On limits of the distribution range of the Crimean rock lizard Darevskia lindholmi (Sauria: Lacertidae)

New data on the boundaries of the distribution range of the Lindholm rock lizard (Darevskia lindholmi), an endemic of the Crimean Peninsula, are presented. This petrophilous lizard inhabit a wide range of biotopes in various landscape levels of the Mountainous Crimea. The upper boundary of D. lindholmi distribution in the southwest of the Main Range of the Crimean Mountains reaches an elevation of 1,520 m a.s.l. (Ai-Petrinskaya Yayla, KemalEgerek Mountain), while on the other high uplands with altitudes above 1.5 km and colder climate (Babugan and Chatyrdag), the species was traced only up to 1,250–1,320 m a.s.l. The northern border of D. lindholmi range in the western part of the Crimean Mountains runs along the Outer Foothill Range (the right bank of the Alma River), while in the eastern part it corresponds the northernmost rocky massifs of the Inner Foothill Range to the north of 45º N latitude. Isolated marginal populations found in the forest-steppe or phrygana-steppe landscapes of the Foothills and arid Southeastern Coast differs significantly in their distance from the main habitat of the species, lizards’ abundance and density. A hypothetical history of the formation of the current range of the Lindholm lizard is discussed.

The mitochondrial phylogeography of the Crimean endemic lizard Darevskia lindholmi (Sauria, Lacertidae): Hidden diversity in an isolated mountain system

The Lindholm rock lizard, Darevskia lindholmi, is the only member of the genus Darevskia whose range is restricted solely to Europe, representing a local endemism found only in the Crimean Mountains. In our study, we investigated the cytochrome b gene (mtDNA) of 101 D. lindholmi sequences from 65 Crimean localities, representing its entire range. We found that D. lindholmi is highly genetically structured, and its range is divided into populations belonging to three mitochondrial lineages. The Lindholm rock lizard populations inhabiting the middle part of the Crimean Mountains (further referred to as the Central lineage) are sharply differentiated from the other two lineages (the Common and the Southwestern lineages), which are present in most of the species range. The genetic distance between the Central lineage and the other two taken together is 4.6%, according to our results, suggesting that the divergence occurred during the Early Pleistocene. The narrowly distributed Southwestern lineage and the widespread Common lineage, on the other hand, are differentiated by 1%. Field observations on the representatives of the main evolutionary groups show that their ecology is also different: the Central lineage is a mesophilic and cold-resistant form, while the other two closely related lineages are more xerophilic and thermophilic. Results of the potential ranges modeling and ecological niche analysis confirm that the genetic lineages occupy different niches of the Crimea. Furthermore, the area of inhabitation of the Central lineage splits the western and eastern parts of the Common lineage range, while the Southwestern lineage is restricted along the coast of the southwestern coast of the peninsula. The long-term co-existence of deeply divergent sister mitochondrial lineages in a relatively small (circa 7,000 km2) isolated mountain system serves as a mesocosm for understanding the speciation process. Our data suggest that the Central lineage warrants further taxonomic investigation.

Stipo pulcherrimae-Festucetalia pallentis Pop 1968 of calcareous petrophytic steppes in Ukraine

Nine hundred and eight-nine relevés from calcareous petrophytic steppes in Ukraine and its adjacent territories were assessed with the help of expert systems to determine the syntaxonomic affiliation of the plant communities at class and order levels. At least 488 relevés belonging to the class Festuco-Brometea were analyzed using the TWINSPAN algorithm, and 8 distinctive clusters were obtained, recognized as alliances of the order Stipo pulcherrimae-Festucetalia pallentis. A new alliance, Bromopsido cappadocicae-Asphodelinion tauricae, was ascribed to the Crimean Mountains and the presence of two alliances, Diantho lumnitzeri-Seslerion albicantis and Genisto tetragonae-Seselion peucedanifoliae, was confirmed as new for this vegetation in Ukraine. Unlike in the Pannonian Basin, Bromo pannonici-Festucion csikhegyensis alliance communities mentioned in the literature do not occur in Ukraine. Centaureo carbonatae-Koelerion talievii has been provisionally transferred from Festucetalia valesiacae to the order Stipo pulcherrimae-Festucetalia pallentis. Furthermore, we distinguished alliances by their geographic locations and their climatic (thermoregime, cryoregime, light in communities) and edaphic (carbonate content, salinity, and acidity) features.

TECTONIC STRUCTURE OF KERCH-TAMAN FOLD ZONE OF AZOV-KUBAN TROUGH

The nrotectonic structure of the Kerch-Taman folded zone, the southern deformed part of the Azov-Kuban piedmont trough, has a number of structural features and a history of deformation development. The authors divide the zone into five transverse segments, differing in the age of the main phase of folding. In the central Taman segment, folding is still going on, while in the framing segments, the main phases of folding took place in the Pliocene. In the peripheral segments, the main phase of folding is older - Early-Middle Miocene. The boundaries of the Kerch-Taman zone, as well as its youngest, the Taman segment, coincide with the transverse fault (fault-flexure) zones. It is shown that the southern frame of the Kerch-Taman folded zone is the continuation of the structures of the Crimean Mountains and the Northwestern Caucasus. The correlation of mud volcanism manifestations with tectonic elements of the Taman segment is noted to be ambiguous.

Geomorphology and Neotectonics of Southwestern Crimea

—The area of southwestern Crimea includes the ending of the Crimean Mountains that arose during the neotectonic activation at the place of the Cretaceous–Paleogene denudation plain and the adjacent shallow-water carbonate sedimentation basin. The Crimean Mountains are one of the links of the Alpine–Himalayan orogenic belt formed during the collision of the Eurasian, African, and Indo–Australian plates. Their area includes late Cenozoic marine terraces of the complete Mediterranean series and a staircase of Neogene, Paleogene and Cretaceous planation surfaces over them. The planation surfaces of different ages resulted from the successive lowering of the World Ocean level. Their subsequent deformations make it possible to outline the area of the neotectonic uplifting and determine its parameters. The main mechanism of the neotectonic activation was the thrust of the East Black Sea microplate under the Scythian one and the formation of a ramp fold structure. The amplitude of the neotectonic uplifting of southwestern Crimea for the past 2 Myr varies from 0 to 800 m, i.e., is up to 0.04 mm/year. The recent neotectonic structure of the area is formed by the northern flank of the ramp fold; it is a monocline of NW dip consisting of “keys” of NW strike separated by the latest faults with vertical displacements of 10 to 120 m. The uplifting of the area and the lowering of the World Ocean level led to a widespread of denudation surfaces. Their good preservation makes it possible to refine the sequence of neotectonic events, whose first pulses reached the study area in the Oligocene, and the main activation phase began in the Pliocene.

Early Mesolithic habitation in the Shpan-Koba grotto (Crimea, Ukraine)

Shpan-Koba Grotto is currently the only stratified Mesolithic and Neolithic site on the plateau of the First Range of the Crimean Mountains (Yayla). Lower Early Mesolithic cultural layers of the site (archaeological unit 3) have exceptional preservation due to the rapid accumulation of sediment and infrequent settlement of the grotto. Their planigraphy, number and composition of the artifacts can tell about the peculiarities of the economical use of the landscapes and rock shelter of the Crimean highlands at that time. Cultural layers of archaeological unit 3 date back to time from 11500 to 7600 years cal BP, which belongs to Early Preborial. At this time, the climate was colder and wetter than today, the plateau of the Yayla around Shpan-Koba was covered by mesofit steppes, pine, birch and juniper grew on the slopes of the mountains. According to archaeozoological data, the fauna of the Yayla included such representatives of steppe landscapes as saiga and horse, and simultaneously typical forest animals — red deer, brown bear and lynx. All cultural layers of the unit 3 are very similar. They belong to the type of "ephemeral": are represented by small fires, few bones of hunted animals and single flint artifacts. The layers were left by the bearers of the Swiderian culture, due to very few flint artifacts, among them: swidrian points, segment, backed blades, straight dihedral burin, end-scrapers. More than half of the found flint artifacts are retouched tools, the rest — blades and flakes, have the traces of use in the form of macro retouch. Such composition of the flint inventory indicates on the hunters (“expeditional”) character of the habitations in the Shpan-Koba grotto. The planigraphy of all layers of the unit 3 was similar too. It corresponds to classic ethnographical «Drop-Toss model» be L. Binford, which describe the organization of the living space around the hearth by a group of people from one to five people (Binford 1978; 1983). The central object in each of the cultural layers was one hearth about one meter in diameter with the thin charcoal lens and little piece of burned clay under it. Three concentric zones were traced around the hearths on the western, southern, and southeastern sides: 1) without artifacts; 2) with little bones and flint artifacts (Drop zone); 3) with bigger bones (Toss zone). The "asymmetrical" location of the finds in relation to the hearths indicates the absence of artificial housing in the grotto. The only exception is the habitation of the 3-5/6 cultural layer, in which a small wall of stones was excavated. For it, the "symmetrical" location of the finds around the hearths can be assumed to be an artificial structure made of plant materials, such as a brush windbreak or a hut. The windbreak could also exist in the habitation of layer 3-2, judging by the lack of a Toss zone in the south-western part. The presence of only one hearth in each of the cultural layers, the location of the artifacts relative to the hearth and their number, the composition of hunting prey, etc., evidence, that Shpan-Koba grotto in the Early Mesolithic was used as dwelling of little group (4—5 people). The occupation, probably, was very short terming, due to small and low-power fires, very few bones of animals and single flint artifacts, which were found in сertain layers. Their purpose was recreation, skinning and butchering of hunting prey, repair of hunting equipment and more. Seasonality of the Early Mesolithic dwellings in the grotto, due to archaeozoological data, fall on warm time — spring and summer (Benecke 1999, s 83, abb. 10). The aim of the swidrian people’s hunting expeditions to Yaila was hunting, first, on the saiga and red deer, which migrated from from the steppes of northern Crimea. The number of the red deer also increased in the First Range of the Crimean Mountains during warm seasons. The bones of the brown bear in many layers suggest that it was also an attractive prey. Key words: Crimea, Ukraine, Early Mesolithic, habitation, seasonal migrations, Swiderian.