Sandstone body character, river styles, and geomorphology of the lower Eocene Willwood Formation, Bighorn Basin, Wyoming, USA

The lower Eocene Willwood Formation of the intermontane Bighorn Basin, Wyoming, USA, is an alluvial red bed succession with a sand content of ca. 20%-25%. The formation has been studied intensively for paleontology, paleoclimate, and sedimentary reconstruction. However, alluvial sandstone bodies and their corresponding river styles remain little characterized and documented. Here, efforts are made to study the characteristics and river styles of sandstone bodies through ca. 300 m of alluvial stratigraphy in the McCullough Peaks outcrop area based on the field data and a georeferenced 3-D photogrammetric model. Four channel facies associations are recognized, and they are ascribed to four river planform styles: distributary channel, massive trunk-shaped channel, braided channel, and sinuous channel, with the latter two styles being the more abundant. The channel sandstone bodies that show the character of sinuous rivers and those of braided rivers differ significantly in average thickness (6.1 m versus 9.0 m) and insignificantly in average width (on average 231 m) and paleoflow directions (on average N003). Braided-character dominated and sinuous-character dominated river styles are seen to alternate in the outcrop, while they show no spatial dependency in the 10 km2 study area. Bighorn Basin margins varied in the early Eocene, with differing tectonic, geological, and topographic characteristics. The observed mixture of river styles may be attributed to differential influences of axial and transverse river systems and/or climate change that controls water discharge and sediment load. An early Eocene geomorphologic reconstruction is constructed summarizing these new and earlier results.

Effects of phylogenetic uncertainty on fossil identification illustrated by a new and enigmatic Eocene iguanian

Fossil identifications made in a phylogenetic framework are beholden to specific tree hypotheses. Without phylogenetic consensus, the systematic provenance of any given fossil can be volatile. Paleobiogeographic and divergence time hypotheses are contingent on the accurate systematic placement of fossils. Thus, fossil diagnoses should consider multiple topologies when phylogenetic resolution or clear apomorphies are lacking. However, such analyses are infrequently performed. Pleurodonta (Squamata: Iguania) is an ancient and frequently-studied lizard clade for which phylogenetic resolution is notoriously elusive. I describe a skull fossil of a new pleurodontan lizard taxon from the Eocene deposits of the Willwood Formation, Wyoming, and use the new taxon as a case-study to explore the effects of phylogenetic uncertainty on fossil identification. The relationships of the new taxon differ considerably among analyses, and resulting interpretations are correspondingly disparate. These results illustrate generalizable and severe issues with fossil interpretations made without consideration of alternative phylogenetic hypotheses.

Stratigraphic relationships along the monoclinal eastern base of Bald Ridge and northwestern edge of Wyoming’s Bighorn Basin, U.S.A.

ABSTRACT This geologic study is focused on a less than 5 square-mile (ca. 13 km2) tract of public land in northwestern Wyoming, 8 miles (12.9 km) south-southwest of the small town of Clark in Park County. The study area is south of Clarks Fork of Yellowstone River along the eastern base of the topographic feature called Bald Ridge, also known structurally as Dead Indian monocline. Since the Middle Eocene, the study area has been along the northwestern margin of the Bighorn Basin. Prior to that time, the study area existed near the west–east center of the basin. Bald Ridge became elevated late in the Laramide orogeny (no older than the Early Eocene) through east-directed faulting of basement rocks via the extensive Line Creek–Oregon Basin thrust system. As that active faulting occurred, the overlying Phanerozoic strata (Lower Cambrian through Lower Eocene) responded with numerous west-directed, out-of-the-basin thrusts as a new western-basin margin developed along the eastern realm of the newly born Absaroka volcanic field. Most of that deformation occurred after deposition of uppermost levels of the Lower Eocene Willwood Formation. The key purpose of the present paper was to improve the accuracy of mapping of the Jurassic into Eocene stratigraphy along the newly restricted, northwestern edge of Wyoming’s Bighorn Basin. The stratigraphic column in a north–south band along the eastern flank of the Beartooth Mountains and continuing southward into the present study area was markedly deformed and deeply eroded late during the Laramide orogeny. The present small, more southerly study area is structurally and erosionally simpler than its more northerly equivalent. Thus, its study adds important geological information to the history of the northern Cody Arch, a convex-westward string of related basement-involved uplifts extending southward to southwest of the city of Cody. Progressively steepening eastward dips of strata characterize a west-to-east transect from the summit of Bald Ridge (capped by the shallowly dipping, Mississippian Madison Limestone) to the western edge of strongly overturned outcrops of the Eocene Willwood Formation. The Upper Cretaceous Meeteetse Formation is the stratigraphic horizon at which the dips attain vertical or slightly overturned orientations. All consequential faults within the newly mapped area are thrusts, and they show generally westward (out-of-the-basin) displacements. Despite those west-directed displacements, their primary cause was tectonic shortening at depth below Bald Ridge that was directed to the northeast or east-northeast. During the Laramide orogeny, certain thrust planes within the east-dipping Phanerozoic rock column cut down-section stratigraphically (but uphill relative to Earth’s surface) and thereby placed younger strata upon older. The cumulative result, as recognized at several levels within the present area of study, was marked thinning of the total section. For example, surface exposures of the mostly Paleocene Fort Union Formation, 4,000 feet (1,219 m) thick only 7 miles (11.3 km) to the east, was completely eliminated from the local surface stratigraphy by that means. The northern end of Bald Ridge is formed by the highly asymmetric Canyon Mouth anticline. That structure differs strongly in the attitude of its hinge line from the general east-northeast dip of strata cloaking Bald Ridge. The Canyon Mouth anticline’s hinge line plunges steeply to the southeast, and dips on its northeastern flanks are vertical to partly overturned. Surprisingly, hinge lines and flanks of all other anticlinal/synclinal structures recognized within the present map area share those same orientations with Canyon Mouth anticline. These consistent but unexpected differences in orientation from unfolded strata may represent very late events in the history of Laramide strain vectors across the study area. Working in northern parts of this study area, an independent group determining radiometric ages of detrital-zircon grains reported close agreements in age with their host localities in the Early Cretaceous Mowry Shale and Frontier Formation. However, under the present paper’s interpretation of the local stratigraphy, the other workers misidentified formational hosts for all three samplings. That resulted in age-determination errors of depositional history within the Upper Cretaceous section of as much as 28.8 million years.

Calcardea junnei Gingerich, 1987 from the late Paleocene of North America is not a heron, but resembles the early Eocene Indian taxon Vastanavis Mayr et al., 2007

We revisit the holotype of Calcardea junnei Gingerich, 1987 from the latest Paleocene (Clarkforkian) of the Willwood Formation (Wyoming, USA). The species is based on a partial skeleton and was originally assigned to the Ardeidae (herons). As we show, this classification cannot be upheld and Calcardea Gingerich, 1987 more closely resembles the taxon Vastanavis Mayr et al., 2007 (Vastanavidae), a parrot-like bird from the early Eocene of India. Even though C. junnei is a large bird, its long wings and short tarsometatarsus argue against a predominantly terrestrial way of living, and the morphology of the tarsometatarsus and pedal phalanges instead suggest strong grasping feet. We conclude that an assignment of Calcardea to the landbird clade (Telluraves) is better supported than its classification into the waterbird clade (Aequornithes), which includes Ardeidae and other ‘ciconiiform’ and ‘pelecaniform’ taxa. Calcardea junnei is one of the oldest known representatives of Telluraves and its morphology shows plesiomorphic features, which contributed to its previous misidentification as a heron. Calcardea exhibits a distinctive osteology and affords a glimpse of a previously unknown late Paleocene avian morphotype.

Fingerprinting snakes: paleontological and paleoecological implications of zygantral growth rings in Serpentes

We introduce a new non-destructive source of skeletochronological data with applications to species identification, associating disarticulated remains, assessing minimum number of individuals (MNI), and collection management of fossil snakes, but with potential implications for all bony vertebrates, extinct or extant. Study of a diverse sample of Recent henophidian snakes confirms that annual growth cycles (AGCs) visible on the surface of the vertebral zygantrum correspond to lines of arrested growth in osteohistological thin sections and accordingly reflect chronological age. None of the specimens considered here showed signs of remodelling of the zygantrum, suggesting that a complete, unaltered age record is preserved. We tested potential influences on AGCs with a single experimental organism, a maleBogertophis subocularis, that was raised at a controlled temperature and with constant access to mice and water. The conditions in which this individual was maintained, including that it had yet to live through a full reproductive cycle, enabled us to determine that its AGCs reflect only the annual solar cycle, and neither temperature, nor resource availability, nor energy diversion to gametogenesis could explain that it still exhibited lines of arrested growth. Moreover, growth lines in this specimen are deposited toward the end of the growth season in the fall, and not in the winter, during which this individual continued to feed and grow, even though this mid-latitude species would normally be hibernating and not growing. This suggests that growth lines are not caused by hibernation, but reflect the onset of a physiological cycle preparingBogertophis subocularisfor winter rest. That being said, hibernation and reproductive cycle could still influence the amount of time represented by an individual growth line. Growth-line number and AGC spacing-pattern, plus centrum length, are used to estimate MNI of the Early Eocene fossil snakeBoavus occidentaliscollected from the Willwood Formation over two field seasons during the late 19th century. We identified eight or nine individuals among specimens previously parcelled among two specimen lots collected during those expeditions.

Detrital Zircon U-Pb geochronology and provenance of the Eocene Willwood Formation, Northern Absaroka Basin, Wyoming

We report the results of U-Pb ages from detrital zircon populations in the lower Eocene synorogenic Willwood Formation in the northern Absaroka Basin, Wyoming. Zircons (n=229) were extracted from three sandstone beds and one ash layer in the Willwood Formation at the base of Jim Mountain in the North Fork Shoshone River Valley. K-S statistical analysis indicates that the three sandstones, which were sampled from the base, middle, and top of the formation, have identical age spectra, indicating that the sandstone provenance remained the same during the duration of Willwood deposition. The zircon age spectra are dominated by Archean zircons (61%), with peak ages at 3270 and 2770 Ma. These sandstones also have very early Paleoproterozoic zircons (∼2450 Ma), which likely were derived from the Tobacco Root Mountains. The final significant age peak is ∼70 Ma, which is likely associated with the Cretaceous Tobacco Root batholith. The Jim Mountain ash, which occurs at the top of the succession, just beneath the allocthonous volcanic rocks of the Heart Mountain slide, has a maximum depositional age of ∼50 Ma. Between 49–50 Ma, as Eocene volcanism in the northern Absaroka Range became more prominent, stratovolcanoes grew and disrupted sediment transport into the Absaroka basin. Lower Wapiti sandstones to the southwest show a mix of Eocene, recycled Proterozoic and Archean grains. The coeval Crandall Conglomerate, which was dismembered by the emplacement of the Heart Mountain slide in the northern Absaroka Range, has a distinct detrital zircon age spectrum. Thus these stream systems that deposited the Crandall did not share the headwaters with the streams that supplied sediment to the Absaroka basin.