Use of growth curve derivatives to illustrate acceleration and deceleration of growth in young plantations under variable competition

2006 ◽  
Vol 36 (10) ◽  
pp. 2515-2522 ◽  
Author(s):  
Michael Newton ◽  
Elizabeth C Cole
Keyword(s):  
Growth Period ◽  
Tsuga Heterophylla ◽  
Douglas Fir ◽  
Early Years ◽  
Western Hemlock ◽  
Richards Equation ◽  
Second Derivative ◽  
Red Alder ◽  
Vegetation Control ◽  
Second Derivatives

Deceleration of growth rates can give an indication of competition and the need for thinning in early years but can be difficult to detect. We computed the first and second derivatives of the von Bertalanffy – Richards equation to assess impacts of density and vegetation control in young plantations in western Oregon. The first derivative describes the response in growth and the second derivative describes the change in growth over time. Three sets of density experiments were used: (i) pure Douglas-fir (Pseudotsuga menziesii (Mirb.) Franco), (ii) mixed Douglas-fir and grand fir (Abies grandis (Dougl. ex D. Don) Lindl.), and (iii) mixed western hemlock (Tsuga heterophylla (Raf.) Sarg.) and red alder (Alnus rubra Bong.). Original planting densities ranged from 475 to 85 470 trees·ha–1 (4.6 m × 4.6 m to 0.34 m × 0.34 m spacing); western hemlock and red alder plots were weeded and unweeded. For the highest densities, the second derivative was rarely above zero for any of the time periods, indicating that the planting densities were too high for tree growth to enter an exponential phase. As expected, the lower the density, the greater and later the peak in growth for both the first and second derivatives. Weeding increased the growth peaks, and peaks were reached earlier in weeded than in unweeded plots. Calculations of this sort may help modelers identify when modifiers for competition and density are needed in growth equations. Specific applications help define onset of competition, precise determining of timing of peak growth, period of acceleration of growth, and interaction of spacing and age in determination of peaks of increment or acceleration or deceleration.

Growth ofPseudotsuga menziesiiandTsuga heterophyllaseedlings along a light gradient: resource allocation and morphological acclimation

1997 ◽  
Vol 75 (9) ◽  
pp. 1424-1435 ◽  
Author(s):  
D. Mailly ◽  
J. P. Kimmins
Keyword(s):  
Leaf Morphology ◽  
Biomass Accumulation ◽  
Tsuga Heterophylla ◽  
Douglas Fir ◽  
Western Hemlock ◽  
Low Light ◽  
Growing Seasons ◽  
Light Levels ◽  
Light Gradient ◽  
Relative Light Intensity

Silvicultural alternatives that differ in the degree of overstory removal may create shady environments that will be problematic for the regeneration of Douglas-fir (Pseudotsuga menziesii (Mirb.) Franco). Gradients of light in the field were used to compare mortality, growth, and leaf morphological acclimation of two conifer species of contrasting shade tolerances: Douglas-fir and western hemlock (Tsuga heterophylla (Raf.) Sarg.). Results after two growing seasons indicated that Douglas-fir mortality occurred mainly at relative light intensity (RLI) below 20%, while western hemlock mortality was evenly distributed along the light gradient. Height, diameter, and biomass of the planted seedlings increased with increasing light for both species but at different rates, and maximum biomass accumulation always occurred in the open. Douglas-fir allocated more resources to stem biomass than western hemlock, which accumulated more foliage biomass. Increases in specific leaf area for Douglas-fir seedlings occurred at RLI ≤ 0.4 and red/far red (R/FR) ratio ≤ 0.6, which appear to be the minimal optimum light levels for growth. Conversely, western hemlock seedlings adjusted their leaf morphology in a more regular pattern, and changes were less pronounced at low light levels. These results, along with early mortality results for Douglas-fir, suggest that the most successful way to artificially regenerate this species may be by allowing at least 20% of RLI for ensuring survival and at least 40% RLI for optimum growth. Key words: light, light quality, leaf morphology, acclimation.

Survival dynamics of mechanically topped Douglas-fir (Pseudotsuga menziesii) and western hemlock (Tsuga heterophylla) snags in Douglas-fir plantations, Oregon, USA

2019 ◽  
Vol 433 ◽  
pp. 105-110
Author(s):  
Matthew E. Hane ◽  
Andrew J. Kroll ◽  
Aaron Springford ◽  
Jack Giovanini ◽  
Mike Rochelle ◽  
...  
Keyword(s):  
Pseudotsuga Menziesii ◽  
Tsuga Heterophylla ◽  
Douglas Fir ◽  
Western Hemlock

Canopy disturbances over the five-century lifetime of an old-growth Douglas-fir stand in the Pacific Northwest

2002 ◽  
Vol 32 (6) ◽  
pp. 1057-1070 ◽  
Author(s):  
Linda E Winter ◽  
Linda B Brubaker ◽  
Jerry F Franklin ◽  
Eric A Miller ◽  
Donald Q DeWitt
Keyword(s):  
Pacific Northwest ◽  
Variable Density ◽  
Tsuga Heterophylla ◽  
Douglas Fir ◽  
Western Hemlock ◽  
Old Growth ◽  
The Pacific ◽  
History Of ◽  
Horizontal Heterogeneity ◽  
Restoration Strategies

The history of canopy disturbances over the lifetime of an old-growth Douglas-fir (Pseudotsuga menziesii (Mirb.) Franco) stand in the western Cascade Range of southern Washington was reconstructed using tree-ring records of cross-dated samples from a 3.3-ha mapped plot. The reconstruction detected pulses in which many western hemlock (Tsuga heterophylla (Raf.) Sarg.) synchronously experienced abrupt and sustained increases in ringwidth, i.e., "growth-increases", and focused on medium-sized or larger ([Formula: see text]0.8 ha) events. The results show that the stand experienced at least three canopy disturbances that each thinned, but did not clear, the canopy over areas [Formula: see text]0.8 ha, occurring approximately in the late 1500s, the 1760s, and the 1930s. None of these promoted regeneration of the shade-intolerant Douglas-fir, all of which established 1500–1521. The disturbances may have promoted regeneration of western hemlock, but their strongest effect on tree dynamics was to elicit western hemlock growth-increases. Canopy disturbances are known to create patchiness, or horizontal heterogeneity, an important characteristic of old-growth forests. This reconstructed history provides one model for restoration strategies to create horizontal heterogeneity in young Douglas-fir stands, for example, by suggesting sizes of areas to thin in variable-density thinnings.

Plant Water Stress and Soil Depletion in Variable-Density, Red Alder/Western Hemlock Coastal Oregon Plantations

2019 ◽  
Vol 66 (3) ◽  
pp. 304-313
Author(s):  
Elizabeth C Cole ◽  
Michael Newton
Keyword(s):  
Water Stress ◽  
Summer Precipitation ◽  
Growing Season ◽  
Variable Density ◽  
Tsuga Heterophylla ◽  
Western Hemlock ◽  
Riparian Ecosystems ◽  
Replacement Series ◽  
Red Alder ◽  
The Impact

Abstract Riparian ecosystems provide critical habitat and functions while being some of the most productive areas in forests. Both conifers and hardwoods contribute to maintenance of habitat and function. To determine the impact of water stress on growth of red alder (Alnus rubra Bong.) and western hemlock (Tsuga heterophylla [Raf.] Sarg.), we installed Nelder type 1a combined with replacement series plots on three Oregon Coast Range sites. Densities ranged from 988 to 85,400 trees/hectare, with ratios (hemlock:alder) of 100:0, 75:25, 50:50, 25:75, and 0:100. In the first 4 years after planting, alder used water in the growing season at greater depths earlier than western hemlock. Higher densities resulted in greater water stress later in the growing season in weeded areas (maintained by herbicide applications), but stress was similar across densities in unweeded areas. Water stress at early ages was correlated with decreased size 14 or 24 years after planting for both species, but these correlations were confounded with other effects of density. Increasing water availability in areas with low summer precipitation could enhance growth of red alder and western hemlock, even in highly productive riparian areas.

Comparing productivity of pure and mixed Douglas-fir and western hemlock plantations in the Pacific Northwest

2006 ◽  
Vol 36 (6) ◽  
pp. 1484-1496 ◽  
Author(s):  
M M Amoroso ◽  
E C Turnblom
Keyword(s):  
Pacific Northwest ◽  
Stand Density ◽  
Tsuga Heterophylla ◽  
Douglas Fir ◽  
Western Hemlock ◽  
Mixed Stand ◽  
Mixed Stands ◽  
Individual Tree ◽  
The Pacific ◽  
Increase In Productivity

We studied pure and 50/50 mixtures of Douglas-fir (Pseudotsuga menziesii (Mirb.) Franco) and western hemlock (Tsuga heterophylla (Raf.) Sarg.) plantations to compare attained total yields between mixed-species stands as opposed to monocultures of equal densities. Whether overall stand density influences this outcome has not been adequately investigated, and to address this we included three density levels (494, 1111, and 1729 trees/ha) in the analysis. At age 12, as components of the mixed stands, Douglas-fir exhibited greater height, diameter, and individual-tree volume than western hemlock at all densities. At 494 and 1111 trees/ha the monocultures had a higher volume per hectare than the mixed stand, but at 1729 trees/ha the mixed stand appeared to be just as productive as the pure stands. The increase in productivity by the mixture at high densities seems to have resulted from the partial stratification observed and most likely also from better use of the site resources. Because of this, less interspecific competition was probably experienced in the mixed stand than intraspecific competition in the pure stands. This study shows the important role density plays in the productivity of mixed stands and thus in comparing mixed and pure stands.

Palynology and paleoecology of postglacial sediments in an anoxic basin, Saanich Inlet, British Columbia

1983 ◽  
Vol 20 (5) ◽  
pp. 873-885 ◽  
Author(s):  
Linda E. Heusser
Keyword(s):  
British Columbia ◽  
Pinus Contorta ◽  
Tsuga Heterophylla ◽  
Douglas Fir ◽  
Western Hemlock ◽  
Thuja Plicata ◽  
Alnus Rubra ◽  
Red Cedar ◽  
Saanich Inlet ◽  
Anoxic Basin

Varved, black clayey silts deposited in the marine waters of Saanich Inlet yield unusually abundant and diverse pollen assemblages derived from the coastal Douglas-fir (Pseudotsuga) and western hemlock (Tsuga heterophylla) forests of southwestern British Columbia. The 12 000 year palynological record chronicles the development of vegetation since ice left Saanich Inlet: the succession of pine (Pinus contorta) and alder (Alnus rubra) woodlands by forests characterized by Douglas-fir and oak (Quercus) and later by western hemlock and red cedar (Thuja plicata). Rapid deposition of annual layers of pollen, charcoal, and other terrigenous particles provides detailed evidence of changes in land use during the past few hundred years: settlement, logging, farming, and urbanization. Vegetational and climatic changes inferred from pollen spectra in the marine sediments of Saanich Inlet compare favorably with changes inferred from correlative pollen assemblages previously described from adjacent parts of Vancouver Island and the Fraser River valley.

Five-year growth responses of Douglas-fir, western hemlock, and western redcedar seedlings to manipulated levels of overstory and understory competition

2006 ◽  
Vol 36 (10) ◽  
pp. 2439-2453 ◽  
Author(s):  
Timothy B Harrington
Keyword(s):  
Growth Rate ◽  
Nitrogen Content ◽  
Douglas Fir ◽  
Western Hemlock ◽  
Volume Index ◽  
Stem Volume ◽  
Western Redcedar ◽  
Foliar Nitrogen ◽  
Relative Growth ◽  
Vegetation Control

Douglas-fir (Pseudotsuga menziesii (Mirb.) Franco), western hemlock (Tsuga heterophylla (Raf.) Sarg.), and western redcedar (Thuja plicata Donn ex D. Don) seedlings were planted in March 2001 within three clearcut-harvested, shelterwood, or thinned stands of mature Douglas-fir near Olympia, Washington. From 2002 to 2005, areas of vegetation control of 0, 4.5, or 9 m2 were maintained with herbicides around a total 162 seedlings per species. Photosynthetically active radiation (PAR) was 34%, 62%, and 100% of full sunlight in thinned stands, shelterwoods, and clearcuts, respectively. Effects of overstory level and vegetation control on seedling growth and resource availability generally were additive. Seedling stem volume index in clearcuts averaged four to eight times that observed in thinned stands, and with vegetation control, it averaged two to four times that observed without it. In thinned stands, relative growth rate of seedling stem volume index had a positive linear relationship with PAR (R2 = 0.38). Foliar nitrogen content of Douglas-fir explained 71% of the variation in relative growth rate. Factors explaining the most variation in foliar nitrogen content differed between thinned stands (PAR, R2 = 0.34) and clearcuts or shelterwoods (midday water potential, R2 = 0.63), suggesting that light and root competition, respectively, were the primary growth-limiting factors for these overstory levels.

Ionic balance and organic acids in western redcedar, western hemlock, and Douglas-fir seedlings grown in low- and high-N soils

1999 ◽  
Vol 29 (6) ◽  
pp. 669-678 ◽  
Author(s):  
J E Graff, Jr. ◽  
R K Hermann ◽  
J B Zaerr
Keyword(s):  
Organic Acids ◽  
Quinic Acid ◽  
Tsuga Heterophylla ◽  
Douglas Fir ◽  
Western Hemlock ◽  
Thuja Plicata ◽  
N Availability ◽  
Ionic Balance ◽  
Total N ◽  
Western Redcedar

Seedlings of western redcedar (Thuja plicata Donn ex. D. Don), western hemlock (Tsuga heterophylla (Raf.) Sarg.), and Douglas-fir (Pseudotsuga menziesii (Mirb.) Franco) were transplanted into soils with low and high levels of available NO3-(and total N). Current-year foliage was sampled after 10 weeks to determine the effect of N availability on foliar cation-anion balance (C-A) and the concentrations of low molecular weight organic acids of the three species. Carboxylate concentrations were estimated by using the difference between sums of cations and anions (C-A): 750 mequiv.·kg-1for western redcedar, 351 mequiv.·kg-1for western hemlock, and 266 mequiv.·kg-1for Douglas-fir. Quinic acid was a primary constituent, accounting for 40% of the total for western redcedar and 75% for western hemlock and Douglas-fir. Oxalic acid was present in greatest concentration in the foliage of western redcedar (65 mequiv.·kg-1) but was a minor constituent in western hemlock and Douglas-fir. The quantified acids accounted for only 15% of the C-A of western redcedar but >80% of the C-A of western hemlock and Douglas-fir. A considerable portion of the C-A balance not accounted for in redcedar may be associated with the accumulation of CaCO3. Litterfall deposition of CaCO3may lead to the consumption of H+ions and enrichment of exchangeable soil Ca in the rooting zone of long-lived western redcedar trees. No statistically significant differences among the soils were detected with regard to C-A or the concentration of organic acids.

Litter decomposition and nutrient release in Douglas-fir, red alder, western hemlock, and Pacific silver fir ecosystems in western Washington

1980 ◽  
Vol 10 (3) ◽  
pp. 327-337 ◽  
Author(s):  
Robert L. Edmonds
Keyword(s):  
Weight Loss ◽  
Nutrient Release ◽  
Douglas Fir ◽  
Western Hemlock ◽  
Silver Fir ◽  
Rapid Weight Loss ◽  
Red Alder ◽  
Cellulose Decomposition ◽  
The Pacific ◽  
Western Washington

Decomposition rates and changes in the nutrient content of needle and leaf litter were examined in Douglas-fir (Pseudotsugamenziesii Mirb. Franco), western hemlock (Tsugaheterophylla (Raf.) Sarg.), Pacific silver fir (Abiesamabilis (Dougl.) Forbes), and red alder (Alnusrubra Bong.) ecosystems in western Washington, U.S.A. Nylon litterbags (1-mm mesh) were placed in the stands in November and December 1974. Bags were collected after 3, 6, 12, and 24 months and weighed, except in the Pacific silver fir stand when bags were collected after 6, 9, 14, and 24 months. Litter was analyzed for C, N, P, K, Ca, Mg, Mn, lignin, and cellulose. Decomposition constants (k values) were determined. Fastest decomposition after 2 years occurred in red alder leaves, followed by Douglas-fir, western hemlock, and Pacific silver fir needles. There were significant differences in weight loss among species after 1 year, but no significant differences were evident after 2 years. Red alder leaves showed rapid weight loss in the 1st year but decomposed little in the 2nd year. Decomposition constants were highly positively correlated with minimum air temperatures and negatively correlated with C:N ratios. Low litter moisture tended to reduce decomposition in summer, particularly in the Pacific silver fir stand. Decomposition proceeded under snow in this ecosystem. The pattern of loss of elements from litterbags after 2 years varied from ecosystem to ecosystem, particularly for N. The following element mobility series resulted for the four ecosystems: red alder (K > Mg > Ca > P > N > Mn), Douglas-fir (K > P > Ca > Mg > Mn > N), western hemlock (K > Ca > Mg > N > Mn > P), and Pacific silver fir (K > Mg > Ca > Mn > P > N).

THE INITIATION AND SPREAD OF PORIA WEIRII ROOT ROT OF DOUGLAS FIR

1965 ◽  
Vol 43 (1) ◽  
pp. 1-9 ◽  
Author(s):  
G. W. Wallis ◽  
G. Reynolds
Keyword(s):  
Root Rot ◽  
Douglas Fir ◽  
Western Hemlock ◽  
Natural Soil ◽  
Limited Extent ◽  
Red Alder ◽  
Red Cedar ◽  
Considerable Resistance ◽  
Western Red Cedar ◽  
Living Tissues

Root rot caused by Poria weirii Murr. occurred when healthy roots of Douglas fir came into contact with inoculum in infected roots of the previous stand. Mycelium grew ectotrophically on the bark of the Douglas fir roots, frequently well in advance of growth in the wood, and penetrated to living tissues directly through sound as well as injured bark. Spread of the disease to adjacent trees took place where healthy and diseased roots were in contact, the mycelium apparently spreading to only a very limited extent through natural soil. It was shown that mycelium could invade roots of trees felled for at least 12 months and Douglas fir heartwood that had been buried in soil for at least 12 months. Viable Poria mycelium was isolated from infected roots as small as 2 cm in diameter 11 years after the trees had been cut. While Douglas fir and western hemlock appeared to be quite susceptible to infection, western red cedar, red alder, and bigleaf maple showed considerable resistance.

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