Wood density variation in naturally regenerated stands of Pinus ponderosa in northern Arizona, USA

In the southwestern United States, land managers are implementing large-scale forest restoration projects involving treatments designed to improve forest health, protect ecosystem services, and reduce the risk of catastrophic wildfire in overstocked ponderosa pine (Pinus ponderosa Dougl. ex P. & C. Laws.) forests. A better understanding of wood properties is necessary to improve the currently limited markets for the woody byproducts generated by these treatments. Therefore, our objective was to investigate variations in ponderosa pine wood density across the northern Arizona landscape. We sampled trees from 18 naturally regenerated stands and used X-ray densitometry to quantify the radial and axial variation within and among trees. We modeled within-stem wood density patterns using generalized additive models, and investigated the effects of climatic variation using response function analyses. Additionally, we tested the effects of site-level predictors on whole-tree wood density. We found high variability in radial wood density profiles among trees in our stands compared with that observed in studies of other species grown in plantations — perhaps due to high genetic variation within naturally regenerated stands. Wood density was negatively correlated with precipitation at annual and most quarterly intervals, except for a positive correlation with late-summer monsoon precipitation. The high wood density variation we found among trees highlights the need for further investigation of controls over wood properties in natural nonplantation forests.

Preliminary Results from a Structural Conversion Thinning Trial in Eastern Canada

In 2008, a thinning trial consisting in the removal of competitors around high growth potential stems (crop trees, CTs) was initiated as the first step of a structural conversion to transform even-aged stands into uneven-aged stands. Two intensities of thinning by CT release and thinning from below were tested in white spruce (Picea glauca [Moench] Voss) plantations and in balsam fir (Abies balsamea [L.] Mill)—dominated naturally regenerated stands. Unlike thinning from below, which aims to remove poor quality stems to improve the growth of residual trees, thinning by CT release aims to reduce competition between dominant trees. Trees thinned by CT release showed a significantly higher diameter at breast height increment than those in the control plots. CT release did not significantly increase stand structure, nor did it reduce stand growth, which helps to address the concerns of regional forest stakeholders. CT release may offer other benefits in the long term, but subsequent interventions are necessary to evaluate if this treatment meets the desired objectives.

The growth of the beech (Fagus sylvatica L.) stand on former agricultural land and its comparison with the naturally regenerated beech stand under comparable conditions

The issue of afforestation of non-forest land (e.g. agricultural land use) is currently a significant trend in land management. Sustainable development and maintenance of its ecological stability are among the reasons. The paper compares the 30-year development of the beech stand originated from artificial regeneration on former agricultural land in the area where the Ipeľ River rises in central Slovakia. Two methods of tending were used on three long-term research plots (LTPs) established at the age of 20 years: negative crown thinning, free crown thinning and the third LTP was left to its spontaneous development. The results confirmed the positive impact of tending on the development of these stands. Compared to naturally regenerated stands under comparable site conditions, their higher quantitative production was found. Regarding the qualitative production expressed by the number of trees of selective quality, the worst results were obtained on the control plot. Based on these results, it is recommended to pay more attention to the planting stock quality used for afforestation.

Comparison of Genetic Diversity in Naturally Regenerated Norway Spruce Stands and Seed Orchard Progeny Trials

Forest ecosystems in Europe are expected to experience changes in temperature and water regimes associated with increased risks of extreme environmental events and disasters. Genetic diversity and relatedness has been linked to resilience of forest stands and landscapes. Genetic diversity indicators were compared between a Norway spruce population naturally regenerated after extensive windthrow and Norway spruce progeny populations derived from two seed orchards. In addition, genetic diversity in an undisturbed stand in a long established national park and a spruce genetic resource stand were analyzed. Populations were genotyped at 11 simple sequence repeat (SSR) loci. Average genetic diversity indicators were similar across populations. However, the total number of alleles, average number of alleles over all loci, effective number of alleles, average gene diversity, and average allelic richness were highest in the naturally regenerated population and lowest in one of the seed orchard progeny populations. The genetic diversity in progeny from seed orchards used for stand renewal is comparable to the genetic diversity in naturally regenerated stands. However, fluctuations in seed production between years can have a large impact on genetic diversity in seed orchard progeny. The use of improved Norway spruce germplasm deployed via clonal seed orchards for forest renewal can maintain similar levels of genetic diversity compared to naturally regenerated stands, while also increasing production and timber quality.

Contrasting Development of Canopy Structure and Primary Production in Planted and Naturally Regenerated Red Pine Forests

Globally, planted forests are rapidly replacing naturally regenerated stands but the implications for canopy structure, carbon (C) storage, and the linkages between the two are unclear. We investigated the successional dynamics, interlinkages and mechanistic relationships between wood net primary production (NPPw) and canopy structure in planted and naturally regenerated red pine (Pinus resinosa Sol. ex Aiton) stands spanning ≥ 45 years of development. We focused our canopy structural analysis on leaf area index (LAI) and a spatially integrative, terrestrial LiDAR-based complexity measure, canopy rugosity, which is positively correlated with NPPw in several naturally regenerated forests, but which has not been investigated in planted stands. We estimated stand NPPw using a dendrochronological approach and examined whether canopy rugosity relates to light absorption and light–use efficiency. We found that canopy rugosity increased similarly with age in planted and naturally regenerated stands, despite differences in other structural features including LAI and stem density. However, the relationship between canopy rugosity and NPPw was negative in planted and not significant in naturally regenerated stands, indicating structural complexity is not a globally positive driver of NPPw. Underlying the negative NPPw-canopy rugosity relationship in planted stands was a corresponding decline in light-use efficiency, which peaked in the youngest, densely stocked stand with high LAI and low structural complexity. Even with significant differences in the developmental trajectories of canopy structure, NPPw, and light use, planted and naturally regenerated stands stored similar amounts of C in wood over a 45-year period. We conclude that widespread increases in planted forests are likely to affect age-related patterns in canopy structure and NPPw, but planted and naturally regenerated forests may function as comparable long-term C sinks via different structural and mechanistic pathways.

Soil enzymatic activity in artificially and naturally regenerated forests after wind damage in north-eastern Poland

The aim of the study was to describe biochemical reactions in the soil based on the activity of urease and dehydrogenases in Scots pine stands damaged during and differentially managed after hurricanes. Soil enzymatic activity was investigated in 2005 and again in 2013 for selected stands in the Pisz Forest District where wind damage occurred in 2002. Most of the damaged stands were cut down and replanted, but 445 ha were left untouched for study purposes. The investigated areas differed with respect to stand damage and management. The enzymatic activity correlated well with the content of organic matter which was higher in organic than in mineral soils. In the area left to regenerate naturally, dehydrogenase and urease activity was higher in 2013 compared to 2005, which suggests the improvement of site conditions. The values of the examined biochemical parameters were correlated with the type of forest regeneration, with the forest soil regeneration being higher in artificially than in naturally regenerated stands. Changes of soil enzymatic activity were correlated with the level of stand damage and the type of management after wind damage.

Sitka Spruce and Western Hemlock Stand and Tree Growth 10 Years after Precommercial Thinning in Southeast Alaska

Ten years after precommercially thinning 16–18-year-old naturally regenerated stands of Sitka spruce (Picea sitchensis [Bong.] Carr.) and western hemlock (Tsuga heterophylla [Raf.] Sarg.) in southeast Alaska, neither species responded in height growth to spacing. Spruce diameter growth increased significantly following all thinning treatments, and among thinning treatments, there was a weak but increasing trend toward more rapid diameter growth at the wider spacings. Spruce basal areas increased almost twice as fast after thinning as without thinning, and radial growth continued to increase, whereas unthinned stands grew at a slower but steady rate. Western hemlock also showed a trend for increased diameter, but growth response was less than for spruce. Shrub control to enhance understory plant development for deer forage did not improve growth of either tree species. Branch diameter of Sitka spruce increased with spacing. Pruning led to epicormic sprouting in the 3 years following pruning with numbers inversely related to spacing. Sprouts developed and persisted in stands thinned to 200 or fewer stems per acre. Overall, pruning led to a small reduction in diameter growth at breast height.