Urine marking at food and caches in captive coyotes
Urine marking of food and caches was studied in a group of captive coyotes, Canis latrans. During feeding observations, coyotes frequently urine marked the food pile and individual food items that had been carried and dropped some distance from the pile. In addition, food items which had been stolen were often marked. However, marking of food did not reserve it for the marking animal. Others usually ignored the urine mark and ate the item. The significant increase in urine marking of food during the breeding season suggested that it was involved in the expression of dominance in intrasexual rivalries. Cache marking was quite different. Urine marking never occurred when food was cached and rarely occurred while the cache still contained food. However, once the cache was emptied, urine marking usually occurred. Thus at caches, urine marking evidently serves a "book-keeping" role, indicating that the caches are no longer worth investigating although food odors might still linger.