Structure and dynamics of trembling aspen – white spruce mixed stands near Fort Nelson, B.C.

2004 ◽  
Vol 34 (2) ◽  
pp. 384-395 ◽  
Author(s):  
Richard Kabzems ◽  
Oscar García

The trembling aspen (Populus tremuloides Michx.) – white spruce (Picea glauca (Moench) Voss) mixed woods near Fort Nelson are distinguished by the large size of individual trees, longevity, and the low occurrence of internal decay in trembling aspen. The development of these forest ecosystems has had limited documentation and may be significantly different than those described in other portions of the boreal forest. At five study stands, stem analysis techniques were used to examine the patterns of height and radial growth over time according to species and structure type. There were two patterns of species establishment that were consistent with the stand structure. In codominant stands, recruitment periods for trembling aspen and white spruce overlapped. The stratified stands were consistently associated with a 29- to 58-year lag in white spruce recruitment. Spruce that were codominant with aspen at the time of sampling had sustained periods of rapid height and diameter growth. White spruce that were later to establish on site had slower rates of height and diameter growth. White spruce ages indicated that a dominant recruitment episode was more common than continuous recruitment. Height and diameter growth of trembling aspen were similar in both stand types. The differences in trembling aspen growth patterns between stands were due to site quality. The white spruce in codominant stands did not appear to go through a period of suppression and then release associated with stand-level trembling aspen mortality, as commonly described for other boreal mixedwoods. The vigor and longevity of trembling aspen in Fort Nelson appear to prolong the period of trembling aspen domination of mixed stands well beyond the time periods observed in other boreal ecosystems.

2011 ◽  
Vol 41 (4) ◽  
pp. 728-739 ◽  
Author(s):  
Robert L. Fleming ◽  
Allister D. Smith

Many of the major questions regarding stand establishment practices involve implications for longer-term ecosystem development. We examined 41-year treatment effects on stand composition and dynamics using a white spruce ( Picea glauca (Moench) Voss) planting, mechanical site preparation (MSP) – herbicide (2,4-D plus 2,4,5-T) trial in a trembling aspen ( Populus tremuloides Michx.) dominated mixedwood. Both barrel and blade MSP with planting increased total and white spruce year 41 stand-level biomass over that in untreated areas. Year 2 herbicide application reduced year 41 trembling aspen biomass without substantially increasing that of white spruce, resulting in total yields similar to those in untreated areas. Barrel MSP increased year 41 trembling aspen biomass over that of untreated areas whereas blade MSP reduced it. Herbicide-related declines in trembling aspen biomass persisted or increased with time whereas white spruce response to herbicide varied with time and MSP. By accounting for inherent growth patterns, age shift calculations gave more balanced temporal depictions of planted white spruce response than effect size or percentage gain calculations. With barrel MSP, stand composition demonstrated a degree of mixedwood homeostasis whereas with blading, trembling aspen composition declined unilaterally from year 20 to 41.


2004 ◽  
Vol 80 (5) ◽  
pp. 583-597 ◽  
Author(s):  
Douglas G Pitt ◽  
Milo Mihajlovich ◽  
Leslie M Proudfoot

Twelve Alberta forest regeneration blocks, situated on representative white spruce (Picea glauca (Moench) Voss) - trembling aspen (Populus tremuloides Michx.) boreal mixedwood sites, planted to white spruce, and operationally released with glyphosate herbicide, were surveyed in the fall of 2002. Stand structure and composition were quantified and compared for treated and untreated portions of each block. The Mixedwood Growth Model (MGM, Department of Renewable Resources, University of Alberta) was used to project these stands over a 100-year horizon and to model the outcomes of several additional silvicultural treatments that could be applied to these blocks. A single release treatment provided 17% and 43% gains in planted white spruce height and stem diameter, respectively, an average of five years after treatment. Treatment shifted stands from being deciduous-dominated, with only 12% conifer basal area, to more than 75% conifer basal area, increasing conifer volumes per hectare nearly three-fold, but retaining conifer-deciduous mixture. Model projections suggest that these stands will produce similar total volumes over an 80-year rotation and that conifer release essentially trades deciduous volume for conifer volume, the degree of release dictating the extent to which this trade-off takes place. A single conifer release treatment led to an average simulated mature stand that contains 21% deciduous basal area, likely meeting mixedwood rather than conifer regeneration criteria. Model simulations of additional silvicultural interventions in these stands suggested that a variety of options exist to satisfy a range of stand or landscape management objectives for spruce-aspen mixedwoods, all within a relatively fixed volume production envelope. A clearer understanding of how early stand conditions translate into stand and landscape management objectives seems prerequisite to solving management conflicts on boreal mixedwood sites. Key words: boreal mixedwoods, vegetation management, conifer release, Mixedwood Growth Model, white spruce, trembling aspen


2007 ◽  
Vol 22 (3) ◽  
pp. 163-170 ◽  
Author(s):  
Ryan J. Klos ◽  
G. Geoff Wang ◽  
Qing-Lai Dang ◽  
Ed W. East

Abstract Kozak's variable exponent taper equation was fitted for balsam poplar (Populus balsamifera L.), trembling aspen (Populus tremuloides Michx.), white spruce (Picea glauca [Moench] Voss), black spruce (Picea mariana [Mill.] B.S.P.), and jack pine (Pinus banksiana Lamb.) in Manitoba. Stem taper variability between two ecozones (i.e., Boreal Shield and Boreal Plains) were tested using the F-test. Regional differences were observed for trembling aspen, white spruce, and jack pine, and for those species, separate ecozone-specific taper equations were developed. However, the gross total volume estimates using the ecozone-specific equations were different from those of the provincial equations by only 2 percent. Although the regional difference in stem form was marginal within a province, a difference of approximately 7 percent of gross total volume estimation was found when our provincial taper equations were compared with those developed in Alberta and Saskatchewan. These results suggest that stem form variation increases with spatial scale and that a single taper equation for each species may be sufficient for each province.


2014 ◽  
Vol 44 (2) ◽  
pp. 162-171 ◽  
Author(s):  
Derek F. Sattler ◽  
Philip G. Comeau ◽  
Alexis Achim

Radial patterns of modulus of elasticity (MOE) were examined for white spruce (Picea glauca (Moench) Voss) and trembling aspen (Populus tremuoides Michx.) from 19 mature, uneven-aged stands in the boreal mixedwood region of northern Alberta, Canada. The main objectives were to (1) evaluate the relationship between pith-to-bark changes in MOE and cambial age or distance from pith; (2) develop species-specific models to predict pith-to-bark changes in MOE; and (3) to test the influences of radial growth, relative vertical height, and tree slenderness (tree height/DBH) on MOE. For both species, cambial age was selected as the best explanatory variable with which to build pith-to-bark models of MOE. For white spruce and trembling aspen, the final nonlinear mixed-effect models indicated that an augmented rate of increase in MOE occurred with increasing vertical position within the tree. For white spruce trees, radial growth and slenderness were found to positively influence maximum estimated MOE. For trembling aspen, there was no apparent effect of vertical position or radial growth on maximum MOE. The results shed light on potential drivers of radial patterns of MOE and will be useful in guiding silvicultural prescriptions.


2006 ◽  
Vol 36 (7) ◽  
pp. 1818-1833 ◽  
Author(s):  
Daniel A MacIsaac ◽  
Philip G Comeau ◽  
S Ellen Macdonald

This study assessed the dynamics of gap development in postharvest regeneration in five stands in northwestern Alberta dominated by trembling aspen (Populus tremuloides Michx.). The pattern of gap development over time was determined from analysis of air photographs taken preharvest and 1, 4, 10, and 12 years postharvest. The area of each stand covered by gaps increased after harvest because of the addition of harvest-related gaps over and above those that had been present prior to harvest. The blocks we studied had a combined gap area of up to 29% of stand area 12 years postharvest. We measured regeneration characteristics, microsite, soil, light, and browse conditions in 30 aspen regeneration gaps (gaps in regeneration that were not gaps preharvest and were not due to obvious harvest-related disturbance) 14 years following harvest. Although deciduous trees within postharvest regeneration gaps were the same age as those outside (i.e., in a fully stocked matrix of newly established even-aged aspen stems), they were often suppressed, with significantly lower density and growth. Within the 14-year-old postharvest regenerating aspen stands, aspen height varied from 1 to 11 m; this substantial variability appeared to be largely due to the influence of browsing. There was little evidence of ongoing regeneration within postharvest regeneration gaps, indicating that these gaps will probably persist over time. This may impact future deciduous stocking and volume. It is unknown what may have initiated the formation of these gaps, although results suggest that they are not due to edaphic conditions or disease in the preharvest stands. There is evidence that bluejoint (Calamagrostis canadensis (Michx.) Beauv.) cover and browsing are important factors in the maintenance of postharvest regeneration gaps. The spatial heterogeneity resulting from gaps could be advantageous, however, either as part of ecosystem-based management emulating natural disturbance or as a template for mixedwood management, where white spruce (Picea glauca (Moench) Voss) are established in gaps.


2019 ◽  
pp. 297-307
Author(s):  
Yuqing Yang ◽  
Shongming Huang ◽  
Robert Vassov ◽  
Brad Pinno ◽  
Sophan Chhin

Climate-sensitive height–age models were developed for top height trees of trembling aspen (Populus tremuloides Michx.), jack pine (Pinus banksiana Lamb.), and white spruce (Picea glauca (Moench) Voss) in natural and reclaimed oil sands stands. We used stem analysis data collected from the Athabasca oil sands region in northern Alberta, Canada, and climate data generated by the ClimateWNA model. Height–age trajectories differed between top height trees in natural and reclaimed stands for jack pine and white spruce, but not for trembling aspen. At a given age, white spruce top height trees were taller and jack pine top height trees were shorter in reclaimed stands than those in natural stands, suggesting that it is easier to achieve similar forest productivity for oil sands sites reclaimed with white spruce stands than for sites reclaimed with jack pine stands. The principal climate variables were growing season (May to September) precipitation averaged over the previous 10 years for trembling aspen and jack pine and summer (June to August) precipitation averaged over the previous 10 years for white spruce. These variables had positive effects on the height–age trajectories.


2005 ◽  
Vol 81 (4) ◽  
pp. 538-547 ◽  
Author(s):  
Douglas G Pitt ◽  
F. Wayne Bell

Stand structure and composition for planted white spruce (Picea glauca (Moench) Voss) and other naturally regenerating commercial species were compared seven years after the testing of five conifer release alternatives on three boreal mixedwood sites. No release resulted in aspen (Populus tremuloides Michx.) -dominated stands with 89% stocking and the highest basal areas (BAs, 5.1 m2/ha) and stem volume indices (SVIs, 10.7 m3/ha) observed. Release by manual or machine cutting increased planted spruce BA and SVI by 67 and 38%, respectively. However, this treatment also caused significant root and stump suckering of aspen, more than doubling stem densities and increasing stocking by 12% over untreated areas. Although cutting reduced the height of aspen from 6 m (untreated) to 2–3 m, equal to or just taller than planted spruce, it is likely that future growth will result in deciduous-dominated mixedwoods. Broadcast foliar application with Release® herbicide temporarily reduced the size of aspen, without causing the increased regeneration observed following cutting. This produced a more varied stand structure that promoted the stature of planted spruce, doubling dominant spruce stocking, BAs, and SVIs, and leading to a more balanced mixedwood. Broadcast release with Vision® herbicide produced conifer-dominated stands with few deciduous stems; these areas contained the lowest observed BAs (1.7 m2/ha) and SVIs (1.9 m3/ha). Relatively low planting densities (1350 sph), coupled with near complete deciduous removal in these plots, created very open-grown conditions that threaten overall productivity and stem quality of the spruce. The five approaches tested are capable of producing a range of stand conditions found in a healthy boreal mixedwood landscape. Key words: boreal mixedwoods, white spruce, trembling aspen, vegetation management, fibre production


2006 ◽  
Vol 36 (9) ◽  
pp. 2331-2340 ◽  
Author(s):  
Suzanne Brais ◽  
David Paré ◽  
Cédric Lierman

To assess nutrient dynamics in decomposing logs of trembling aspen (Populus tremuloides Michx.), white birch (Betula papyrifera Marsh.), white spruce (Picea glauca (Moench) Voss), and jack pine (Pinus banksiana Lamb.), we monitored mass losses and changes in N and P contents in dead boles from a chronosequence of sites following stand-replacing disturbances. To assess the importance of wood decomposition to nutrient cycling, we compared net estimates of nutrient release from logs with net nutrient immobilization in live-tree biomass of stands as a function of time since disturbance. Mineralization rates were 0.060, 0.053, 0.038, and 0.020·year–1 for trembling aspen, white birch, white spruce, and jack pine logs, respectively. Trembling aspen boles released large quantities of N and P during the first year of decomposition (51 kg·ha–1 of N and 7 kg·ha–1 of P, assuming a bole volume of 150 m3·ha–1). White birch boles acted initially as a nutrient sink and delayed the release of immobilized nutrients until a period when the stand's net nutrient immobilization rates were highest. Jack pine boles appeared to be intermediate in terms of their contribution as a sink or a source of nutrients but, in mature stands, provided up to 40% of N and 26% of P immobilized annually in tree biomass. As pure stands of white spruce are rare in boreal Quebec, information on nutrient accumulation in white spruce stands was not available.


2001 ◽  
Vol 79 (1) ◽  
pp. 76-92 ◽  
Author(s):  
Trevor C Lumley ◽  
L Dennis Gignac ◽  
Randolph S Currah

Fallen logs of trembling aspen (Populus tremuloides Michx.) and white spruce (Picea glauca (Moench) Voss) at various stages of decomposition were sampled from undisturbed and 1-, 14-, and 28-year-old post-fire and post-harvest sites in northern Alberta and studied for differences in the associated microfungus communities. Wood samples were plated directly onto each of six different media and, from these, fungal species were identified and enumerated over a 24-month period. Approximately 10 000 isolates were obtained, representing 292 species of filamentous microfungi, including 41 ascomycetes, 29 zygomycetes, and 222 mitosporic fungi. The most commonly isolated species were Trichoderma viride (Gray) Pers., Rhinocladiella atrovirens Nannf., Penicillium pinophilum Hedgcock, and Mortierella ramanniana (Moller) Linnem. Cluster analysis and ordination of microfungus communities in logs showed that the tree species of the log had the greatest influence on the species composition of communities. Fungus community composition was also correlated with the stage of decomposition. Species richness was highest in logs from undisturbed sites and lowest in logs from the most recently disturbed sites. Species diversity (Shannon-Weaver) was only slightly higher at undisturbed sites than at disturbed sites. The most significant environmental factor was log moisture, which increased proportionately with stage of decomposition and was significantly correlated with climatic factors.Key words: ecology, fungi, ordination, succession, species richness, diversity.


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