scholarly journals First record of Mesozoic terrestrial vertebrates from Lithuania: phytosaurs (Diapsida: Archosauriformes) of probable Late Triassic age, with a review of phytosaur biogeography

2012 ◽  
Vol 150 (1) ◽  
pp. 110-122 ◽  
Author(s):  
STEPHEN L. BRUSATTE ◽  
RICHARD J. BUTLER ◽  
GRZEGORZ NIEDŹWIEDZKI ◽  
TOMASZ SULEJ ◽  
ROBERT BRONOWICZ ◽  
...  

AbstractFossils of Mesozoic terrestrial vertebrates from Lithuania and the wider East Baltic region of Europe have previously been unknown. We here report the first Mesozoic terrestrial vertebrate fossils from Lithuania: two premaxillary specimens and three teeth that belong to Phytosauria, a common clade of semiaquatic Triassic archosauriforms. These specimens represent an uncrested phytosaur, similar to several species within the generaPaleorhinus,Parasuchus,RutiodonandNicrosaurus. Because phytosaurs are currently only known from the Upper Triassic, their discovery in northwestern Lithuania (the Šaltiškiai clay-pit) suggests that at least part of the Triassic succession in this region is Late Triassic in age, and is not solely Early Triassic as has been previously considered. The new specimens are among the most northerly occurrences of phytosaurs in the Late Triassic, as Lithuania was approximately 7–10° further north than classic phytosaur-bearing localities in nearby Germany and Poland, and as much as 40° further north than the best-sampled phytosaur localities in North America. The far northerly occurrence of the Lithuanian fossils prompts a review of phytosaur biogeography and distribution, which suggests that these predators were widely distributed in the Triassic monsoonal belt but rarer in more arid regions.

2009 ◽  
Vol 83 (5) ◽  
pp. 783-793 ◽  
Author(s):  
B. Senowbari-Daryan ◽  
G. D. Stanley

Stromatomorpha californica Smith is a massive, calcified, tropical to subtropical organism of the Late Triassic that produced small biostromes and contributed in building some reefs. It comes from the displaced terranes of Cordilleran North America (Eastern Klamath terrane, Alexander terrane, and Wrangellia). This shallow-water organism formed small laminar masses and sometimes patch reefs. It was first referred to the order Spongiomorphidae but was considered to be a coral. Other affinities that have been proposed include hydrozoan, stomatoporoid, sclerosponge, and chambered sponge. Part of the problem was diagenesis that resulted in dissolution of the siliceous spicules and/or replaced them with calcite. Well-preserved dendroclone spicules found during study of newly discovered specimens necessitate an assignment of Stromatomorpha californica to the demosponge order Orchocladina Rauff. Restudy of examples from the Northern Calcareous Alps extends the distribution of this species to the Tethys, where it was an important secondary framework builder in Upper Triassic (Norian-Rhaetian) reef complexes. Revisions of Stromatomorpha californica produce much wider pantropical distribution, mirroring paleogeographic patterns revealed for other tropical Triassic taxa. Review of Liassic material from the Jurassic of Morocco, previously assigned to Stromatomorpha californica Smith var. columnaris Le Maitre, cannot be sustained. Species previously included in Stromatomorpha are: S. stylifera Frech (type species, Rhaetian), S. actinostromoides Boiko (Norian), S. californica Smith (Norian), S. concescui Balters (Ladinian-Carnian), S. pamirica Boiko (Norian), S. rhaetica Kühn (Rhaetian), S. stromatoporoides Frech, and S. tenuiramosa Boiko (Norian). Stromatomorpha rhaetica Kühn described from the Rhaetian of Vorarlberg, Austria shows no major difference from S. californica. An example described as S. oncescui Balters from the Ladinian-Carnian of the Rarau Mountains, Romania, is very similar to S. californica in exhibiting similar spicule types. However, because of the greater distance between individual pillars, horizontal layers, and the older age, S. oncescui is retained as a separate species. The net-like and regular skeleton of Spongiomorpha sanpozanensis Yabe and Sugiyama, from the Upper Triassic of Sambosan (Tosa, Japan), suggests a closer alliance with Stromatomorpha, and this taxon possibly could be the same as S. californica.


2008 ◽  
Vol 276 (1656) ◽  
pp. 507-513 ◽  
Author(s):  
Walter G Joyce ◽  
Spencer G Lucas ◽  
Torsten M Scheyer ◽  
Andrew B Heckert ◽  
Adrian P Hunt

A new, thin-shelled fossil from the Upper Triassic (Revueltian: Norian) Chinle Group of New Mexico, Chinlechelys tenertesta , is one of the most primitive known unambiguous members of the turtle stem lineage. The thin-shelled nature of the new turtle combined with its likely terrestrial habitat preference hint at taphonomic filters that basal turtles had to overcome before entering the fossil record. Chinlechelys tenertesta possesses neck spines formed by multiple osteoderms, indicating that the earliest known turtles were covered with rows of dermal armour. More importantly, the primitive, vertically oriented dorsal ribs of the new turtle are only poorly associated with the overlying costal bones, indicating that these two structures are independent ossifications in basal turtles. These novel observations lend support to the hypothesis that the turtle shell was originally a complex composite in which dermal armour fused with the endoskeletal ribs and vertebrae of an ancestral lineage instead of forming de novo. The critical shell elements (i.e. costals and neurals) are thus not simple outgrowths of the bone of the endoskeletal elements as has been hypothesized from some embryological observations.


2017 ◽  
Vol 7 (1) ◽  
Author(s):  
Michelle R. Stocker ◽  
Li-Jun Zhao ◽  
Sterling J. Nesbitt ◽  
Xiao-Chun Wu ◽  
Chun Li

Abstract Following the end-Permian extinction, terrestrial vertebrate diversity recovered by the Middle Triassic, and that diversity was now dominated by reptiles. However, those reptilian clades, including archosaurs and their closest relatives, are not commonly found until ~30 million years post-extinction in Late Triassic deposits despite time-calibrated phylogenetic analyses predicting an Early Triassic divergence for those clades. One of these groups from the Late Triassic, Phytosauria, is well known from a near-Pangean distribution, and this easily recognized clade bears an elongated rostrum with posteriorly retracted nares and numerous postcranial synapomorphies that are unique compared with all other contemporary reptiles. Here, we recognize the exquisitely preserved, nearly complete skeleton of Diandongosuchus fuyuanensis from the Middle Triassic of China as the oldest and basalmost phytosaur. The Middle Triassic age and lack of the characteristically-elongated rostrum fill a critical morphological and temporal gap in phytosaur evolution, indicating that the characteristic elongated rostrum of phytosaurs appeared subsequent to cranial and postcranial modifications associated with enhanced prey capture, predating that general trend of morphological evolution observed within Crocodyliformes. Additionally, Diandongosuchus supports that the clade was present across Pangea, suggesting early ecosystem exploration for Archosauriformes through nearshore environments and leading to ease of dispersal across the Tethys.


2001 ◽  
Vol 38 (5) ◽  
pp. 803-812 ◽  
Author(s):  
David S Berman ◽  
Robert R Reisz ◽  
Thomas Martens ◽  
Amy C Henrici

A new species of the sphenacodontid synapsid Dimetrodon, D. teutonis, is described on the basis of a single, adult specimen consisting of a large portion of the presacral vertebral column. The holotype was collected from the Lower Permian Tambach Formation, lowermost formational unit of the Upper Rotliegend, of the Bromacker quarry locality in the midregion of the Thuringian Forest near Gotha, central Germany. This is the first record of the genus outside of North America and, therefore, provides not only additional biological evidence of a continuous Euramerican landmass during the Early Permian, but also the absence of any major physical or biological barrier to faunal interchange of terrestrial vertebrates. An estimated weight of 14 kg for D. teutonis is half that of the smallest, previously recognized species, D. natalis. Sphenacodontid phylogeny indicates that the diminutive size of D. teutonis represents an autapomorphy and is in general accord with the absence of large-sized, basal synapsid predators at this truly terrestrial upland locality. It is speculated that the diminutive size of D. teutonis was probably an adaptation to a truly terrestrial, relatively uplands existence like that represented by the Bromacker locality. Here it subsisted on small vertebrates (and possibly large invertebrates) of the Bromacker assemblage, in which the dominant members in both size and abundance were herbivorous diadectids, and it was unlikely to encounter large predators.


2013 ◽  
Vol 50 (2) ◽  
pp. 142-147
Author(s):  
George D. Stanley ◽  
Thomas E. Yancey ◽  
Hannah M.E. Shepherd

One of the most distinctive components of the Late Triassic warm-water biota are alatoform, reclining bivalves of the genus Wallowaconcha. Wallowaconcha raylenea was first described from shallow-water, fine-grained Upper Triassic carbonate rocks of the Wallowa terrane, northeastern Oregon, and later found in coeval limestone in the Yukon. Fossils of the family Wallowaconchidae are easily recognized and readily distinguished from other fossil groups by their large size (over a metre in length), alatoform morphology, and especially the chambered wing-like extensions likely associated with photosymbiosis. Several different taxa of Norian age inhabited lagoon and reef-related settings on four separate terranes of western North America (Antimonio terrane, Sonora, Mexico; Wallowa terrane, northeastern Oregon; Stikine terrane in the Yukon; Chulitna terrane of Alaska), which during Triassic time existed as volcanic islands in the eastern Panthalassa Ocean. Outside eastern Panthalassa in the eastern Tethys, two other species of Wallowaconcha come from distant localities in Asia and Arabia. We here report for the first time, in presumed Rhaetian limestone of the upper part of the Parson Bay Formation, Vancouver Island, newly discovered examples of Wallowaconcha. They are from Wrangellia and, based on size and shape of the chambers, are assignable to W. raylenea but unlike other examples they appear to be Rhaetian in age. This species of giant bivalve inhabited warm-water locales outboard of North America during the Late Triassic, and its presence provides possible paleobiogeographic links of Wrangellia with both Stikinia and the Wallowa terrane.


1989 ◽  
Vol 63 (3) ◽  
pp. 374-381 ◽  
Author(s):  
Erik Flügel ◽  
Baba Senowbari-Daryan ◽  
George D. Stanley

An Upper Triassic metaspondyle dasycladacean alga,Diplopora oregonensisn. sp., is described from the Hurwal Formation, southern Wallowa Mountains, northeastern Oregon. It occurs in the accreted Wallowa terrane, which is interpreted as far-travelled relative to the craton of North America. The fossil alga is found in limestone clasts within a limestone–chert–volcanic clast conglomerate of the Hurwal Formation. The new species is related toDiploplora borzaiBystricky, known from the Upper Triassic of the Carpathian Mountains and Sicily, but is distinguished by very small branches and a distinct segmentation of the thalli.Diplopora oregonensisis the first Triassic dasycladacean alga known from the United States, and perhaps from all of North America. The absence of calcareous green algae from rocks of cratonal North America, as well as from most Triassic displaced terranes of the eastern and western Pacific, is in stark contrast to counterparts in the former Tethys region of central Europe, where dasycladacean algae were abundant and contributed significantly to the sediment. This paucity of algae may be related to differences in environment, but more likely is linked to the paleogeographic situation and dispersal abilities of the algae. The similarity of the Oregon dasyclads to species in western Europe, coupled with the lack of dasyclad algae in any other part of North America, is evidence in support of a far-travelled nature for the Wallowa terrane.


2019 ◽  
Vol 15 (2) ◽  
pp. 20180922 ◽  
Author(s):  
Michelle R. Stocker ◽  
Sterling J. Nesbitt ◽  
Ben T. Kligman ◽  
Daniel J. Paluh ◽  
Adam D. Marsh ◽  
...  

Crown-group frogs (Anura) originated over 200 Ma according to molecular phylogenetic analyses, though only a few fossils from high latitudes chronicle the first approximately 60 Myr of frog evolution and distribution. We report fossils that represent both the first Late Triassic and the earliest equatorial record of Salientia, the group that includes stem and crown-frogs. These small fossils consist of complete and partial ilia with anteriorly directed, elongate and distally hollow iliac blades. These features of these ilia, including the lack of a prominent dorsal protuberance and a shaft that is much longer than the acetabular region, suggest a closer affinity to crown-group Anura than to Early Triassic stem anurans Triadobatrachus from Madagascar and Czatkobatrachus from Poland, both high-latitude records. The new fossils demonstrate that crown anurans may have been present in the Late Triassic equatorial region of Pangea. Furthermore, the presence of Early Jurassic anurans in the same stratigraphic sequence ( Prosalirus bitis from the Kayenta Formation) suggests that anurans survived the climatic aridification of this region in the early Mesozoic. These fossils highlight the importance of the targeted collection of microfossils and provide further evidence for the presence of crown-group representatives of terrestrial vertebrates prior to the end-Triassic extinction.


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