The effect of calcium supply on bur set and seed formation in subterranean clover (Trifolium subterraneum L.)

1973 ◽  
Vol 24 (6) ◽  
pp. 839 ◽  
Author(s):  
PG Ozanne ◽  
KMW Howes

In a glasshouse experiment subterranean clover plants were grown in soil at various levels of calcium nutrition. The developing burs were buried in separate soil containers to which no nutrients, calcium sulphate, calcium carbonate, or potassium sulphate were applied. When subterranean clover was growing in soil low in calcium, top yield was little affected; but bur and seed formation were greatly reduced. However, when the reflexed flowers were buried in soil to which a calcium salt had been added, bur and seed yield increases of up to 500% were obtained The increase in seed yield resulted from a greater percentage of flowers forming bur, more seed per bur, heavier seed, and less rotten seed. Similar results were obtained in a second glasshouse experiment in which a different soil was used to grow the clover. The beneficial effect of calcium ions around the developing bur was independent of the associated anions, or of the soil pH; nor could it be produced by the substitution of potassium or magnesium for calcium. It seems likely that commercial seed yields and subterranean clover persistence in many areas may be reduced by calcium deficiency in apparently lush healthy pastures. The similarity of this disorder to calcium deficiency in peanuts suggests that other crop and pasture species forming buried seed may also be especially susceptible.

1987 ◽  
Vol 38 (3) ◽  
pp. 537 ◽  
Author(s):  
BS Dear ◽  
J Lipsett

There has been little research on the boron (B) status of subterranean clover, despite strong indications of deficiency of B in southern Australia where clover decline has been reported. This paper describes glasshouse experiments to vary the level of B supply to clover grown on soils low in B. On a soil farmed in a cereal clover rotation, herbage yields of clover increased by 25% with applied B, but seed yields, negligible without B, increased 21-fold. On two other acid soils, from pastures, there were also large responses to B in seed yield (1.5- and 5.1-fold respectively); liming increased the responses (8- and 55.2-fold). However, herbage production was less responsive to B (25% and 1.7-fold with liming); on the unlimed soils, application of B depressed yield (- 1 and -21%). This is attributed to there being enough B to sustain herbage growth, but inadequate for seed formation. These results show that herbage yield, conventionally employed in the past, is not a satisfactory index of B status for total performance in subterranean clover. The response to B in seed yield was associated with increases in: numbers of seed set (75% to 22.6-fold, depending on the particular combination of soil and liming); size of seed (10% to 1.2-fold); proportion of buried seed (20%-70%). It is suggested that these favour establishment and persistence of clover and that clover decline may well involve deficiency of B. It was shown that concentration of B in the plant does not necessarily relate predictably to yield of herbage and that reduction in seed yield may not be heralded by foliar symptoms, since seed requirement exceeds that of herbage.


1967 ◽  
Vol 7 (24) ◽  
pp. 25 ◽  
Author(s):  
GB Taylor ◽  
RC Rossiter

Seed production and persistence of the Carnamah, Northam A, Dwalganup, and Geraldton strains of subterranean clover (Trifolium subterraneum L.) were examined in undefoliated swards in the wheatbelt of Western Australia. The early flowering characteristic of Carnamah was not always associated with higher seed yields. Only when there was a well-defined, early finish to the growing season, or when flowering was very much earlier in Carnamah (viz., following an early 'break' to the season), did this strain clearly outyield both Northam A and Geraldton. The seed yield of Dwalganup was generally inferior to that of the other strains. Factors affecting regeneration are discussed. Under low rainfall conditions, poorer germination-regulation of Carnamah, compared with Geraldton and Northam A, would be expected to result in poorer persistence unless offset by higher seed yields in the Carnamah strain.


2008 ◽  
Vol 48 (8) ◽  
pp. 1133 ◽  
Author(s):  
B. S. Dear ◽  
B. F. Hackney ◽  
G. M. Dyce ◽  
C. A. Rodham

Swards of four cultivars of subterranean clover (Trifolium subterraneum L.) were cut at three different times to determine the effect on forage yield and quality, seed set and seedling regeneration in two successive seasons in southern New South Wales. The four cultivars of subterranean clover (Seaton Park LF, Junee, Goulburn and Clare) were cut on 23–25 September (early cut), 8–10 October (mid cut) or 22–23 October (late cut), to simulate an early silage, late silage or hay cut. Additional treatments imposed included either grazing or leaving the regrowth after cutting and raising the cutting height from 3 to 6 cm. Forage yields ranged from 3.5 to 9.3 t dry matter (DM)/ha in the first year and from 2.0 to 5.9 t DM/ha in the second year. Herbage yield was influenced by both cultivar and harvest time with the highest yields achieved with the mid cut. Lower forage yields at the later cut were attributed to losses due to respiration and decay under dense leaf canopies. Changes in forage quality were consistent across both years, with in vivo DM digestibility declining from 76–79% to 69–70% as cutting time was delayed. Crude protein fell from 22–24% to 14–17% over the same period, depending on cultivar. Seed yields in both years were influenced by both cutting time and cultivar with a positive relationship (R2 = 0.45–0.61) between herbage present in late spring after a period of regrowth and subsequent seed yield. The early flowering cultivar Seaton Park LF had the highest seed yield in both years and the more erect cultivar Clare had the lowest. Seed yields declined with later cutting time but increased by an average of 39% when the cutting height was raised from 3 to 6 cm. Seedling regeneration reflected seed yield responses with the largest seedling regeneration occurring in treatments cut early. The study found that forage conservation in early October is likely to yield more and be of higher quality than swards cut later in the season. Seed set is greatly reduced by all cutting strategies to levels unprofitable for seed harvesting but may be adequate for pasture regeneration.


1995 ◽  
Vol 35 (4) ◽  
pp. 475 ◽  
Author(s):  
GA Sandral ◽  
BS Dear ◽  
NE Coombes

The effect of broadleaf herbicides on seed set by Trifolium subterraneum (subterranean clover) cultivars was examined at 2 sites (Wagga Wagga and Canowindra) over 2 years. Five commonly used herbicide treatments (bromoxynil, MCPA, 2,4-DB, MCPA + terbutryn, MCPA + diuron) were applied at 2 rates to 7 cultivars of subterranean clover. Significant site x cultivar x herbicide interactions were observed. Seed yields were either unaffected or depressed by up to 66% at the higher rainfall site (Canowindra), whereas at the lower rainfall site (Wagga Wagga) some herbicide x cultivar combinations showed increases in seed yield up to 115%. The increase in seed yield was greater at the lower herbicide rate. The variation in seed yield with herbicide treatment was largely a result of a change (P<0.001) in the number of seeds set (R = 0.94 at Wagga Wagga; R = 0.85 at Canowindra). Seed size was also correlated (P<0.001) with seed yield at both sites but explained less of the variation (R = 0.23 at Wagga Wagga; R = 0.47 at Canowindra). The reduction in herbage yield as a result of herbicide application was a poor indicator of the subsequent seed yield response at both sites. Cultivar Trikkala was consistently most tolerant to the herbicide treatments, showing either no change in seed yield at the wetter site or large increases in seed yield at the lower rainfall site. In contrast, the seed yield of Dalkeith was depressed by 2,4-DB at both sites in both years by 39-66%. Increases in seed yield, which were most pronounced in cultivars of midseason maturity (Trikkala, Junee, Seaton Park) and least in later maturing cultivars (Karridale, Clare) and in the very early flowering Dalkeith, were attributed in part to a water-saving effect of the herbicide treatments. Cultivar maturity ranking was negatively correlated (P<0.001) with seed yield (R=-0.73 at Wagga Wagga; R=-0.45 at Canowindra). The germination percentage of seed produced by the cultivars was unaffected by herbicide treatment, although the number of abnormal radicles formed by germinating seed was higher in the 2,4-DB and MCPA treatments.


1975 ◽  
Vol 15 (72) ◽  
pp. 69 ◽  
Author(s):  
GH Walton

Four defoliation treatments before flowering were imposed on twelve genotypes from three subspecies of Trifolium subterraneum. The experiment was conducted on a sandy soil with irrigation supplementing the natural rainfall. The severest defoliation doubled the percentage and tripled the absolute yield of buried seed. Highly significant differences were found between the genotypes within subspecies, indicating that selection for subterranean clover pastures with a range of ability to bury burrs is possible. No differences in genotype or subspecies response to defoliation nor differences of absolute seed yields were obtained. It is suggested that sward density is very important when experiments evaluating subterranean clover genotypes include defoliation treatments of the swards.


1981 ◽  
Vol 32 (3) ◽  
pp. 445 ◽  
Author(s):  
RC Rossiter ◽  
MJ Palmer

Two experiments using various strains of subterranean clover are described. The first experiment was conducted in the field with mixtures of Dwalganup/Northam A, DwalganupI/Daliak and Yarloop/Seaton Park, and the second in an open-sided glasshouse with Yarloop/Seaton Park. In the field experiment, the plots were grazed continuously from late July until early November when the swards were completely dry; in the glasshouse experiment, the swards were defoliated weekly until shortly after the beginning of flowering. Mixtures were arranged according to the de Wit replacement series. Seed yields for the Dwa1ganup/Northam A mixture showed the classical (de Wit) competition for the same resources, with Northam A the better competitor. On the other hand, for the Dwalganup/Daliak and Yarloop/Seaton Park mixtures there was no clear evidence of a competitive advantage of one strain over the other pair member, at least under our defoliated conditions. Whether or not this absence of 'competition' (as usually understood) is common in clover-strain mixtures is as yet unknown. We emphasize, however, that long-term success of clover strains in mixtures is not invariably associated with competitive advantage.


Soil Research ◽  
2000 ◽  
Vol 38 (3) ◽  
pp. 735 ◽  
Author(s):  
Robert Summers ◽  
Martin Clarke ◽  
Tim Pope ◽  
Tim O'Dea

Bauxite residue from alumina refining was used to coat granules of single superphosphate to reduce the leaching of phosphorus in coarse, sandy soils for pastures in high rainfall areas of south-western Australia (>800 mm annual average). The impact of coating the superphosphate on the leaching of phosphorus was measured in a glasshouse experiment and the effectiveness of the fertiliser using dry herbage yield of subterranean clover (Trifolium subterraneum) was measured in a field experiment. The glasshouse experiment measured the effect of coating the superphosphate with bauxite residue at 0, 5, 10, 15, 20, 25, 30, 35, and 40% by weight. A coating of 25% (by weight) was chosen for the field experiment. In the glasshouse experiment, the coated granules were applied to columns of soil, where subterranean clover was grown under leaching conditions. A coating of 30%, by weight, reduced leaching of single superphosphate by about half. Increasing the coating of bauxite residue also increased the phosphorus uptake and increased the plant growth. In the field trial, the effectiveness of single superphosphate with a bauxite residue coating of 25% by weight was increased on average by 100% in Year 1, 303% in Year 2, and 158% in Year 3, relative to freshly applied single superphosphate. The bauxite residue coating also increased the phosphorus content of the herbage in a similar manner to the increases in yield. Limited soil phosphorus tests showed only minor increases in the residues of phosphorus where the superphosphate had been coated with bauxite residue.


2007 ◽  
Vol 58 (2) ◽  
pp. 123 ◽  
Author(s):  
B. S. Dear ◽  
G. A. Sandral ◽  
J. M. Virgona ◽  
A. D. Swan ◽  
B. A. Orchard ◽  
...  

The effect of the density of 3 perennial species, phalaris (Phalaris aquatica L.), wallaby grass (Austrodanthonia richardsonii Kunth), and lucerne (Medicago sativa L.), on seed set, regeneration, and the relative competitiveness of 3 cultivars of subterranean clover (Trifolium subterraneum L.) was examined in 2 environments in the south-eastern Australian wheatbelt. Seed yields of subterranean clover were inversely related to perennial density at both sites over the first 2 years, the relationship varying with perennial species. Phalaris depressed the seed yield of clover more than lucerne and wallaby grass in the second and third year at equivalent densities. Clover seed yield was positively related to clover herbage yield in late spring at both sites, and inversely related to perennial herbage yield. Clover seed yield displayed an increasing linear relationship with the proportion of light reaching the clover understorey in spring, which in turn was inversely related to perennial density and perennial herbage yield. Clover seedling regeneration in mixed swards in autumn was positively related to the size of the summer seed bank, but negatively related to perennial density. Clover seedling survival following a premature germination at Kamarah was inversely correlated to the density of phalaris and lucerne in the sward. The relative competitiveness of the 3 subterranean clover cultivars varied between sites, with climatic conditions (rainfall and growing-season length) having a greater effect on the relative cultivar performance than companion perennial species or density. The later maturing subterranean clover cv. Goulburn became the dominant cultivar at the wetter site, constituting 72% of the seed bank, but declined to only 3–8% of the seed bank at the drier site. The proportion of the early flowering cultivar Dalkeith in the seed bank increased over time at the drier site and was highest (53%) in plots with the highest perennial density. We concluded that although perennial pasture species will depress clover seed yield and subsequent regeneration, these effects could be minimised by reducing perennial densities and exploiting variations in competitiveness between perennial species as identified in this study. Sowing earlier maturing subterranean clover cultivars would only be an advantage in increasing clover content in low-rainfall environments. The findings suggest that clover seed reserves and regeneration could also be increased by using grazing management to reduce the level of shading of clover by perennials, a factor associated with reduced clover seed yield.


1981 ◽  
Vol 32 (5) ◽  
pp. 783 ◽  
Author(s):  
WJ Collins

The effects of length of growing season and defoliation on seed yield and hard-seededness were examined in two strains of subterranean clover (Seaton Park, Midland B) grown in swards in the field. All plots were sown at the same time and the length of growing season was varied by altering the time of finish of the season (by withholding water). There were three length of growing season treatments: T1 (short), T2 (intermediate) and T3 (control). The defoliation treatments were D0, uncut (control), and D1, defoliated at weekly intervals until the commencement of flowering. Reducing the length of growing season drastically reduced seed yield. Thus when the growing season was only 3 weeks shorter than the control (i.e. T2 compared with T3), seed yields averaged over strains and defoliations were reduced by at least one half. With a further reduction of 2 weeks in the length of the season (T1) seed yields were only about one-third of those obtained in the control (T3). The reductions in seed yield were due to reductions in both the number of mature burrs produced and to a lesser extent in mean weight per seed. Although defoliation increased seed yield in all growing seasons, the effect when measured on a relative scale was greater in T2 than in either T1 or T3. But on an absolute scale the size of the response was greater in T2 and T3 than in T1. The rate of breakdown of hard-seededness was faster in Seaton Park than in Midland B, but it decreased in both strains with increasing length of growing season.


1978 ◽  
Vol 29 (6) ◽  
pp. 1167 ◽  
Author(s):  
WJ Collins ◽  
RC Rossiter ◽  
MA Ramas

Swards of three strains of subterranean clover (Woogenellup, Clare and Seaton Park) were subjected to three shading treatments (30%, 55% and 100% of daylight) from the beginning of flowering onwards. For the 100% daylight (unshaded) treatment the incoming daily solar radiation averaged between 20 and 30 MJ/m2. In the unshaded treatment, seed yields for the three strains ranged from about 120 to almost 200 g/m2. These yields were invariably decreased by shading. Moreover the evidence indicated a curvilinear relationship between seed yield and light supply such that at 50% of daylight there was a reduction of rather more than 50% in seed yield. Shading mainly affected the number of inflorescences produced per unit area, although other components of seed yield were also affected adversely. Even late shading (towards the end of flowering) led to lower seed yield, partly associated with lower inflorescence numbers and partly with lower seed numbers per burr. The results are considered to have significance for clover regeneration in mixed clover/grass pastures and also in the cloveritree associations of agro-forestry.


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