scholarly journals Communities of the class Festuco-Brometea Br.-Bl. et Tx. ex Soó 1947 in Mesyagutovskaya and Krasnoufimskaya forest-steppes (the Middle Cis-Ural region)

2019 ◽  
pp. 29-78 ◽  
Author(s):  
N. V. Zolotareva ◽  
A. Yu. Korolyuk ◽  
S. M. Yamalov

There is a series of forest-steppe islands in the forest zone of the Middle Cis-Ural region. Two of these (Krasnoufimskaya and Mesyagutovskaya forest-steppes) of the considerable size and are of great interest for the study of steppe and meadow-steppe vegetation diversity in the northern locations of the Cis-Urals region. In the recent decades the studies of Mesyagutovskaya forest-steppe herbaceous vegetation with their syntaxonomy were done (Yamalov, Kucherova, 2009; Yamalov et al., 2013). For Krasno­ufimskaya forest-steppe such research was not carried out. Our aim was to make the revision of previously described syntaxa on this territory using the new data and to determine their position within the class Festuco-Brometea according to modern views. Existence of the forest-steppe landscape in the forest zone of Middle Cis-Urals region is due tothebarrier effect of the Ufa Plateau, intercepting part of the rainfall coming from the Atlantic transport. The climate of the study area is continental with moderately cold winter and moderately warm summer. Bedrocks here are limestones, sandstones and slates. The extrazonal northern forest-steppe extends for two degrees in the latitudinal direction. Mountainousness increases from the north to the south, while the climate becomes drier and warmer (Table 1). The central part of Krasnoufimskaya forest-steppe is flat; its agricultural development began in 17th century, so therefore the steppe areas were preserved only on the slopes of river valleys and hills. The territory of Mesyagutovskaya forest-steppe is more mountainous, therefore, the number of preserved steppe plots here is greater than in the Krasnoufimsk area (Fig. 2, 8). The initial dataset includes 295 relevés collected in the course of the field study in 2007–2014 in some locations of the Sverdlovsk Region and the Bashkor­tostan Republic. After the revision according to EuroVegChecklist Expert System (Mucina et al., 2016) the relevés previously assigned to the class Moli­nio-Arrhenatheretea were excluded, thus the data of this paper are based on the analysis of 260 relevés. All relevés were classified using the modified TWINSPAN algorithm (Roleček et al., 2009) in JUICE 7.0 (Tichý, 2002). According to TWINSPAN results the dataset was divided into 9 groups which formed two clusters (Fig. 1). Each of them is differentiated by group of species: cluster A — by the meso-xerophyte species of meadow steppes and xeric meadows, including 18 diagnostic species of the order Brachypodietalia pinnati (syn. Brometalia erecti) (Willner et al., 2017, 2019): Amoria montana, Brachypodium pinnatum, Carex caryophyllea, Dactylis glomerata, Festuca pratensis, Lathyrus pratensis, Leucanthemum vulgare, Ranunculus polyanthemos, Trifolium pratense, Vicia cracca, etc.; cluster B — by the widespread steppe species, including diagnostic ones of the class Festuco-Brometea (Anemone sylvestris, Artemisia latifolia, Campanula sibirica, Koeleria cristata, Scabiosa ochroleuca, Stipa capillata) and the order Helictotricho-Stipetalia (Helictotrichon desertorum, Onosma simplicissima, Oxy­tropis pilosa, Pilosella echioides, Potentilla humifusa, Salvia stepposa) (Korolyuk, 2017). Thus, cluster A combines meadow steppes and xeric meadows on rich soil habitats, and these communities correspond to the order Brachypodietalia pinnati. Cluster B combines petrophytic steppes on dry rocky habitats of insolated slopes, and these communities correspond to the order Helictotricho-Stipetalia. Ordination showed that such ecological factors as habitat moisture, soil fertility and rockiness are important in differentiation of syntaxa of the class Festuco-Brometea (Fig. 13). The vegetation of the class Festuco-Brometea in the northern forest-steppe of the Cis-Urals region belongs to 2 orders, 6 associations, 9 subassociations, 2 variants and 3 communities. Among them 1 association (Centaureo sibiricae–Stipetum pulcherrimae Yamalov ass. nov. hoc loco), 9 subassociations (Leucanthemo vulgaris–Stipetum pennatae typicum subass. nov. hoc loco, L. v.–S. p. festucetosum pratensis subass. nov. hoc loco, L. v.–S. p. vicietosum tenuifoliae subass. nov. hoc loco, Drabo sibiricae–Primu­letum macrocalycis typicum subass. nov. hoc loco, D. s.–P. m. potentilletosum humifusae subass. nov. hoc loco, Fragario viridis–Caraganetum fruticis poetosum angustifoliae Yamalov subass. nov. hoc loco, Stipo pennatae–Centauretum sibiricae typicum subass. nov. hoc loco, S. p.–C. s. polygonatetosum odorati subass. nov. hoc loco, S. p.–C. s. polygaletosum sibiricae subass. nov. hoc loco) and 3 communities (Spiraea crenata, Festuca valesiaca–Alyssum tortuosum, Anthemis subtinctoria–Aster alpinus) are new. One association (Stipo pennatae–Centauretum sibiricae Yamalov et al. ex Zolotareva, Korolyuk, Yamalov ass. nov. hoc loco) has been validated. Syntaxa of the order Brachypodietalia pinnati represent the base of the class Festuco-Brometea in the study area. Currently, only meadow steppes of the ass. Leucanthemo vulgaris–Stipetum pennatae are widespread throughout the extrazonal forest-steppe. Steppe-meadows of the ass. Drabo sibiricae–Primuletum macrocalycis) are found in the Mesyagutovskaya area; in the northern part the same landscape positions are occupied by communities of the class Molinio-Arrhenatheretea. The steppe shrubs in the northern and the southern parts of the forest-steppe area are also different: in the south these are dominated by Caragana frutex and corresponds to the ass. Fragario viridis–Caraganetum fruticis, in the north (Krasnoufimskaya area) Caragana frutex is not found and shrub communitiesaredominated by Spiraea crenata (Fig. 2). Syntaxa of the order Helictotricho-Stipetalia occur on limestones exposures. Currently, the petrophytic steppes of the ass. Stipo pennatae–Centauretum sibiri­cae are known overall in the extrazonal forest-steppe, while the communities of the ass. Centaureo sibiricae–Stipetum pulcherrimae are registered only in the southern part of Mesyagutovskaya forest-steppe; northwards locations of this association diagnostic species are rare, some of these are absent in the Krasnoufimskaya forest-steppe area. The other syntaxa of petrophytic steppes with many relict and endemic species occur locally on the slopes of ancient reeves due to the unique combination of factors (Fig. 8).

2002 ◽  
Vol 57 (3) ◽  
pp. 391-400 ◽  
Author(s):  
Anastasia K. Markova ◽  
Alexandra N. Simakova ◽  
Andrei Yu Puzachenko ◽  
Lev M. Kitaev

AbstractAbundant mammal (45 localities) and plant (52 sections) fossil data collected from the Russian Plain deposits of the Middle Valdai Briansk Interstade indicate a period of noticeable warming. These materials were jointly analyzed. The data were organized in the database software PARADOX and then moved to the GIS program ARC/INFO. Mathematical methods as well as traditional research methods were used for the analysis. The reconstructed biogeographical provinces illustrate the specific environmental and climatic conditions of the Briansk Interstade. Analogues of modern natural zones did not exist at this time on the Russian Plain. The Briansk landscapes reflect the moderate-cool climate of this period, when the majority of northern subarctic plant and mammal species extended their ranges considerably to the south to the Central Russian Plain. At the same time, steppe species penetrated farther north and west. This combination shows that the continuous forest zone was destroyed during the Briansk Inter-stade. Five biogeographical provinces have been reconstructed from the north to the south for the Russian Plain during the Briansk warming on the basis of joint analysis of mammalian and botanical data.


Author(s):  
V. P. Starikov

Steppe lemming of the South Trans-Ural region is a rare species of open spaces. To the north, its distribution is limited to the subzone of herbbunchgrass steppes and the southern strip of the forest-steppe subzone. The presented work summarizes the material, which gives the full species composition of the parasitic arthropods of this rodent, and gives quantitative characteristics of ectoparasites. In total 155 individuals and one nest of steppe lemming were examined for ectoparasites. There are 808 parasitic arthropods of 20 species, including 14 species of gamasid mites, 2 species of ixodid ticks and 4 species (subspecies) of fleas. According to the number of individuals of ectoparasites feeding on the steppe lemming, the dominance of gamasid mites is strongly pronounced, both prevailing multi-host species and specific, typical for other rodents (gray and red-backed voles, northern mole vole, Eurasian water vole, northern birch mouse). The contribution of ixodid ticks and fleas to the parasitofauna of the steppe lemming is small. Steppe lemming can be involved in circulation of a causative agent of tularemia in the South Trans-Ural region, but as an additional carrier in maintenance of a foci in biotopes, where it contacts the habitats of the Eurasian water vole.


2016 ◽  
Vol 3 (3) ◽  
pp. 325-331
Author(s):  
Жигилева ◽  
O. Zhigileva ◽  
Усламина ◽  
I. Uslamina

Objective of research: to study the nematode infestation of the sable and pine marten in Western Siberia. Materials and methods: Altogether 169 individuals of the sable and 18 — of the pine marten were investigated using the method of partial dissection of the gastrointestinal tract and lungs. Animals were caught according to the hunting license in 2009-2011 and 2014-2015, in 8 areas of Western Siberia from the south taiga to the northern forest-steppe. Results and discussion: We have found four species of parasitic nematodes, two of which (Crenosoma petrovi, Filaroides martis) were localized in lungs and two species (Capillaria putorii, Strongyloides martis) — in the intestine of the sable and the pine marten. Prevalence of lung nematodes was 80%, intestine nematodes — 40%. Intensity of infestation ranged from 26 to 358 nematodes per animal. F. martis is the most pathogenic and most common helminth in martens in Western Siberia. 53,3% of pine martens and 26,5% of sables were infected with this nematode species. Nematode S. martis is the most abundant in the intestine. In pine martens the extensity of invasion was 42,8%, the abundance index — 8.29 helminths per animal; in sables — 9,1% and 1.18, respectively. Pine martens were infected by F. martes and S. martis more than sables (p — 0,05). Prevalence increases in the direction from the North East (taiga) to the South West (sub-taiga), which determines the intensity of the helminthiases focus in biocenoses of Western Siberia. Infection of pine martens in the forest-steppe was not detected. In 2014–2015 compared with the previous study period (2009–2011) the extensity of invasion with intestinal nematodes increased by 1,5–2 times, and with highly pathogenic lung parasites — by 3 times. The increase in the helminth infestation may be explained by the growth of animal population density.


Author(s):  
V. A. Sapega

The assessment of productivity and the parameters of adaptability allowed to use and perspective mid-season and mid-late varieties of spring wheat in the conditions of the Northern Trans-Ural region carried out. The varieties were tested in 2014-2016 on the steam predecessor in three climatic zones: subtaiga (II zone), northern forest-steppe (III zone) and southern forest-steppe (IV zone). The greatest interaction a genotype-environment is noted when calculating correlation dependence between productivity of varieties in various years of their test in the conditions of the southern forest-steppe zone (from r=-0,07±0.28 to r=0,41±0,25). Assessment of productivity and parameters of adaptability of spring wheat varieties in 9 environments (3 years x 3 GSU) has shown that Aviada allowed to use characterized the greatest maximum and average productivity in group of mid-season varieties (respectively 5,86 and 3,78 t/hectare), and in the mid-late group – a perspective Quintus (respectively 6,98 and 4,58 t/hectare). The considerable variability of productivity is noted.  Its size was from 23,0 % (Ikar) to 29,9 % (Aviada) in group of mid-season varieties, and at the mid-late group – from 25,6 % (Riks) to 31,6 % (Quintus). Resistance to stress of varieties was low irrespective of group of their ripeness. Ikar (-2,32) had the greatest indicator of resistance to stress in group mid-season varieties, and in the mid-late group – Riks (-2,54). Aviada (4,24 t/hectare) was characterized the greatest average productivity in contrast conditions in mid-season group and in the mid-late group – Quintus (4,73 t/hectare). Three groups of varieties were allocated on responsiveness for change of conditions:  high responsiveness (mid-season – the Lutescens 70, Aviada; mid-late – the Melody, Quintus), plastic (mid-season – Chernyava 13, Skent 3, Omskaya 36, Tyumenskaya 25, Tyumenskaya 29; mid-late – Ingala, Yamalskaya) and poorly responsiveness (mid-season – Ikar, Sigma; mid-late – Riks, the Stepnaya Niva). The low gomeostasis is revealed at all varieties irrespective of group of ripeness. The variety Ikar (Hom=6,59) was the best on a gomeostasis in mid-season group, and at the mid-late group – Riks (Hom=5,48). In general all varieties were characterized by the low general adaptive ability (GAA). Its rather high values in group of mid-season varieties are revealed at Aviada (OAC=0,19), and at the  mid-late group – Quintus (OAC=0,50). Tyumenskaya 25 and Tyumenskaya 29 are recognized the best mid-season varieties in the conditions of the Northern Trans-Ural region on the basis of complex assessment of productivity and parameters of adaptability, and at the mid-late group – Melody and Ingala.


Author(s):  
Aleksander Tairov

The paper publishes new materials of the early Sarmatian time from the North-East periphery of the Sarmatian World – the Southern forest-steppe and steppe zones of the Trans-Ural region within the bounds of the modern Chelyabinsk region. The territory of the Southern forest-steppe is represented by the materials of the ruined burial, which is situated on the shore of the lake Smolino within the bounds of Chelyabinsk. It is dated by the early 4th century BC. There are three accidental finds from this region: iron daggers as well. The daggers with curved bar-shaped pommels and arcuate cross-guards are classified as daggers of “transitional type” and are dated by the 4th century BC. The third dagger with a semicircular pommel and a straight cross-guard is included in the group of classical Prokhorovka daggers and is dated the 3rd – 2nd centuries BC. This paper presents the materials of two peculiar burial complexes from the steppe zone of the Southern Trans-Urals. The burial near Mogutovka village is the only one burial (without a barrow) of the early Sarmatian time, which was investigated in the Southern Trans-Ural region. It is situated on the first terrace above the floodplain of the Kamysty-Ayat river. The localization of the burial is not typical for the sites of the early Trans-Ural nomads of the 1st millennium BC. The early Sarmatian burial of the 3rd – 2nd centuries BC was excavated in Druzhinskiy burial site. It was made in the burial pit, which combines signs of a catacomb and an alcove. A child buried here was dressed in a shirt, whose collar and sleeves were decorated with beading. Publishing materials will help to expand the point of view on burial rites and material culture of the early Trans-Ural nomads of the early Sarmatian time, which exists in scientific literature.


2015 ◽  
pp. 129-147
Author(s):  
M. P. Tishchenko ◽  
A. Yu. Korolyuk ◽  
N. I. Makunina

The Tobol and the Irtysh watershed occupy the western part of West-Siberian plain. Area studied covers latitudinal band between 55° and 57° n. l. It includes the southern part of subtaiga subzone and northern part of forest-steppe zone. This territory has been used for agriculture since XVIIth century. In the middle of XXth century the local landscapes were represented by endless arable lands alternated with tiny patches of forests on the watersheds and swamps and solonchaks prevailing at lowlands. In the end of XXth century the large areas of arable lands, especially in northern part of mentioned area, were abandoned. Nowadays this area is covered by fallow meadows on different stages of demutation. Vegetation studies of the region have started in the end of XIXth century. One of the best scientific works on local grasslands was written by B. N. Gorodkov (1915). The spectrum of natural plant communities demonstrates some changes fr om south to north. These changes enclose both zonal and intrazonal vegetation. The southern part of the area studied belongs to forest-steppe zone. Its zonal vegetation is represented by small-leaved forests, grasslands (steppe meadows) and meadow steppes. Zonal vegetation of northern part — subtaiga subzone — includes only small-leaved forests. The watershed grasslands replace the cut-off forests. Intrazonal vegetation of southern part consists of various grasslands on salty soils and swamps predominated in the northern part. The border of forest-steppe and subtaiga represents a combination of zonal and intrazonal communities disturbed by man. The goal of this article is to reveal the diversity of grasslands on drained soils of mentioned area. The analysis of 151 geobotanical relevйs obtained fr om subtaiga and northern forest-steppe of Omskaya and Tyumenskaya administrative districts has been carried out. The syntaxonomy of the Tobol-and-the Irtysh watershed grasslands is represented by 6 associations, 4 subassociations and 2 communities of 4 alliances, 4 order and 2 classes. Subtaiga and northern forest steppe are charac­terized by different spectra of grasslands. This can be explained by a presence of wide range of saline soils in forest-steppe and their absence in subtaiga. The floristic composition of grasslands of forest-steppes zone includes the plants tolerant to soil salinity such as Artemisia pontica, Galatella biflora, Plantago maxi­ma, Eryngium planum, Saussurea amara, Limonium gmelinii. The listed species are rare in subtaiga grasslands while Festuca pratensis, Phleum pratense, Agrostis gigantea of alliance Festucion pratensis are dominated in subtaiga grassland meadows. Steppe meadows of this territory are represented by two classes. Northern forest-steppe subzone demonstrates the northern lim it of Festuco-Brometea (alliance Galatellion biflorae of order Festucetalia valesiacae). The associations Galatello biflorae–Calamagrostietum epigeii and Galio borealis–Artemisietum ponticae occupy the zonal positions. Their communities are characterize by high activity of meadow-steppe mesoxerophytes that are the diagnostic of the class Festuco-Brometea. The meadow mesophytes dominate in subtaiga steppe meadows absolutely; meadow-steppe species are rare. They are marked with a lack of diagnostic species of alliance Galatellion biflorae. They are ascribed to the class Molinio–Arrhenatheretea, order Galietalia veri and alliance Trifolion montani as the communities Filipendula stepposa–Cenolophium denudatum and Astragalus danicus–Potentilla anserinа. Forest steppe meadows of alliance Aconito barbati–Vicion unijugae (order Carici macrou­rae–Crepidetalia sibiricae, class Molinio-Arrhena­theretea) are usual for northern, subtaiga subzone of the Tobol and the Irtysh watershed. Diagnostic species list of the communities belonging to the order Carici macrourae–Crepidetalia sibiricae on West-Siberian plain is impoverished in comparison with the uplands and the mountains of South Siberia wh ere this order was described by Ermakov et al. (1999). We have revealed 2 new associations of alliance Aconito barbati–Vicion unijugae: Aegopodio podagrariae–Brachypodietum pinnati and Dracocephalo ruyschiani–Heracleetum sibiricae. In subtaiga subzone communities of these associations occupy the small well drained habitats between mesophytic forests and arable lands. The first association unites the xerophytic forest meadows with Aegopodium podagraria dominance, occurring only on right bank of the Ishim River. The second one includes forest meadows of subtaiga widely spread on the Tobol and the Irtysh watershed. Association Dracocephalo ruyschiani–Heracleetum sibiricae comprises two subassociations: D. r.–H. s. typicum and D. r.–H. s.molinietosum caeruleae. One of diagnostic species of the last mentioned subassociation is Molinia caerulea that is a common species of boreal wet meadows of Europe. The eastern border of this species distribution is on the left riverside of the Ishim River. Meadows of alliance Festucion pratensis of order Arrhenatheretalia are characteristic only for subtaiga, they belong to two associations. Meadows of subassociation Cirsio setosi–Phleetum pratensis pastinacetosum sylvestris are widespread. They are wide spread over the abandoned arable lands and rather monotonous by their floristic composition. The floristic peculiarity of plant communities is abundance of diagnostic species of classes Artemisietea и Chenopodietea. Meadows of Agrostio tenuis–Festucetum pratensis Yamalov 2005 occur on sandy soils; they present one of the first stages of pine forest demutation. Some of species are not characteristic for typical subassociation described by S. M. Yamalov (2005), therefore the meadows of this association on the Tobol and the Irtysh watershed are referred to a new subassociation A. t.–F. p. senecionetosum jacobaea.


1922 ◽  
Vol 59 (10) ◽  
pp. 433-441 ◽  
Author(s):  
Henry Fairfield Osborn

In studying the evolution and distribution of the Proboscidea and the arrival of man in Great Britain (Osborn, 1922, 1922a, 1922b), the writer has recently had occasion to review the faunal researches of Lydekker and Newton, and the collection of Mr. A. C. Savin, in connexion with the very interesting question of the geologic correlation of Great Britain with the Upper Pliocene fauna of Europeto the south-east, and of the Scandinavian Glaciation I on the north, with the advent of the northern forest, the boreal, and the Arctic mammal fauna of Great Britain.


2020 ◽  
Vol 53 (3) ◽  
pp. 283-293
Author(s):  
R. R. Suleymanov ◽  
V. V. Ovsyannikov ◽  
A. G. Kolonskih ◽  
E. V. Abakumov ◽  
A. Ya. Kungurtsev ◽  
...  

Author(s):  
S. L. Zhygalova

According to our own observations, herbarium materials and literature data, the distribution of Iris pineti- cola Klokov — an endemic forest-steppe species included in the Red Data Book of Ukraine in the status of “vulnerable” was analyzed. Data on its taxonomic status are given, distribution map is made (geographical coordinates are restored on the basis of materials of the National Herbarium of Ukraine (KW), herbarium of V. N. Karazin Kharkiv National University (CWU), as well as literature data), range and population data are analyzed. It is established that I. pineticola is distributed in the Right-Bank and Left-Bank Forest-Steppe, occasionally entering the north part of the steppe and the valley of the Siversky Donets. Grows in forests, artificial pine plantations on sandy terraces of rivers. According to own data and analyzed literature, I. pineticola populations are stable. In addition to inclusion in the Red Data Book of Ukraine, at the regional level I. pineticola is protected in Donetsk and Luhansk regions of Ukraine, also protected in the territories of NPP “Holy Mountains” and landscape reserve of national importance “Red Coast”.


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