Proboscidea from the Baynunah Formation
The Baynunah Formation provides the only late Miocene record of fossil proboscideans from the Arabian Peninsula, identifiable as a deinothere, tetralophodont gomphothere(s), and stegotetrabelodont elephant. Ichnofossils show the oldest evidence for modern-type elephant herd structure. The elephant sample is particularly abundant, and newly-recovered fossils yield critical evidence about their systematics and biology. Comparative results show that the Baynunah stegotetrabelodont belongs in a new species, and that in many ways it is the most primitive of all elephants —supporting a geological age of 8.0-6.0 Ma—consistent with having evolved in Afro-Arabia from Tetralophodon between 9.0-8.0 Ma. Body size analyses reveal that these elephants were exceptionally tall and heavy, possibly exceeding 10,000 kg. Tooth emergence is reconstructed as dp2-dp3-dp4-p3-m1-p4- m2-m3. The taxonomic composition and dietary preferences of the Baynunah proboscideans indicate the presence of heterogeneous local habitats in the region during the late Miocene, though the high abundance of the stegotetrabelodont, and the rarity of the deinothere and gomphothere(s) indicate a dominance of grassland habitats and a lesser representation of wooded areas.