An unusual ephemeropteran larva from the Lower Cretaceous dysodiles of Lebanon

2019 ◽  
Vol 2 (2) ◽  
pp. 192-198 ◽  
Author(s):  
DANY AZAR ◽  
CLOTILDE DE LA FERTÉ ◽  
LAYLA EL HAJJ ◽  
ANDRÉ NEL ◽  
SIBELLE MAKSOUD
Keyword(s):  
Fore Wing ◽  
Lower Cretaceous ◽  
Hind Wing ◽  
Incertae Sedis

Libanoephemera inopinatabranchia gen. and sp. nov., an ephemeropteran nymph from the Lower Barremian dysodiles of Lebanon, is herein characterized, described, illustrated and assigned to an incertae sedis family. It differs from all known ephemeropteran nymphs and is characterized by hypognathous mouthparts, hind wing pads not covered by fore wing pads, legs with five-segmented tarsi and presence of five pairs of gills on abdominal segments; abdominal segments I and III–V with two-branched gills; segment VI bearing complex gills with six branches.

Description of Caenoephemera shangyuanensis, gen.nov., sp.nov. (Ephemeroptera), from the Yixian Formation

2001 ◽  
Vol 133 (6) ◽  
pp. 747-754 ◽  
Author(s):  
Qi-Bin Lin ◽  
Di-Ying Huang
Keyword(s):  
Fore Wing ◽  
Lower Cretaceous ◽  
Hind Wing ◽  
Upper Jurassic ◽  
Yixian Formation ◽  
Mayfly Nymph ◽  
Tracheal Gills

AbstractA fossil mayfly nymph Caenoephemera shangyuanensis, gen.nov., sp.nov., discovered in the lower part of the Upper Jurassic – Lower Cretaceous Yixian Formation of Jianshangou, Beipiao district, West Liaoning, China, is described. It has several archaic features typical of Paleozoic mayfly nymphs, such as 5 tarsal segments and hind wing pads not covered by fore wing pads, but also exhibits specialized modern features, such as a short metathorax, abdominal tracheal gills, and cerci. It appears to be intermediate in structure between Paleozoic and extant mayfly nymphs.

Gymnosperms from the Lower Cretaceous Crato Formation (Brazil). I. Araucariaceae andLindleycladus (incertae sedis)

Fossil Record ◽  
2004 ◽  
Vol 7 (1) ◽  
pp. 155-174 ◽  
Author(s):  
Lutz Kunzmann ◽  
Barbara A. R. Mohr ◽  
Mary E. C. Bernardes-de-Oliveira
Keyword(s):  
Lower Cretaceous ◽  
Incertae Sedis

scholarly journals Notes on Cretaceous amber Braconidae (Hymenoptera), with descriptions of two new genera

2016 ◽  
pp. 1 ◽  
Author(s):  
Michael S. Engel
Keyword(s):  
Late Cretaceous ◽  
New Genus ◽  
Hind Wing ◽  
New Combination ◽  
New Genera ◽  
New Genus And Species ◽  
Incertae Sedis

A new genus and species of basal cyclostome Braconidae is described and figured from a male preserved in mid-Cretaceous amber from northern Myanmar.  <strong><em>Rhetinorhyssalus morticinus</em></strong> Engel, new genus and species, is interesting for its combination of primitive features such as a minute apical costal cell and anal stubs in the forewing, while lacking 2Cu in the hind wing, a putatively derived trait.  As such, the genus may represent a lineage diverging from the braconid stem subsequent to many protorhyssalines, while remaining basal relative to generalized cyclostome groups such as Rhyssalinae.  In addition, the Late Cretaceous <em>Diospilus allani</em> Brues, in Campanian Canadian amber, is transferred to <strong><em>Diorhyssalus</em></strong> Engel, new genus, and its similarity to <em>Rhetinorhyssalus</em> is discussed.  This transfer results in the new combination, <em>Diorhyssalus allani</em> (Brues).  Both genera are tentatively considered as subfamily <em>incertae sedis</em>.

scholarly journals Completely preserved cockroaches of the family Mesoblattinidae from the Upper Jurassic-Lower Cretaceous Yixian Formation (Liaoning Province, NE China)

2013 ◽  
Vol 64 (4) ◽  
pp. 291-304 ◽  
Author(s):  
Dandan Wei ◽  
Dong Ren
Keyword(s):  
Lower Cretaceous ◽  
Hind Wing ◽  
Upper Jurassic ◽  
Morphological Characters ◽  
Yixian Formation ◽  
Liaoning Province ◽  
Tergal Glands ◽  
The Family ◽  
Left And Right ◽  
First Time

Abstract Although cockroaches were the dominant insects in various Paleozoic and Mesozoic insect assemblages, their general morphology was extremely conservative. One of the most common of them, the Jurassic-Cretaceous family Mesoblattinidae, is described here for the first time on the basis of completely preserved specimens. Ninety-two specimens of Perlucipecta aurea gen. et sp. n. reveal details of head, mandible, male tergal glands and terminal hook; cercal, leg and antennal sensilla. Its congener, P. vrsanskyi is described from the same sediments of the Yixian Formation (Upper Jurassic-Lower Cretaceous). The forewing venation variability of P. aurea, analysed for the first time in this family is nearly identical (CV = 6.23 %) with variability of two species of family Blattulidae that occur at the same locality (CV = 6.22 %; 5.72 %). The transitional nature of morphological characters represented by asymmetry between left and right wings (simple/branched forewing SC and hind wing M) in P. aurea documents the phylogenetic relation between the families Mesoblattinidae and Ectobiidae

New Protosmylinae (Neuroptera: Osmylidae) from the early Eocene of western North America, with taxonomic remarks

Zootaxa ◽  
2021 ◽  
Vol 4980 (1) ◽  
pp. 142-156
Author(s):  
VLADIMIR N. MAKARKIN ◽  
S. BRUCE ARCHIBALD ◽  
ROLF W. MATHEWES
Keyword(s):  
North America ◽  
Hind Wing ◽  
Late Eocene ◽  
Early Eocene ◽  
Western North America ◽  
Green River ◽  
Incertae Sedis ◽  
Three New Species ◽  
Okanagan Highlands ◽  
Provincial Park

The osmylid subfamily Protosmylinae is revised based on our emended diagnosis: Petrushevskia Martynova, 1958 and Mesosmylidus Jepson et al., 2012 are excluded (both considered Osmylidae incertae sedis), and Sogjuta Martynova, 1958 is transferred to it from the Mesosmylininae. The late Eocene genus Protosmylus Krüger, 1913 is considered a junior synonym of Osmylidia Cockerell, 1908 based on a distinct apomorphy (deeply forked MA in the hind wing), syn. nov. Three new species of Osmylidia from the early Eocene of North America are described: O. donnae sp. nov. from Quilchena, O. glastrai sp. nov. from Republic, Washington, USA, and an unnamed species of Osmylidia is reported from Driftwood Canyon Provincial Park, British Columbia (all localities of the Okanagan Highlands series), and O. taliae sp. nov. from the Green River Formation of Colorado, USA. 

scholarly journals Morphology-based phylogenetic analysis of the treehopper tribe Smiliini (Hemiptera: Membracidae: Smiliinae), with reinstatement of the tribe Telamonini

Zootaxa ◽  
2011 ◽  
Vol 3047 (1) ◽  
pp. 1 ◽  
Author(s):  
MATTHEW S. WALLACE
Keyword(s):  
Phylogenetic Analysis ◽  
Common Ancestor ◽  
Hind Wing ◽  
Morphological Characters ◽  
Dorsal Margin ◽  
Incertae Sedis ◽  
Plant Associations ◽  
Original Dataset ◽  
The Common ◽  
Morphological Differences

Members of the Smiliini, the nominotypical tribe of the large New World subfamily Smiliinae, are predominately Nearctic in distribution. This tribe included 169 mostly tree-feeding species in 23 genera. A parsimony-based phylogenetic analysis of an original dataset comprising 89 traditional and newly discovered morphological characters for 69 species, including representatives of 22 of the 23 described genera of Smiliini and five other previously recognized tribes of the subfamily, resulted in a single most parsimonious tree with three major clades. The broad recent concept of Smiliini (including Telamonini as a junior synonym) was not recovered as monophyletic by the analysis. Instead, the analysis supported narrower definitions of both Telamonini, here reinstated from synonymy, and Smiliini. A key and diagnoses are given to define these tribes, along with discussions of their phylogeny, biogeography, and host plant associations. The genera Antianthe Fowler, Hemicardiacus Plummer, Smilirhexia McKamey, and Tropidarnis Fowler are placed as Smiliinae, incertae sedis. Based on the phylogeny, several genera from both tribes including Atymna Stål, Cyrtolobus Goding, Heliria Stål, and Telamona Fitch are not monophyletic. Diagnostic characters emphasizing the morphological differences between the Smiliini and Telamonini include the dorsal margin of the head, the shape of the pronotum, the size of the pronotal humeral angles, the presence or absence of pronotal longitudinal rugae, the size of forewing cells, variations in the fusion of veins R and M apically in both the foreand hind wing, and the shape of the apex of the female second valvulae. Mapping geographic distribution onto the phylogeny suggests that the common ancestor of the ingroup (all three clades) occurred in Central America and Mexico, with multiple dispersals to temperate North America. Many Smiliini and Telamonini feed on various species of oak (Quercus) and the close evolutionary association between these insects and their hosts is discussed.

The second genus of the ‘libelluloid’ family Araripephlebiidae (Odonata, Clavilabiata)

2020 ◽  
Vol 3 (3) ◽  
pp. 240-244
Author(s):  
ANDRÉ NEL ◽  
JEAN-MARC POUILLON
Keyword(s):  
Lower Cretaceous ◽  
Hind Wing ◽  
Longitudinal Vein ◽  
Anal Area

The Lower Cretaceous Crato Formation is a well-known Konservat Lagerstätte with a very rich entomofauna. The Odonata are especially very diverse and extensively studied (Bechly, 1998, 2000, 2007, 2010; Nel et al., 1998; Bechly et al., 2001; Bechly & Ueda, 2002) with representatives of all the extant anisopteran main subgroups. This fauna is especially interesting because it comprises some of the oldest and most ‘basal’ groups of the highly diverse extant ‘libelluloid’ dragonflies, or Clavilabiata Bechly, 1996. Among these, the monospecific family Araripephlebiidae Bechly, 1998 is remarkable in the highly specialized hind wing cubito-anal area that contains a curious supplementary longitudinal vein more or less parallel to AA and CuA, unique among the Odonata. Nevertheless, this family remained rather poorly known by three described specimens, only females.

EVIDENCE FOR MONOPHYLY AND RELATIONSHIPS OF CHALCIDOIDEA, MYMARIDAE, AND MYMAROMMATIDAE (HYMENOPTERA: TEREBRANTES)

1986 ◽  
Vol 118 (3) ◽  
pp. 205-240 ◽  
Author(s):  
Gary A.P. Gibson
Keyword(s):  
Fore Wing ◽  
Hind Wing ◽  
Sister Group ◽  
Character State ◽  
Family Status ◽  
Lateral Edge ◽  
The Family ◽  
State Evolution ◽  
Internal Character ◽  
Relationship Of

AbstractTwenty-three characters or character systems of adults and larvae of Terebrantes are analyzed for evidence of monophyly and phyletic relationships of Chalcidoidea, Mymaridae, and Mymarommatidae. The taxa are considered to be a monophyletic group based on 3 hypothesized synapomorphies: mesotrochanteral depressor without fu2-tr2 or mesoscutal portion of t2-tr2; axillar phragma as site of origin for all or part of t,-tr2 muscle; and independent basal ring absent from male genitalia. The family Mymaridae is considered to be monophyletic based on at least 3 apomorphies: fore wing with hypochaeta; head with frontal, median, and supraorbital sulci; and toruli distinctly closer to inner margin of eye than to each other. Chalcidoidea, including Mymaridae, is considered to be a monophyletic taxon based on 3 apomorphies: prepectus externally visible, at least dorsally adjacent to lateral edge of mesoscutum; mesothoracic spiracle positioned at exposed lateral edge of mesoscutum; and multiporous plate sensilla of antenna with unique structure, as described in text. Mymarommatidae is considered to be the monophyletic sister group of Chalcidoidea based on several apomorphies, including 4 autapomorphies: head composed of frontal and occipital sclerites, which are connected by pleated membrane along hyperoccipital region; hind wing stalk-like, without membrane and terminated in bifurcation that clasps fore wing; fore wing with reticulate pattern formed by raised lineations of membrane; and axillar portion of t2-tr2 muscle absent. Phyletic relationship of Serphitidae with Mymarommatidae is deemed inconclusive because relevant internal character states of amber fossil serphitids cannot be determined. It is suggested that mymarommatids be accorded family status, but not be assigned to superfamily until phyletic relationships are more accurately determined in Terebrantes. A matrix summarizes character-state distribution of most characters analyzed for Terebrantes, and a cladogram illustrates hypotheses of character-state evolution and proposed relationships.

scholarly journals Morphological characterization and wing description of Vespa orientalis orientalis queens

2017 ◽  
Vol 33 (2) ◽  
pp. 251-259
Author(s):  
Hossam Abou-Shaara
Keyword(s):  
Fore Wing ◽  
Wing Length ◽  
Hind Wing ◽  
Morphological Characteristics ◽  
Morphological Characterization ◽  
Head Width ◽  
Vespa Orientalis ◽  
Computer Based ◽  
Bee Colonies ◽  
Tibia Length

Oriental hornets, Vespa orientalis, are dangerous enemy to bee colonies in some countries of the world. There are more than one subspecies of V. orientalis. Few studies have investigated the morphological characteristics of these subspecies. Morphological characterization can help in confirming and discriminating between the subspecies, and to follow any changes in their morphology over time. In this study, some body characteristics of V. orientalis orientalis queens from Egypt were measured including head width, fore wing length and width, hind wing length and width, femur length, tibia length and approximate stinger length. Also, fore wing characteristics using wing coordinates for 20 landmarks were studied. Computer based techniques were applied to take these measurements. The data of the current study can be utilized for comparisons with other subspecies.

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