedlings/m2). In both years the shallow cultivated treatment recorded the highest number of seedlings. About 90% of biserrula and serradella seed remained hard after 2 years on the soil surface, compared with only about 10% of subterranean clover. Serradella softened more rapidly when buried 2 cm below the soil surface (8–12% hard) than it did on the soil surface (84–92% hard) (P<0.05). In contrast, the softening of subterranean clover decreased with increasing depth. Biserrula was intermediate, although it too softened most rapidly at 2 cm (78–95% hard compared with 82–97% on the surface) (P<0.05). Rate of seed softening in all species decreased with increasing depth of burial below 2 cm. Of the 4 accessions of biserrula, an accession from Greece (83% hard after 2 years exposure) was significantly softer than the other accessions. The results indicate that biserrula is very hardseeded, although there is sufficient variation in hardseededness for the selection of somewhat softer lines. Its pattern of softening suggests that biserrula may be successful in the ley farming system (crop/pasture rotations) of southern Australia.

Effect of tillage practices on the fate of hard seeds of subterranean clover in a ley farming system

1985 ◽  
Vol 25 (3) ◽  
pp. 568 ◽  
Author(s):  
GB Taylor
Keyword(s):  
Subterranean Clover ◽  
Soil Surface ◽  
Trifolium Subterraneum ◽  
Farming System ◽  
Conventional Tillage ◽  
No Tillage ◽  
Minimum Tillage ◽  
Tillage Practices ◽  
Tillage Treatment ◽  
Clover Seed

In a rotation of 1 year pasture/l year crop, a subterranean clover (Trifolium subterraneum cv. Daliak) pasture was either left untilled or subjected to minimum or conventional tillage. One set of tillage treatments was imposed in each ofthree crop years while another set of treatments was imposed in only the first crop year. Regenerating clover plants were prevented from setting seed. In the first crop, 44% of the clover seeds were buried below 2 cm of soil by minimum tillage; this proportion was 65% in the conventional tillage treatment. In the first pasture regeneration year, seedling densities were highest in the no-tillage treatment. Conversely, there were more residual seeds in the tilled treatments and, in the second and third pasture regeneration years, this led to higher seedling densities than in the no-tillage treatment. The effects of tillage were more marked in the conventional than in the minimum-tillage treatment. Clover establishment was improved by repeat tillage operations which returned some of the buried seeds closer to the soil surface. Although more seedlings overall were obtained from the no-tillage treatment, the disadvantage of fewer seedlings in the tilled treatments was offset by the spread of seedling establishment over a number of pasture years. This spread, which would be more marked with harder-seeded cultivars, could be desirable in environments in which clover seed production is unreliable.

Effect of depth of burial on the longevity of hard seeds of subterranean clover and annual medics

1988 ◽  
Vol 28 (1) ◽  
pp. 77 ◽  
Author(s):  
GB Taylor ◽  
MA Ewing
Keyword(s):  
Subterranean Clover ◽  
Soil Surface ◽  
Trifolium Subterraneum ◽  
Barrel Medic ◽  
Hard Seeds ◽  
Burr Medic ◽  
Seed Reserves ◽  
Annual Medics ◽  
Depth Of Burial ◽  
Tillage Operations

Burrs of 3 cultivars of subterranean clover (Trifolium subterraneum) and 1 cultivar each of burr medic (Medicago polymorpha) and barrel medic (M. truncatula), which had experienced 1 summer at the soil surface, were placed on the soil surface and at depths of 2, 6 and 10 cm in the soil. The numbers of residual hard seeds were determined each year for up to 4 years. There was a marked reduction in the rate of seed softening in all 3 clover cultivars with increasing depth of burial. Whereas <20% of the seeds of the hardest seeded clover cultivar, Nungarin, survived 3 years at the soil surface, there was no significant decline in seed numbers during 4 years of burial at 10 cm. Even with cv. Geraldton, in which only 5% of seeds remained after 1 year of placement at the soil surface, 75% of seeds survived 4 years of burial at 10 cm. Hard seeds of both medic varieties were considerably more resilient than clover seeds at the soil surface, particularly during the first summer following seed set. However, burial had much less effect on their longevity, with no significant effect of burial to 2 cm in either medic, or of burial to 6 cm in the case of barrel medic. These results support earlier findings which showed that tillage operations associated with crop establishment which result in the burial of substantial proportions of subterranean clover seeds can lead to useful soil seed reserves. The much lesser effect of burial on seed softening of the medics, compared with subterranean clover, suggests that tillage operations will be less advantageous to medic persistence in leys.

Plant population dynamics in subterranean clover and murex medic swards. I. Size and composition of the seed bank

1994 ◽  
Vol 45 (4) ◽  
pp. 913 ◽  
Author(s):  
MJ Blumenthal ◽  
RL Ison
Keyword(s):  
High Temperature ◽  
Seed Bank ◽  
Subterranean Clover ◽  
Soil Surface ◽  
Trifolium Subterraneum ◽  
Controlled Environment ◽  
Plant Population Dynamics ◽  
Embryo Dormancy ◽  
Alternating Temperature ◽  
Total Seed

It is possible that the use of harder-seeded varieties of sub. clover (Trifolium subterraneum) and the recently domesticated murex medic (Medicago murex) may improve annual legume persistence in pasture leys. To determine the mechanisms of production and persistence in these two species, the size and composition of the seed bank were compared in four varieties of sub. clover and two lines of murex medic over a three-year period at Forbes in central-west N.S.W. The rate of breakdown of hardseed and the loss of high temperature dormancy was also studied in the same genotypes under conditions of alternating temperature in controlled environment cabinets. In terms of the size of the total seed bank, the sub. clover cultivar with the highest level of hardseed (Dalkeith) was the most successful genotype studied. This was the result of slower rates of hardseed breakdown compared to the other sub. clover cultivars, and better seed production compared to the two murex medic lines. Losses of seed from the seed bank were substantial, and were not all accounted for by germinated seedlings or grazing over summer. The sub. clover cultivars Junee and Seaton Park had the highest levels of high temperature embryo dormancy and the lowest proportion of seedlings germinating over summer. The level of sub. clover hardseed breakdown in the field corresponded well with the 60/15�C laboratory estimates of hardseed breakdown, even though soil surface temperatures over summer at Forbes were rarely in the range of 60/15�C.

Plant population dynamics in subterranean clover and murex medic swards. 3. Effect of pod burial, summer grazing and autumn cultivation on emergence

1996 ◽  
Vol 36 (5) ◽  
pp. 533 ◽  
Author(s):  
MJ Blumenthal ◽  
RL Ison
Keyword(s):  
Soil Water ◽  
Seedling Recruitment ◽  
Subterranean Clover ◽  
Soil Surface ◽  
Trifolium Subterraneum ◽  
Farming Systems ◽  
Plant Population Dynamics ◽  
Eastern Australia ◽  
Ley Farming ◽  
Summer Grazing

Murex medic (Medicago murex Willd.) seedling recruitment is more sensitive to soil water at the time of emergence than subterranean clover (Trifolium subterraneum L.). Murex medic pods normally lay on the soil surface. Shallow burial of pods may be beneficial when soil moisture is marginal for germination and emergence. In addition, the tightly coiled structure of murex medic pods may also act as a barrier to water uptake by the seed. Two methods of burying murex medic pods were investigated in the field: (i) trampling by sheep hooves through summer grazing; and (ii) through light cultivation in autumn. A glasshouse experiment was also conducted to examine the interaction between the length of time that the soil stays moist and pod burial for CD26 and CD53 murex medic and Dalkeith, Junee, Seaton Park and Woogenellup subterranean clover. In the glasshouse, pod burial was important for the attainment of maximum emergence in all genotypes when soil water was limiting. However, pod structure did not appear to have a limiting role in germination and emergence in murex medic. When tested in the field, pod burial by sheep trampling through summer grazing improved emergence in CD26, possibly because the smaller more open pod was more easily trampled than that of CD53. Summer grazing in CD53 and Dalkeith and autumn cultivation in all genotypes did not improve emergence; possible reasons for this are discussed so to is the role of murex medic in ley farming systems in eastern Australia.

Superphosphate utilization in clover ley farming. I. Effects on pasture and sheep production

1974 ◽  
Vol 25 (4) ◽  
pp. 525 ◽  
Author(s):  
GD Kohn
Keyword(s):  
Agricultural Research ◽  
Subterranean Clover ◽  
Trifolium Subterraneum ◽  
Wagga Wagga ◽  
First Year ◽  
Stocking Rate ◽  
Wool Production ◽  
Ley Farming ◽  
Hordeum Leporinum ◽  
Sheep Production

From 1962 to 1966 an experiment was carried out at the Agricultural Research Institute, Wagga Wagga, N.S.W., with dry Merino sheep on an annual clover-grass pasture, in which the stocking rate and the rates of topdressing with superphosphate were varied. In the first year Trifolium subterraneum (subterranean clover) responded to superphosphate, particularly at the lowest stocking rates. In the following 2–3 years there was a change to Hordeum leporinum Link (barley grass) dominance. A greater amount of subterranean clover persisted over the years on the unfertilized plots than on the fertilized plots. Clover burr on these treatments was associated with better liveweight performance by sheep over the dry summer-autumn period. However, after the autumn rains germinating barley grass on the topdressed plots allowed rapid liveweight gains to be made by the sheep on those treatments. Greasy wool production per head, averaged over four shearings, was 4.81, 4.36 and 3.86 kg at 5, 10 and 15 sheep per ha respectively. Superphosphate had no significant effect on wool production. Liveweights decreased as the stocking rate increased. However, at a given stocking rate, the liveweight changes were influenced by changes in pasture composition induced by topdressing with superphosphate, as much as by total available pasture. It appears that in this clover ley farming area topdressing can be omitted in the pasture phase of a wool production system after 700 kg superphosphate ha-I have been applied to the soil.

Effect of burial on the softening of hard seeds of subterranean clover

1984 ◽  
Vol 35 (2) ◽  
pp. 201 ◽  
Author(s):  
GB Taylor
Keyword(s):  
Subterranean Clover ◽  
Soil Surface ◽  
Trifolium Subterraneum ◽  
Temperature Fluctuations ◽  
The Other ◽  
Soil Tillage ◽  
Microbial Decomposition ◽  
Buried Seeds ◽  
Hard Seeds ◽  
Depth Of Burial

Burrs of eight varieties of subterranean clover (Trifolium subterraneum L.), which had experienced one summer at the soil surface, were placed on the soil surface and at depths of 2, 6 and 10 cm in the soil. The numbers of residual hard seeds were determined after 1, 2 and 3 years. The effects of laboratory treatment at a diurnally fluctuating temperature of 60/15�C on the softening of buried seeds and of seeds stored in the laboratory for 1 and 3 years were determined. Rate of seed softening in all varieties decreased with increasing depth of burial, apparently because the soil insulated the seeds from high soil surface temperatures. Few seeds of the varieties Northam and Geraldton softened during 3 years of burial at 6 or 10 cm; while, at the other extreme, few seeds of Yarloop survived 3 years at any depth. Some evidence was found for microbial decomposition of hard seeds in the field. Seeds softened more readily at 60/15�C (in the laboratory) as the preceding periods of either laboratory storage or field burial increased. Such storage or burial experiences have a preconditioning effect on hard seeds, making them more amenable to softening once they are subjected to wide diurnal temperature fluctuations. The results indicate that soil tillage associated with cropping should build up a useful soil seed reserve of the harder seeded varieties.

Effects of initial clover seeding rate and length of ley on pasture production, soil nitrogen and crop yields in a ley farming system

1976 ◽  
Vol 16 (81) ◽  
pp. 484 ◽  
Author(s):  
ER Watson ◽  
P Lapins ◽  
RJW Barron
Keyword(s):  
Soil Nitrogen ◽  
Crop Yields ◽  
Wheat Yield ◽  
Subterranean Clover ◽  
Trifolium Subterraneum ◽  
Farming System ◽  
Botanical Composition ◽  
Seeding Rate ◽  
Pasture Production ◽  
Ley Farming

Several factors relating to the use of subterranean clover (Trifolium subterraneum) in a ley farming system were studied at Kojonup in the south-west of Western Australia viz; the effect of seeding rates of clover on the botanical composition and herbage production of a grazed pasture over five years; the build-up of soil nitrogen over this period, and the effect of 5, 8 and 11 years of clover on wheat yield and on subsequent regeneration of the pasture. In the final year (year 12) the effects of lime on mineralization of soil nitrogen and on crop yields were also studied. Seeding rates of clover in excess of 6 kg ha-1 affected botanical composition in the first year, and yield of pasture in the first two years only. Build-up of soil nitrogen over the first five years (when all plots were grazed) was significantly greater (P < 0.05) where clover had been sown at the higher rates (< 6 kg ha-1) but this did not result in increased wheat yields at the end of this period. In the 12th year, when all plots were cropped, there were no significant differences in grain yield between treatments cropped for the first time (after 11 years of clover) and those cropped either once or twice in the preceding six years. Cropping reduced pasture production in the early part of the post crop year but the proportion of clover in the pasture increased. The application of lime (in year 12) resulted in a significant increase in grain, and straw yields and nitrogen uptake on all treatments. The results suggest that crop yields can be sustained in short term rotations provided there is sufficient clover present in the pasture phase.

Response of defoliated swards of subterranean clover and yellow serradella to superphosphate applications

1991 ◽  
Vol 31 (6) ◽  
pp. 777
Author(s):  
MDA Bolland
Keyword(s):  
Field Experiment ◽  
Sandy Soil ◽  
Dry Matter ◽  
Maximum Yield ◽  
Subterranean Clover ◽  
Soil Surface ◽  
Trifolium Subterraneum ◽  
Herbage Yield ◽  
Ornithopus Compressus ◽  
The Relationship

The effect of superphosphate applications (0, 25, 50, 75, 100 and 125 kg P/ha to the soil surface) on the dry matter (DM) herbage production of dense swards of subterranean clover (Trifolium subterraneum cv. Junee) and yellow serradella (Ornithopus compressus cv. Tauro) was measured in a field experiment on deep, sandy soil in south-western Australia. The swards were defoliated with a reel mower at weekly intervals from 88 to 158 days after sowing, to a height of 2 cm for the first 9 cuts, 4 cm for the tenth cut and 5 cm for the eleventh cut. Yellow serradella was more productive than subterranean clover. Consequently, for the relationship between yield and the level of phosphorus (P) applied, yellow serradella supported larger maximum yields and required less P than subterranean clover, to produce the same DM herbage yield. Maximum yields of yellow serradella were 12-40% larger. To produce 70% of the maximum yield for yellow serradella at each harvest, yellow serradella required about 50% less P than subterranean clover. However, when yields were expressed as a percentage of the maximum yield measured for each species at each harvest, the relationship between yield and the level of P applied was similar for both species, and they had similar P requirements.

Effect of perennial pasture species on surface soil moisture and early growth and survival of subterranean clover (Trifolium subterraneum L.) seedlings

1997 ◽  
Vol 48 (5) ◽  
pp. 683 ◽  
Author(s):  
B. S. Dear ◽  
P. S. Cocks
Keyword(s):  
Mineral Soil ◽  
Subterranean Clover ◽  
Late Summer ◽  
Soil Surface ◽  
Trifolium Subterraneum ◽  
Annual Grass ◽  
Growth And Survival ◽  
Eastern Australia ◽  
Standing Dead ◽  
Clover Seed

Subterranean clover seedling numbers and growth in swards containing 1 of 5 perennial pasture species [phalaris (Phalaris aquatica) cv. Sirolan, cocksfoot (Dactylis glomerata) cv. Currie, lucerne (Medicago sativa) cv. Aquarius, wallaby grass (Danthonia richardsonii) cv. Taranna, and lovegrass (Eragrostis curvula) cv. Consol] were compared with those in typical annual pastures and pure clover swards in the wheatbelt of eastern Australia. Presence of a perennial species or the volunteer annual grass (Eragrostis cilianensis) increased the rate of drying of the soil surface (0–5 cm) after late February and May rain, compared with subterranean clover swards. Perennials differed in the rate they dried the soil surface, with the more summer-active lucerne and consul lovegrass drying the profile more rapidly than phalaris. The amount of water in the surface 5 cm, 6 days after the rainfall event on 27–28 February, was strongly negatively correlated (r = –0·75, P < 0·01) with the amount of green perennial biomass, but not related to standing dead material or surface residues. Where perennials were present, a smaller proportion (2–4%) of the clover seed pool produced seedlings in response to late summer rain, compared with pure clover swards (18%). A higher proportion of the seed pool produced seedlings (19–36%) following rain in late autumn but there was no difference between species. The more summer-active perennials (cocksfoot, danthonia, and lucerne) markedly depressed the survival of emerged clover seedlings following both germinations. Of the seedlings that emerged in early March, the proportion remaining by 29 March was 57% in phalaris, 21% in lucerne, 13% in danthonia, and 1% in cocksfoot, compared with a 78% increase in seedlings in pure subterranean clover swards. By 15 May, all perennials had <2 clover seedlings/m2 surviving, compared with 37 in the annual pasture and 964 plants/m2 in pure subterranean clover. Following the May germination, the highest proportion of emerged seedlings surviving until 29 May was in the phalaris swards (40%) and least in the cocksfoot and danthonia swards (2–4%). Presence of a perennial or annual grass decreased (P < 0·05) relative water content of clover seedlings on 15 March from 74% in pure clover swards, to 48% in annual pasture, 34% in phalaris, and 29% in lucerne swards. Clover seedlings growing in pure subterranean swards on 15 March (17 days after germinating rain) were 4 times larger than those in lucerne and twice as large as those in either phalaris or annual pasture. Seed size did not differ between treatments, but available mineral soil nitrogen was significantly higher (P < 0·001) in pure subterranean clover swards (32 mg N/g) compared with perennials (3–13 mg N/g). Strategies such as heavy grazing in late summer to reduce green biomass of the perennials or sowing the perennials at lower densities may reduce the adverse effects that perennials have on subterranean clover seedlings in these drier environments.

The short-term effects of rock phosphate and super-phosphate on a subterranean clover pasture.

1955 ◽  
Vol 6 (4) ◽  
pp. 553 ◽  
Author(s):  
RC Rossiter ◽  
PG Ozanne
Keyword(s):  
Rock Phosphate ◽  
Subterranean Clover ◽  
Trifolium Subterraneum ◽  
Phosphorus Recovery ◽  
Available Phosphorus ◽  
First Year ◽  
Short Term ◽  
Pasture Production ◽  
Total Plant ◽  
Second Year

A 2-year field experiment is described, in which an annual-type pasture was grown on a soil of lateritic origin with various initial rates of rock phosphate and superphosphate. The soil was acutely deficient in plant-available phosphorus at the outset. Application of superphosphate led to the expected increases in total pasture production, but rock phosphate also gave substantial yield increases, even during the first season. Differential species effects were noted; subterranean clover (Trifolium subterraneum L.) and cape-weed (Cryptostemma calendula Druce) responded about equally to superphosphate, but the clover responded to rock phosphate to a greater extent than did cape-weed. Both relative efficiency for total plant growth and percentage utilization of applied phosphorus were much higher with the soluble phosphatic fertilizer than with rock phosphate, especially in the first year. However, phosphorus recovery from rock phosphate was as high in the second year as in the first, whereas there was a marked decrease in the second year from superphosphate.

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