scholarly journals Integrative taxonomy ofMetrichiaRoss (Trichoptera: Hydroptilidae: Ochrotrichiinae) microcaddisflies from Brazil: descriptions of twenty new species

PeerJ ◽  
2016 ◽  
Vol 4 ◽  
pp. e2009 ◽  
Author(s):  
Allan P.M. Santos ◽  
Daniela M. Takiya ◽  
Jorge L. Nessimian
Keyword(s):  
New Species ◽  
Dna Sequences ◽  
Mitochondrial Gene ◽  
Dna Barcode ◽  
First Record ◽  
Integrative Taxonomy ◽  
Morphological Data ◽  
Mato Grosso ◽  
Intraspecific Divergence ◽  
Coi Sequences

Metrichiais assigned to the Ochrotrichiinae, a group of almost exclusively Neotropical microcaddisflies.Metrichiacomprises over 100 described species and, despite its diversity, only one species has been described from Brazil so far. In this paper, we provide descriptions for 20 new species from 8 Brazilian states:M. acuminatasp. nov.,M. azulsp. nov.,M. bonitasp. nov.,M. bracuisp. nov.,M. caracasp. nov.,M. circuliformesp. nov.,M. curtasp. nov.,M. farofasp. nov.,M. forcepssp. nov.,M. formosinhasp. nov.,M. goianasp. nov.,M. itabaianasp. nov.,M. longissimasp. nov.,M. peludasp. nov.,M. rafaelisp. nov.,M. simplessp. nov.,M. talhadasp. nov.,M. teresp. nov.,M. ubajarasp. nov., andM. vulgarissp. nov.DNA barcode sequences (577 bp of the mitochondrial gene COI) were generated for 13 of the new species and two previously known species ofMetrichiaresulting in 64 sequences. In addition, COI sequences were obtained for other genera of Ochrotrichiinae (Angrisanoia,Nothotrichia,Ochrotrichia,Ragatrichia, andRhyacopsyche). DNA sequences and morphological data were integrated to evaluate species delimitations. K2P pairwise distances were calculated to generate a neighbor-joining tree. COI sequences also were submitted to ABGD and GMYC methods to assess ‘potential species’ delimitation. Analyses showed a conspicuous barcoding gap amongMetrichiasequences (highest intraspecific divergence: 4.8%; lowest interspecific divergence: 12.6%). Molecular analyses also allowed the association of larvae and adults ofMetrichia bonitasp. nov.from Mato Grosso do Sul, representing the first record of microcaddisfly larvae occurring in calcareous tufa (or travertine). ABGD results agreed with the morphological delimitation ofMetrichiaspecies, while GMYC estimated a slightly higher number of species, suggesting the division of two morphological species, each one into two potential species. Because this could be due to unbalanced sampling and the lack of morphological diagnostic characters, we have maintained these two species as undivided.

scholarly journals Integrative taxonomy of the new millipede genus Coxobolellus, gen. nov. (Diplopoda : Spirobolida : Pseudospirobolellidae), with descriptions of ten new species

2020 ◽  
Author(s):  
Piyatida Pimvichai ◽  
Henrik Enghoff ◽  
Somsak Panha ◽  
Thierry Backeljau
Keyword(s):  
New Species ◽  
New Genus ◽  
Phylogenetic Analyses ◽  
Mitochondrial Gene ◽  
Sequence Divergence ◽  
Morphological Characters ◽  
Integrative Taxonomy ◽  
Morphological Data ◽  
Morphological Evidence ◽  
Morphological Differences

Pseudospirobolellidae is a poorly known family of spirobolidan millipedes with only two genera and five described species. Yet, the descriptive taxonomy and molecular systematics of this group have been largely neglected. Therefore, the present work presents an integrative taxonomic study of new pseudospirobolellid taxa in Thailand. To this end, two mitochondrial gene fragments (COI and 16S rRNA) combined with morphological characters were used to define the genus Coxobolellus, gen. nov. with 10 new species, viz. C. albiceps, sp. nov., C. compactogonus, sp. nov., C. fuscus, sp. nov., C. nodosus, sp. nov., C. serratus, sp. nov., C. simplex, sp. nov., C. tenebris, sp. nov., C. tigris, sp. nov., C. transversalis, sp. nov. and C. valvatus, sp. nov. The interspecific COI sequence divergences among the new species ranged from 6 to 15%. The intergeneric COI sequence divergence between species of Coxobolellus, gen. nov., Benoitolus birgitae and Pseudospirobolellus sp. ranged from 20 to 23%. Three major morphological differences separate Coxobolellus, gen. nov. from Benoitolus and Pseudospirobolellus, namely (1) the protruding process on the 3rd (and 4th) coxae on male legs, (2) the posterior gonopod telopodite divided into two parts, and (3) a conspicuous opening pore at the mesal margin at the end of the coxal part of the posterior gonopod. Thus, the new genus is well supported by both mtDNA and morphological evidence, while the delimitation of the 10 new species is supported by the congruence between mtDNA and morphological data. Yet, with respect to the relationships of Benoitolus birgitae, morphological data suggest a similarity with Coxobolellus, gen. nov. and Pseudospirobolellus, whereas mtDNA data place this species in the Pachybolidae. Further phylogenetic analyses are needed to explore this apparent incongruence and test the monophyly of Pseudospirobolellidae.

scholarly journals Accelerating the knowledge of Peruvian Chalcididae (Insecta,  Hymenoptera, Chalcidoidea) with integrative taxonomy

2019 ◽  
Vol 7 ◽  
Author(s):  
Bruno Cancian de Araujo ◽  
Marcelo Tavares ◽  
Thales Renan Brotto ◽  
Juliana Silva-Freitas ◽  
Max Estefani Santos ◽  
...  
Keyword(s):  
New Species ◽  
Dna Barcoding ◽  
Dna Sequences ◽  
Fold Increase ◽  
Integrative Taxonomy ◽  
Morphological Data ◽  
Peruvian Amazon ◽  
Insect Biodiversity ◽  
Canopy Fogging ◽  
Molecular Dataset

We present the first regional inventory of the fauna of Chalcididae in the Peruvian Amazon, with a nearly 6-fold increase in the number of species recorded for the country. A total of 418 specimens of Chalcididae were collected between 2000 and 2017 at the Panguana Reserve, Peruvian Amazon, 400 of which were obtained using Malaise traps and the remaining 18 specimens by canopy fogging. The morphological analyses indicated that these specimens represent 183 species of Chalcididae in 10 different genera, with 173 new to Peru and 134 potentially new species. We submitted 268 specimens, representing 167 species, to DNA barcoding. Of these, 141 specimens yielded sequences, 136 of them with a minimum of 300 bp. Sixty specimens were assigned a BIN by the Barcode of Life Database System (BOLD), resulting in 50 BINs. A cluster analysis of 138 individuals that yielded DNA sequences longer than 100 bp revealed 118 MOTUs (molecular operative taxonomic units), all of them highly congruent with the morphological data. Prior to the present study, 37 species in 9 genera of Chalcididae were known from Peru. With our results, this number was increased to 210 species in 13 genera. The present study is the result of a joint effort between the SNSB - Zoologische Staatssammlung München, Germany (ZSM) and the Insect Biodiversity Laboratory of the Universidade Federal do Espírito Santo, Vitória, Brazil (LaBI-UFES), intending to apply an accelerated taxonomic treatment of the Chalcididae of the Panguana reserve using traditional morphological approaches in combination with DNA barcoding. The complete molecular dataset and associated voucher information is publicly available through BOLD. The new species that were discovered as part of the study are being formally described elsewhere as part of taxonomic treatments of Neotropical and world generic revisions at LaBI-UFES.

Uncovering the hidden biodiversity of natural history collections: Insights from DNA barcoding and morphological characters of the Neotropical genus Orthocomotis Dognin (Lepidoptera: Tortricidae)

Zootaxa ◽  
2017 ◽  
Vol 4250 (6) ◽  
pp. 541 ◽  
Author(s):  
JÓZEF RAZOWSKI ◽  
VOLKER PELZ ◽  
SEBASTIAN TARCZ
Keyword(s):  
Natural History ◽  
Host Plants ◽  
Mitochondrial Gene ◽  
Dna Barcode ◽  
Molecular Study ◽  
Morphological Characters ◽  
Morphological Data ◽  
Rapid Speciation ◽  
Natural History Collections ◽  
Coi Sequences

We used a 227-bp fragment of the mitochondrial gene cytochrome oxidase I (DNA “barcode”) in conjunction with morphological data to study specimens of the Neotropical genus Orthocomotis Dognin, 1906, acquired from natural history collections. We examined over 20 species of Orthocomotis from 17 localities in Colombia, Ecuador, and Peru. The analysis identified 32 haplotypes among the 62 specimens and found no haplotypes shared among species. The molecular study revealed not only the usefulness of short COI sequences in discriminating among Orthocomotis species but also showed distinctness of four clusters which correspond to those based on morphological (genitalia) characters. Moreover, the molecular results suggest the occurrence of rapid speciation in Orthocomotis. We hypothesize that this may be linked to the great biodiversity of potential host plants in Neotropical ecosystems. 

Chameleonfishes in Bangladesh: hipshot taxonomy, sibling species, elusive species, and limits of species delimitation (Teleostei: Badidae)

Zootaxa ◽  
2019 ◽  
Vol 4586 (2) ◽  
pp. 301 ◽  
Author(s):  
SVEN KULLANDER ◽  
MICHAEL NORÉN ◽  
MD. MIZANUR RAHMAN ◽  
ABDUR ROB MOLLAH
Keyword(s):  
New Species ◽  
Dna Sequences ◽  
Species Delimitation ◽  
Genetic Difference ◽  
Morphological Data ◽  
Valid Species ◽  
Colour Pattern ◽  
Stabilizing Selection ◽  
Difference Threshold ◽  
Coi Sequences

Five species of Badidae are reported from Bangladesh, with morphological diagnoses and mitochondrial DNA sequences (cytochrome b, cytb; and cytochrome c oxidase subunit I, coi). Dario kajal is recorded from Bangladesh for the first time with a precise locality. Badis badis is reported from several localities in central Bangladesh. Badis chittagongis is redescribed on the basis of samples from the region of Cox′s Bazar, including Maheskhali Island. Badis pallidus, new species, is described from the Sangu and Karnafuli River drainages in Bangladesh. It is most similar to B. chittagongis, but differs slightly in colouration and meristics, and is separated by 3.8% uncorrected p-distance in coi from B. chittagongis. Badis chittagongis and B. pallidus are almost identical in morphology, colour pattern and meristics, but occupy different habitats and are reciprocally allopatric. Pronounced genetic difference but similar morphology in these two species may be due to strong stabilizing selection for cryptic colouration in Badis. Badis rhabdotus is a new species from northeastern Bangladesh and adjacent Meghalaya in India. It is distinguished from congeneric species by the colour pattern, including well-defined narrow vertical bars; posterior bars curved; and meristics. Species delimitation analysis of an alignment comprising all coi sequences available from GenBank longer than 600 bp and attributed to species of Badidae (21 June 2018) plus our coi sequences and outgroup sequences of Nandus nandus, using pairwise p-distance and the computer software GMYC, ABGD, and bPTP, produced similar results. Among 103 coi sequences of Badidae, unidentified or tagged with one of 18 valid species names, uncorrected p-distance suggests 27 OTUs at 2% difference threshold; ABGD found between 15 and 55 OTUs; GMYC with single evolutionary rate 33 OTUs, with multiple evolutionary rates 32 OTUs; PTP, mPTP and bPTP 27–28 OTUs. Phylogenetic analysis based on coi and cytb sequences support previous analyses and previously proposed species groups. Inadequate recent species descriptions and many misidentifications or provisional identifications of published DNA sequences hamper progress in species-level systematics in Badis. Based on published morphological data, Badis triocellus cannot be distinguished from B. singenensis; Badis dibruensis and B. pancharatnaensis cannot be distinguished from B. badis; Badis andrewraoi, B. autumnum, B. kyanos, and B. soraya are insufficiently well distinguished from each other. 

Isostenosmylus ammirabilis sp. nov., a remarkable new species of lance lacewing (Neuroptera: Osmylidae) from the Colombian Andes

Zootaxa ◽  
2020 ◽  
Vol 4803 (3) ◽  
pp. 561-575
Author(s):  
ADRIAN ARDILA-CAMACHO ◽  
CALEB CALIFRE MARTINS ◽  
JORGE ARI NORIEGA
Keyword(s):  
New Species ◽  
Cytochrome C ◽  
Cytochrome C Oxidase ◽  
Mitochondrial Gene ◽  
Dna Barcode ◽  
Neotropical Region ◽  
Andean Region ◽  
Oxidase Subunit ◽  
Colombian Andes ◽  
South Of Brazil

Isostenosmylus Krüger, 1913 is the richest genus of Osmylidae of the Neotropical region, with 17 described species so far, which are distributed mainly in the Andean region and in the South of Brazil and Paraguay. A new remarkable Colombian species of Isostenosmylus—I. ammirabilis sp. nov.—is herein described and illustrated. DNA barcode of mitochondrial gene cytochrome c oxidase subunit I (COI) for this species is also provided. Taxonomic keys for the genus are updated. 

scholarly journals Integrative taxonomy of Nearctic and Palaearctic Aleocharinae: new species, synonymies, and records (Coleoptera, Staphylinidae)

ZooKeys ◽  
2021 ◽  
Vol 1041 ◽  
pp. 27-99
Author(s):  
Adam J. Brunke ◽  
Mikko Pentinsaari ◽  
Jan Klimaszewski
Keyword(s):  
New Species ◽  
North America ◽  
Dna Barcode ◽  
Integrative Taxonomy ◽  
Eastern Canada ◽  
Rove Beetles ◽  
Adventive Species ◽  
The Status ◽  
Taxonomic Studies ◽  
Holarctic Species

A long tradition of separate Nearctic and Palaearctic taxonomic studies of the diverse aleocharine rove beetles (Coleoptera: Staphylinidae) has obscured the recognition of Holarctic species and detection of adventive species in both regions. Recently, integrated study of the two regions through detailed morphological comparisons and development of an authoritatively identified DNA barcode reference library has revealed the degree to which these two aleocharine faunas are interconnected, both naturally and through human activity. Here this approach is adopted to recognize new species, reveal Holarctic species, and recognize adventive species in both North America and Europe. The following new species are described: Isoglossa triangularis Klimaszewski, Brunke & Pentinsaari, sp. nov. from British Columbia; Gnypeta impressicollis Klimaszewski, Brunke & Pentinsaari, sp. nov., from Ontario, Maryland and North Carolina; Aloconota pseudogregaria Klimaszewski, Brunke & Pentinsaari, sp. nov., from Ontario and Virginia; and Philhygra pseudolaevicollis Klimaszewski, Brunke & Pentinsaari, sp. nov. from eastern Canada. Dasygnypeta velata and Philhygra angusticauda are revealed to be Holarctic species, resulting in the following synonymies: Dasygnypeta velata (Erichson, 1839) = Gnypeta minuta Klimaszewski & Webster, 2008, syn. nov. and Philhygra angusticauda (Bernhauer, 1909) = Atheta (Philhygra) pinegensis Muona, 1983, syn. nov. The Nearctic species Hylota ochracea (and genus Hylota), Thecturota tenuissima, and Trichiusa robustula are newly reported from the Palaearctic region as adventive, resulting in the following synonymies: Hylota ochracea Casey, 1906 = Stichoglossa (Dexiogyia) forticornis Strand, 1939, syn. nov.; Thecturota tenuissima Casey, 1893 = Atheta marchii Dodero, 1922, syn. nov.; and Trichiusa robustula Casey, 1893 = T. immigrata Lohse, 1984, syn. nov. The Palaearctic species Amarochara forticornis, Anomognathus cuspidatus, Oligota pumilio, and Parocyusa rubicunda are newly confirmed from the Nearctic region as adventive, resulting in the following synonymies: Parocyusa rubicunda (Erichson, 1837) = Chilopora americana Casey, 1906, syn. nov. and Anomognathus cuspidatus (Erichson, 1839) = Thectura americana Casey, 1893, syn. nov. The genus Dasygnypeta, sensu nov. is newly reported from North America, Paradilacra is newly reported from eastern North America, and Haploglossa is newly reported from Canada, resulting in the following synonymy: Paradilacra densissima (Bernhauer, 1909) = Gnypeta saccharina Klimaszewski & Webster, 2008, syn. nov. Native Cyphea wallisi is newly reported from across Canada and C. curtula is removed from the Nearctic fauna. The status of both Gyrophaena affinis and Homalota plana is uncertain but these species are no longer considered to be adventive in North America. Three new combinations are proposed: Dasygnypeta baranowskii (Klimaszewski, 2020) and D. nigrella (LeConte, 1863) (both from Gnypeta) and Mocyta scopula (Casey, 1893) (from Acrotona). Dolosota Casey, 1910, syn. nov. (type species Eurypronota scopula Casey), currently a subgenus of Acrotona, is therefore synonymized with Mocyta Mulsant & Rey, 1874. Additionally, four new Canadian records and 18 new provincial and state records are reported.

Review of the genus Cobitis in the Middle East, with the description of eight new species (Teleostei: Cobitidae)

Zootaxa ◽  
2018 ◽  
Vol 4535 (1) ◽  
pp. 1 ◽  
Author(s):  
JÖRG FREYHOF ◽  
ESRA BAYÇELEBİ ◽  
MATTHIAS GEIGER
Keyword(s):  
New Species ◽  
Middle East ◽  
Process Analysis ◽  
Dna Barcode ◽  
Black Spot ◽  
Morphological Characters ◽  
Morphological Data ◽  
Colour Pattern ◽  
Caudal Fin ◽  
River Drainage

The diversity of Cobitis in the Middle East is reviewed, resulting in the recognition of 30 species, of which eight are described herein as new. Two species, C. amphilekta and C. kellei, seem to be extinct. Hypotheses on species-level diversity derived from distance and Poisson tree process analyses of DNA barcode data are tested against morphometric and morphological characters including colour patterns. For species pairs separated by small K2P distances in COI sequence data we follow a practitioner-oriented diagnostic species concept, in which we recognise species only if differentiated morphologically (including by colour pattern). For all 30 species we provide diagnoses and identification keys. Cobitis afifeae, new species, from the Büyük Menderes River drainage in the Aegean Sea basin, is distinguished by having two laminae circularis in the male, a row of blotches below Z4, a small, roundish or comma-shaped black spot at the upper caudal-fin base, and elevated mental lobes. Cobitis aliyeae, new species, from the lower Seyhan and Ceyhan River drainages, is distinguished by having two laminae circularis in the male, the blotches in Z2 and Z4 anterior to the dorsal-fin origin usually well separated from each other, and the pigmentation in Z1 well distinguished from the pigmentation in Z2. Cobitis anabelae, new species, from the lower Orontes River drainage, is distinguished by having two laminae circularis in the male, the pigmentation in Z2 formed by small, brown spots, always much smaller than blotches in Z3, much smaller than the pupil diameter, Z2 and Z3 well separated, and no pigmentation below Z4. Cobitis erkakanae, new species, from the Gölbasi Lakes, adjacent to the Ceyhan River drainage, is distinguished by having two laminae circularis in the male, no blotches below Z4, the blotches in Z2 and Z4 being horizontally elongated and often fused with adjacent ones, and the caudal fin with 4–6 wide, regularly-shaped, brown bands. Cobitis emrei, new species, from the Lake Sapanca basin is distinguished by having one lamina circularis in the male, a large black spot at the upper caudal-fin base, and Z3 fully covered by very small spots forming a sand-like pattern. Cobitis joergbohleni, new species, from the Sultan marshes in Central Anatolia is distinguished by having two laminae circularis in the male, and the flank colour pattern being completely disorganised, not following the Gambetta zones. Cobitis pirii, new species, from the endorheic Lake Eğirdir basin and the Mediterranean Aksu and Köprü Rivers, is distinguished by having two laminae circularis in the male, a simple external part of the suborbital spine and two distinct rows of small blotches in Z4, one along the lateral midline and one distinctly below. Cobitis troasensis, new species, from the Tuzla River drainage, is distinguished by having one lamina circularis in the male and 25–36 small, comma-shaped brown blotches in Z4. A lectotype is designated for Cobitis battalgilae. As First Revisers, priority is given to Cobitis fahireae over C. kurui. Cobitis damlae and C. kurui are treated as synonyms of C. fahireae. Cobitis strumicae and C. taenia are recorded for the first time from Anatolia and C. saniae is newly documented from the Black Sea basin in Georgia. The Poisson tree process analysis of COI data proposed 31 groups, most of which could be distinguished by morphological characters. Cobitis troasensis is described based on morphological data alone.  

scholarly journals Establishment of six new Rhabdoblatta species (Blattodea, Blaberidae, Epilamprinae) from China

ZooKeys ◽  
2019 ◽  
Vol 851 ◽  
pp. 27-69
Author(s):  
Rong Yang ◽  
Zhenzhen Wang ◽  
Yanshuang Zhou ◽  
Zongqing Wang ◽  
Yanli Che
Keyword(s):  
Phylogenetic Analysis ◽  
New Species ◽  
Molecular Species ◽  
Species Delimitation ◽  
Female Genitalia ◽  
Morphological Data ◽  
Coi Sequences ◽  
First Time

This study examined 504 Rhabdoblatta specimens sampled from China, of which, 86 Rhabdoblatta specimens were used for COI sequencing. A phylogenetic analysis using the ML method and MOTUs estimations by ABGD and GMYC based on COI sequences was performed. Eighteen Rhabdoblatta species were identified when these data were combined with morphological data. Six new species were established among these samples, i.e., Rh.similsinuatasp. n., Rh.densimaculatasp. n., Rh.gyroflexasp. n., Rh.chaulformissp. n., Rh.maculatasp. n., and Rh.ecarinatasp. n. For the first time, females including female genitalia of 14 known Rhabdoblatta species are described worldwide. Our study shows that combining molecular species delimitation methods with morphological data helps to delimit species and understand cockroach biodiversity.

Integrative description of five Pseudechiniscus species (Heterotardigrada: Echiniscidae: the suillus-facettalis complex)

Zootaxa ◽  
2020 ◽  
Vol 4763 (4) ◽  
pp. 451-484 ◽  
Author(s):  
MILENA ROSZKOWSKA ◽  
DARIA GROBYS ◽  
TOMASZ BARTYLAK ◽  
MAGDALENA GAWLAK ◽  
HANNA KMITA ◽  
...  
Keyword(s):  
New Species ◽  
Phylogenetic Tree ◽  
Genetic Data ◽  
Integrative Taxonomy ◽  
Genetic Distances ◽  
Morphological Features ◽  
Species Discrimination ◽  
Coi Sequences ◽  
Morphology And Morphometry ◽  
Species Specific

Pseudechiniscus is a morphologically homogeneous heterotardigrade genus with a relatively low number of morphological features useful for the species discrimination. The species of the Pseudechiniscus suillus-facettalis complex are some of the most challenging tardigrades to identify. Here, we examine several populations from Antarctica, Italy, Madagascar and Norway that would have most likely been attributed to Pse. suillus prior to the recent redescription of the species. Populations were analysed using integrative taxonomy—a combination of classical morphology and morphometry, as well as genetic data. Besides minute differences in dorsal sculpture and morphometry, we found clear, species-specific differences in ventral sculpture which are very useful in discrimination of Pseudechiniscus species. Based on morphology (mainly ventral sculpture) and significant genetic distances in COI and ITS-2 sequences, we describe five new species: Pse. angelusalas sp. nov. from Madagascar, Pse. dastychi sp. nov. from Antarctica, Pse. ehrenbergi sp. nov. from Italy as well as Pse. indistinctus sp. nov. and Pse. lacyformis sp. nov. from Norway. Finally, we provide an updated phylogenetic tree of the genus Pseudechiniscus based on COI sequences. 

Morphological and molecular evidence of some Minuartia species (Caryophyllaceae) from Turkey, with a desciption of a new species

Phytotaxa ◽  
2019 ◽  
Vol 391 (2) ◽  
pp. 122 ◽  
Author(s):  
MURAT KOÇ ◽  
ERGIN HAMZAOĞLU ◽  
AHMET AKSOY
Keyword(s):  
New Species ◽  
Dna Sequences ◽  
Related Species ◽  
Phylogenetic Relationships ◽  
Morphological Data ◽  
Molecular Evidence ◽  
A New Species

The genus Minuartia is represented in Turkey by 34 taxa. Some interesting specimens were collected from Antalya province, and examined. These specimens resemble Minuartia meyeri, and M. multinervis from which differ by characters (macro-, and micromorphological) of inflorescence, alar pedicels, petals, sepals, capsules and seeds. Moreover, by using the DNA sequences of the ITS genes, phylogenetic relationships between this collected species, and the related species were investigated. As a result of the evaluation of molecular, and morphological data, we proposed to described the population from Antalya as a new species for the science. A description, pictures, distribution, habitat, and IUCN category are given.

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