The essays in this volume are analyzed to assess the degree to which they portray scientific and beyond-science interactions. The Long-Term Ecological Research (LTER) program represents a scientific or intellectual movement based on articulation of the program’s highly respected founders, resource allocation for individual and collective pursuits, use of LTER sites for recruitment, and commonly held themes or foci for research. Interdisciplinary scientific interactions within the LTER program have influenced researchers’ ideas, networks, and productivity but have also presented challenges, particularly for junior participants. Interactions beyond the scientific community focus on one-dimensional flows of information as well as on collaborative, multidirectional partnerships with a variety of stakeholders. This analytical chapter explores social interactions catalyzed by experiences of scientists associated with the LTER program. I analyze the essays by LTER scientists in this volume using a broad, three- tiered structure: (1) the degree to which insights from the essays suggest that the LTER program represents a scientific or intellectual movement within environmental sciences examining ecological dynamics; (2) the extent of interdisciplinary interactions with scientists across broader fields of study, including associated reactions and challenges; and (3) interactions with others beyond science. Findings are examined across different career stages of respondents. Direct quotations are used to illustrate findings and to provide evidence for conclusions based on the LTER scientists’ own words. The LTER program was initiated 34 years ago (Waide [Chapter 2]; Gholz, Marinelli, and Taylor [Chapter 3]). Given the growth of the LTER program, in terms of the number and geographic distribution of sites, as well as the contributions of engaged scientists and students, there is no doubt of the influence of the LTER program on the science of ecology and general understanding of ecosystems around the world (Robertson et al. 2012). In this chapter, I examine the social interactions of scientists in the LTER program through the lenses provided by their essays in this volume to explore three dimensions—interactions within the environmental sciences focused on ecological dynamics, broader interdisciplinary interactions, and interactions with stakeholders beyond science.