Postcranium of the paradigm elasmosaurid plesiosaurianLibonectes morgani(Welles, 1949)

2014 ◽  
Vol 152 (4) ◽  
pp. 694-710 ◽  
Author(s):  
SVEN SACHS ◽  
BENJAMIN P. KEAR
Keyword(s):  
Upper Cretaceous ◽  
Cervical Vertebrae ◽  
Axial Skeleton ◽  
Body Plan ◽  
Pectoral Girdle ◽  
Postcranial Skeleton ◽  
Line Drawings ◽  
Phylogenetic Placement

AbstractElasmosauridae constitutes one of the most immediately recognizable plesiosaurian radiations. Their distinctive body plan represents the popular model for Plesiosauria, and is typified by an osteological morphology especially adapted for hyper-elongation of the neck. Nevertheless, many archetypal elasmosaurids are known only from incomplete and/or inadequately documented material, a problem that has contributed to their uncertain intra-clade relationships. A prime example of this isLibonectes morganifrom the Upper Cretaceous of Texas, USA, which is frequently presented as an elasmosaurid structural proxy because of its three-dimensionally preserved holotype skull. Perplexingly though, both the taxonomic diagnosis and phylogenetic placement ofL. morganirely primarily upon the cervical vertebrae, together with the pectoral girdle and forelimb, yet most of these elements are now lost and figured only as line drawings. We therefore reviewed the remnant postcranial skeleton ofL. morganifirst-hand with the objective of clarifying its defining character states. Our observations showed that the existing diagnosis ofL. morganiis indeed inadequate. Moreover, the only identifiable autapomorphies occurred within the axial skeleton. This concurred with an examination of character scores used in published plesiosaurian phylogenies, and highlights the persistent significance of postcranial elements for discriminating elasmosaurid taxa.

New information on the pectoral girdle and vertebral column in Lupeosaurus (Reptilia, Pelycosauria)

1989 ◽  
Vol 26 (7) ◽  
pp. 1343-1349 ◽  
Author(s):  
Stuart S. Sumida
Keyword(s):  
Cross Section ◽  
Vertebral Column ◽  
Cervical Vertebrae ◽  
Lower Permian ◽  
Pectoral Girdle ◽  
Postcranial Skeleton ◽  
Posterior Edge ◽  
Articular Facet ◽  
New Information

UCLA VP 1651, a new specimen from the Lower Permian Admiral Formation of Archer County, Texas, provides information on heretofore unknown portions of the postcranial skeleton of the large pelycosaur Lupeosaurus. Presacral neural spines are elongate and have a subcircular cross section. Laterally directed tubercles are not present on any of the neural spines. Cervical vertebrae appear to have elongate neural spines and sharply keeled centra. Transverse processes are positioned relatively high above the bases of the centra through the length of axial column. The pattern of tilting of the neural spines is similar to that in the primitive edaphosaur Ianthasaurus hardestii. The proximal end of the first sacral rib contributes significantly to the intervertebral articular facet usually formed by the centra. The sacral ribs do not appear to fuse with one another distally. The ventral clavicular plate is greatly expanded anteriorly. The posterior edge of the subcoracoscapular fossa is very well defined. The available morphological information does not warrant a separate familial designation for the genus Lupeosaurus. Despite the lack of laterally directed tubercles of the neural spines, Lupeosaurus appears to be referrable to the Edaphosauridae. However, in the absence of cranial materials, this association must remain tentative. Although this description adds new information for only a restricted portion of the skeleton, it does appear to confirm the existence of a distinct, albeit rare, pelycosaur from the extensively studied fauna of the Lower Permian.

Infundibulicybe rufa sp. nov. (Tricholomataceae), a reddish brown species from southwestern China

Phytotaxa ◽  
2016 ◽  
Vol 266 (2) ◽  
pp. 134 ◽  
Author(s):  
QI ZHAO ◽  
YAN-JIA HAO ◽  
JIAN-KUI LIU ◽  
KEVIN D. HYDE ◽  
YANG-YANG CUI ◽  
...  
Keyword(s):  
Internal Transcribed Spacer ◽  
Phylogenetic Analyses ◽  
Biosphere Reserve ◽  
Its Region ◽  
Southwestern China ◽  
New Taxon ◽  
Line Drawings ◽  
Phylogenetic Placement ◽  
Molecular Phylogenetic ◽  
Pileus Surface

Infundibulicybe rufa sp. nov., is described from Jiuzhaigou Biosphere Reserve, southwestern China. It is characterized by the combination of the following characters: umbilicate to slightly infundibuliform, reddish brown pileus; decurrent, cream lamellae; cylindrical stipe concolorous with the pileus surface. Molecular phylogenetic analyses using the nuclear ribosomal internal transcribed spacer (ITS) region indicates that I. rufa is closely related to I. mediterranea and I. bresadolana. A description, line drawings, phylogenetic placement and comparison with allied taxa for the new taxon are presented.

CHILD SACRIFICE IN TULA: A BIOARCHAEOLOGICAL STUDY

2021 ◽  
Vol 32 (1) ◽  
pp. 84-99 ◽  
Author(s):  
Angélica María Medrano Enríquez
Keyword(s):  
Young Children ◽  
Cervical Vertebrae ◽  
Human Remains ◽  
Young Age ◽  
Human Sacrifice ◽  
Postcranial Skeleton ◽  
Compound A ◽  
Cut Marks ◽  
Early Postclassic

AbstractRecent salvage excavations conducted in the Early Postclassic city of Tula, Hidalgo, exposed a residential compound containing an open patio, beneath which was discovered a massive burial, designated Feature 5. Human remains involving a minimum of 49 individuals were discovered, many of whom were arranged in a sitting or squatting position. The majority of individuals, including all of the latter, were young children. The bioarchaeological analysis suggests that they were sacrificed. Many individuals exhibited anthropogenic modification, including cut marks on the skull and postcranial skeleton, indicating the children had been flayed. Some individuals were represented only by the skull and cervical vertebrae, suggesting decapitation. The individuals appeared to be in bad health, a common attribute of children sacrificed to Tlaloc by the Aztecs. According to ethnohistorical sources, children offered to Tlaloc commonly had their throats slit, although this practice was not identified among the individuals in Feature 5, possibly owing to their young age. Sacrifice also may have been made to Xipe Totec, as suggested by the evidence of flaying and the presence of a large hollow sculpture of the deity located in an adjacent residential compound, a deity who has been also linked to human sacrifice related to regeneration and fertility.

The locomotor system

2013 ◽  
Author(s):  
Martin E. Atkinson
Keyword(s):  
Vertebral Column ◽  
Cervical Vertebrae ◽  
Lumbar Vertebrae ◽  
Posterior Wall ◽  
Axial Skeleton ◽  
Intervertebral Discs ◽  
Central Canal ◽  
The Body ◽  
Thoracic Vertebrae ◽  
Locomotor System

The locomotor system comprises the skeleton, composed principally of bone and cartilage, the joints between them, and the muscles which move bones at joints. The skeleton forms a supporting framework for the body and provides the levers to which the muscles are attached to produce movement of parts of the body in relation to each other or movement of the body as a whole in relation to its environment. The skeleton also plays a crucial role in the protection of internal organs. The skeleton is shown in outline in Figure 2.1A. The skull, vertebral column, and ribs together constitute the axial skeleton. This forms, as its name implies, the axis of the body. The skull houses and protects the brain and the eyes and ears; the anatomy of the skull is absolutely fundamental to the understanding of the structure of the head and is covered in detail in Section 4. The vertebral column surrounds and protects the spinal cord which is enclosed in the spinal canal formed by a large central canal in each vertebra. The vertebral column is formed from 33 individual bones although some of these become fused together. The vertebral column and its component bones are shown from the side in Figure 2.1B. There are seven cervical vertebrae in the neck, twelve thoracic vertebrae in the posterior wall of the thorax, five lumbar vertebrae in the small of the back, five fused sacral vertebrae in the pelvis, and four coccygeal vertebrae—the vestigial remnants of a tail. Intervertebral discs separate individual vertebrae from each other and act as a cushion between the adjacent bones; the discs are absent from the fused sacral vertebrae. The cervical vertebrae are small and very mobile, allowing an extensive range of neck movements and hence changes in head position. The first two cervical vertebrae, the atlas and axis, have unusual shapes and specialized joints that allow nodding and shaking movements of the head on the neck. The thoracic vertebrae are relatively immobile. combination of thoracic vertebral column, ribs, and sternum form the thoracic cage that protects the thoracic organs, the heart, and lungs and is intimately involved in ventilation (breathing).

The Axial Skeletons of Sunghir 1, 2, and 3

2014 ◽  
Author(s):  
Erik Trinkaus ◽  
Alexandra P. Buzhilova ◽  
Maria B. Mednikova ◽  
Maria V. Dobrovolskaya
Keyword(s):  
Radiocarbon Dating ◽  
Cervical Vertebrae ◽  
Costal Cartilage ◽  
Lumbar Vertebrae ◽  
Axial Skeleton ◽  
Age At Death ◽  
Vertebral Bodies ◽  
Length Scaling

Given their burial positions, on their backs with the trunks and limbs extended, the Sunghir 1 to 3 individuals should have retained major portions of their axial skeletons. This is the case for Sunghir 2 and 3, both of whom retain all of the cervical vertebrae, most of their thoracic and lumbar vertebrae, and major portions of their sacra. Sunghir 2 preserves portions of 23 of the 24 ribs, and Sunghir 3 retains at least a small piece of each of her 24 ribs. Moreover her left fifth and sixth ribs lack only their costal cartilage surfaces. Only Sunghir 3 preserves any elements of the sternum, two partial and separated sternebral segments. In contrast, despite the apparent presence of major portions of the axial skeleton in situ, little remains of the Sunghir 1 vertebrae, ribs, or sternum. The cervical vertebrae are absent, unless pieces of them are mixed with the collection of what appear to be thoracic and lumbar fragments. Only two vertebrae remain reasonably intact, the T1 and T2. There are eight pieces of vertebral bodies, one of which has a pathological growth (chapter 17). The ribs consist of small pieces, except for a largely intact left first rib. Although evident in the in situ photographs, nothing remains of the manubrium. There is also a piece of distal middle rib, which is of use for the age-at-death assessment. Some of the vertebral and rib pieces have been sacrificed over the years for direct radiocarbon dating (e.g., Kuzmin et al. 2004). Others pieces, heavily fissured and hence probably descending into fragments during excavation, were only partially retained. There are nonetheless a few aspects of the Sunghir axial skeletons, beyond age assessments (chapter 6), the pathological lesions on the Sunghir 1 vertebrae (chapter 17), use of the sacra in the pelves (chapter 14), and body length scaling for Sunghir 2 and 3 (chapter 11), that are of interest.

A new marine clupeoid fish from the Lower Cretaceous of the Sergipe-Alagoas Basin, northeastern Brazil

Zootaxa ◽  
2009 ◽  
Vol 2164 (1) ◽  
pp. 21-32 ◽  
Author(s):  
FRANCISCO J. DE FIGUEIREDO
Keyword(s):  
Fossil Record ◽  
Upper Cretaceous ◽  
Democratic Republic ◽  
Democratic Republic Of Congo ◽  
Axial Skeleton ◽  
Northeastern Brazil ◽  
Unique Combination ◽  
Caudal Vertebrae ◽  
Clupeoid Fish ◽  
Marine Deposit

Clupeomorph fishes are largely diversified and widespread in Cretaceous strata of northeastern Brazil. They are represented by basal clupeiforms, †ellimmichthyiforms, and advanced clupeoids. In this paper, a new clupeoid fish, †Nolfia riachuelensis sp. nov., is described based on a specimen found in marine shale yielded in the Taquari Member (Albian) of Riachuelo Formation, Sergipe-Alagoas Basin. Although partially preserved (most of skull is lacking), the fish shows many informative characters, particularly in the axial skeleton. The fish is morphologically very similar to †Nolfia kwangoensis, a clupeoid fish originally described from marine deposit from the Upper Cretaceous (Cenomanian) of Kipala, Democratic Republic of Congo, with which it shares: 18 caudal vertebrae; long and cylindrical pleural ribs; posteriormost pleural ribs supported by long parapophyses and gradually decreasing in size posteriorly; very reduced ural centra; presence of long and distally congruent neural spines of third, second, and first preural centra; three uroneural bones (the first probably forming the pleurostyle). Otherwise, †Nolfia riachuelensis sp. nov. differs from other clupeomorphs by a unique combination of features. The number of supraneural bones, aligned rod-like postcleithra, less than 30 rays in the anal fin, and derived absence of dorsal scutes indicate its placement within Clupeidae. As far as is known, †Nolfia riachuelensis sp. nov. is the most ancient clupeoid fish known from the fossil record.

scholarly journals The postcranial skeleton of Vagaceratops irvinensis (Dinosauria, Ceratopsidae)

2014 ◽  
Vol 1 ◽  
pp. 1 ◽  
Author(s):  
Robert B Holmes
Keyword(s):  
Latissimus Dorsi ◽  
Vertebral Column ◽  
Cervical Vertebrae ◽  
Dorsal Surface ◽  
Ventral Surface ◽  
Advanced Age ◽  
Time Of Death ◽  
Postcranial Skeleton ◽  
Hind Limbs ◽  
Left Front

The postcranial skeleton of Vagaceratops (= Chasmosaurus) irvinensis (CMN 41357), lacking only the tail, most of the left front and left hind limbs, and portions of the pelvis, is preserved in articulation. It is one of the most complete, best articulated ceratopsid skeletons known. Both the manus and vertebral column exhibit conspicuous pathologies, possibly an indication of advanced age at the time of death. The vertebral column comprises a syncervical, six additional cervical vertebrae, and 12 dorsals. A partial synsacrum is represented by two dorsosacrals, four sacrals, two caudosacrals, and a partial third caudosacral centrum. The ribcage, although crushed, is nearly complete. The sternum is unusually wide compared with other ceratopsids. As in other chasmosaurines, the humerus bears a prominent deltopectoral crest that forms the anterior edge of the broad, rectangular proximal humeral expansion. The medial tuberosity is separated from the dorsal surface of the humerus by a distinct notch. The insertion for the latissimus dorsi is conspicuous. The deltopectoral crest extends a full half of the distance to the distal end of the humerus. Epipodials of the forelimb are relatively short compared to the corresponding propodial. The ulna has a long, distinctly triangular olecranon, broadly rounded anterolateral process, prominent medial process, and a deeply concave trochlear notch. The terminal phalanges on the fourth and fifth manual digits are relatively large, and unlike other ceratopsids have distinct distal ?articular facets. The fourth trochanter of the femur is relatively proximal in position. This study and other recent studies of ceratopsid postcrania suggest that potentially useful taxonomic variation is present in the number of dorsosacrals, size of the groove on the ventral surface of the sacrum, morphology of the last dorsal and dorsosacral ribs, morphology of the scapula and proximal expansion of the humerus, morphology of the ulna, ratio of humerus/epipodium, morphology of the fifth manual digit, and position of the fourth trochanter of the femur.

scholarly journals Problematica old and new

2008 ◽  
Vol 363 (1496) ◽  
pp. 1503-1512 ◽  
Author(s):  
Ronald A Jenner ◽  
D.Timothy J Littlewood
Keyword(s):  
Molecular Phylogenetics ◽  
Body Plan ◽  
Phylogeny Reconstruction ◽  
Phylogenetic Placement ◽  
Genomic Markers ◽  
Body Plan Evolution ◽  
Metazoan Phylogeny

Problematica are taxa that defy robust phylogenetic placement. Traditionally the term was restricted to fossil forms, but it is clear that extant taxa may be just as difficult to place, whether using morphological or molecular (nucleotide, gene or genomic) markers for phylogeny reconstruction. We discuss the kinds and causes of Problematica within the Metazoa, as well as criteria for their recognition and possible solutions. The inclusive set of Problematica changes depending upon the nature and quality of (homologous) data available, the methods of phylogeny reconstruction and the sister taxa inferred by their placement or displacement. We address Problematica in the context of pre-cladistic phylogenetics, numerical morphological cladistics and molecular phylogenetics, and focus on general biological and methodological implications of Problematica, rather than presenting a review of individual taxa. Rather than excluding Problematica from phylogeny reconstruction, as has often been preferred, we conclude that the study of Problematica is crucial for both the resolution of metazoan phylogeny and the proper inference of body plan evolution.

Aspects of the postcranial skeleton of the Lower Permian lungfish, Gnathorhiza

1989 ◽  
Vol 63 (6) ◽  
pp. 919-930 ◽  
Author(s):  
Walter W. Dalquest ◽  
M. John Kocurko ◽  
John V. Grimes
Keyword(s):  
Axial Skeleton ◽  
Lower Permian ◽  
Fish Scales ◽  
Early Permian ◽  
Postcranial Skeleton ◽  
Caudal Fin ◽  
North Central ◽  
New Information ◽  
Central Texas ◽  
Median Fins

A newly discovered locality of aestivation burrow casts containing the lungfish, Gnathorhiza serrata, is reported from the early Permian Arroyo Formation of Wilbarger County, north-central Texas. Remains preserved in the burrow casts provide sections of mummified Gnathorhiza and new information about the postcranial skeleton of this fish. Scales of Gnathorhiza resemble those of the modern lungfishes such as Lepidosiren in their microanatomy. No traces of paddle-like pectoral or pelvic fins were found and paired fins of Gnathorhiza may have resembled those of Lepidosiren. The axial skeleton and median fins of Gnathorhiza seem to resemble those of Lepidosiren except that the tail area and caudal fin of Gnathorhiza were stout and strong rather than slender and tapering. Gnathorhiza, which aestivated tail-down in its burrow, may have required a stout tail for support.

scholarly journals A comprehensive anatomical and phylogenetic evaluation of Dilophosaurus wetherilli (Dinosauria, Theropoda) with descriptions of new specimens from the Kayenta Formation of northern Arizona

2020 ◽  
Vol 94 (S78) ◽  
pp. 1-103 ◽  
Author(s):  
Adam D. Marsh ◽  
Timothy B. Rowe
Keyword(s):  
Phylogenetic Analyses ◽  
Cervical Vertebrae ◽  
Pop Culture ◽  
Single Species ◽  
Axial Skeleton ◽  
Late Triassic ◽  
Monophyletic Clade ◽  
Navajo Nation ◽  
Northern Arizona ◽  
Nasal Process

AbstractDilophosaurus wetherilli was the largest animal known to have lived on land in North America during the Early Jurassic. Despite its charismatic presence in pop culture and dinosaurian phylogenetic analyses, major aspects of the skeletal anatomy, taxonomy, ontogeny, and evolutionary relationships of this dinosaur remain unknown. Skeletons of this species were collected from the middle and lower part of the Kayenta Formation in the Navajo Nation in northern Arizona. Redescription of the holotype, referred, and previously undescribed specimens of Dilophosaurus wetherilli supports the existence of a single species of crested, large-bodied theropod in the Kayenta Formation. The parasagittal nasolacrimal crests are uniquely constructed by a small ridge on the nasal process of the premaxilla, dorsoventrally expanded nasal, and tall lacrimal that includes a posterior process behind the eye. The cervical vertebrae exhibit serial variation within the posterior centrodiapophyseal lamina, which bifurcates and reunites down the neck. Iterative specimen-based phylogenetic analyses result in each of the additional specimens recovered as the sister taxon to the holotype. When all five specimens are included in an analysis, they form a monophyletic clade that supports the monotypy of the genus. Dilophosaurus wetherilli is not recovered as a ceratosaur or coelophysoid, but is instead a non-averostran neotheropod in a grade with other stem-averostrans such as Cryolophosaurus ellioti and Zupaysaurus rougieri. We did not recover a monophyletic ‘Dilophosauridae.’ Instead of being apomorphic for a small clade of early theropods, it is more likely that elaboration of the nasals and lacrimals of stem-averostrans is plesiomorphically present in early ceratosaurs and tetanurans that share those features. Many characters of the axial skeleton of Dilophosaurus wetherilli are derived compared to Late Triassic theropods and may be associated with macropredation and an increase in body size in Theropoda across the Triassic-Jurassic boundary.

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