Phylogeny of the weevil genus Rhinusa Stephens based on adult morphological characters and host plant information (Coleoptera: Curculionidae)

Zootaxa ◽  
2010 ◽  
Vol 2627 (1) ◽  
pp. 39 ◽  
Author(s):  
ROBERTO CALDARA ◽  
DAVIDE SASSI ◽  
IVO TOŠEVSKI
Keyword(s):  
Host Plant ◽  
Feeding Habits ◽  
Cladistic Analysis ◽  
Morphological Characters ◽  
Closely Related Species ◽  
Weak Support ◽  
Plant Associations ◽  
The Family ◽  
Related Plant ◽  
Species Groups

A phylogenetic analysis of the species belonging to the weevil genus Rhinusa Stephens, 1829 (Coleoptera: Curculionidae: Curculioninae: Mecinini) was carried out. Rhinusa weevils feed on plants of the closely related families Scrophulariaceae and Plantaginaceae. Based on a cladistic analysis of six outgroup and 33 ingroup taxa, and 39 adult morphological and 8 ecological characters, eight well supported species groups and two monobasic groups belonging to three separate and more inclusive assemblages were recognized. The first assemblage (A) includes nine species belonging to two groups (R. bipustulata and R. tetra groups), whereas the second and third assemblages (B and C) include a total of 14 species belonging to two groups (R. antirrhini and R. linariae groups) and six groups (R. pilosa, R. herbarum, R. neta, R. vestita, R. mauritii and R. melas groups), respectively. Two of the three main assemblages (A and B) are well supported as monophyletic entities, whereas the third assemblage (C) has weak support contingent in part upon the exclusion of host plant associations. Assemblage A includes all species living on species of Scrophulariaceae, with two groups occurring on two closely related plant genera, Scrophularia (R. bipustulata group) and Verbascum (R. tetra group), respectively. The other two assemblages include species living exclusively on species of the family Plantaginaceae, tribe Antirrhineae. These patterns suggest a well conserved and phylogenetically congruent association among the weevils and their hosts. Optimizing host plant preferences onto the morphological phylogeny indicates that feeding on Plantaginaceae was the plesiomorphic condition for the genus Rhinusa. In general there are no strict relationships between groups of weevils and their specific feeding habits; however, species of the R. antirrhini group are all feeding on seed capsules. In contrast, in other groups the larvae of closely related species display significant variations in host plant parasitism. Some species of the R. tetra group feed on seed capsules whereas others are stem borers. In turn, certain species of the R. neta group feed on seed capsules yet others are inquilines of gall forming species of Rhinusa. The latter habit is present in multiple convergent groups such as the R. linariae and R. pilosa groups.

The morphology, higher-level phylogeny and classification of the Empidoidea (Diptera)

Zootaxa ◽  
2006 ◽  
Vol 1180 (1) ◽  
pp. 1 ◽  
Author(s):  
BRADLEY J. SINCLAIR ◽  
JEFFREY M. CUMMING
Keyword(s):  
Feeding Habits ◽  
Cladistic Analysis ◽  
Sister Group ◽  
Morphological Characters ◽  
Male And Female ◽  
Ground Plan ◽  
Incertae Sedis ◽  
Higher Taxa ◽  
The Family

A cladistic analysis of the Empidoidea and basal lineages of the Cyclorrhapha, based on morphological characters, confirms the monophyly of both groups as well as that of the                    Eremoneura. The resulting final trees are used to revise the classification of the Empidoidea to include the following five families: Empididae, Hybotidae, Atelestidae (including Nemedininae n. subfam.), Brachystomatidae rev. stat. (comprising the subfamilies Brachystomatinae, Ceratomerinae and Trichopezinae), and Dolichopodidae s.lat. The family Microphoridae is not recognized, and the Microphorinae and Parathalassiinae are assigned to the Dolichopodidae s.lat. The Dolichopodidae s.str. includes 15 subfamilies that were previously recognized within the family. Within the Empidoidea we found support for Atelestidae as the sister group to the Hybotidae and for the monophyly of Parathalassiinae + Dolichopodidae s.str. The Empididae remains poorly defined and the genera Homalocnemis Philippi, Iteaphila Zetterstedt, Anthepiscopus Becker, and Oreogeton Schiner are classified as incertae sedis within the                   Empidoidea. In addition, the following higher taxa are proposed: Symballophthalmini n. tribe, Bicellariini n. tribe, Oedaleinae rev. stat., and Trichininae rev. stat., which are all assigned to the Hybotidae. The genus Sematopoda Collin is tentatively assigned to Trichopezinae, and Xanthodromia Saigusa is transferred from Hemerodromiinae to Brachystomatinae.        All morphological characters are extensively discussed and illustrated, including details of the antennae, mouthparts, internal thoracic structures, wings, and male and female terminalia. In addition, a key to families and unplaced genus groups of the Empidoidea is provided. Feeding habits are also discussed in terms of the empidoid ground plan condition.

THE NEPTICULIDAE (LEPIDOPTERA) OF CANADA

1979 ◽  
Vol 111 (S107) ◽  
pp. 1-129 ◽  
Author(s):  
Christopher Wilkinson ◽  
M. J. Scoble
Keyword(s):  
Host Plant ◽  
A Priori ◽  
Relevant Literature ◽  
Taxonomic Revision ◽  
Diagnostic Features ◽  
The North ◽  
Plant Associations ◽  
The Family ◽  
Species Groups ◽  
Plant Choice

AbstractA taxonomic revision, including descriptions, diagnoses, and bibliographies, is given of the genera and species of the family Nepticulidae (Lepidoptera) so far collected in Canada. In addition, extensive material from the U.S.A. and a number of specimens from Europe have been examined. Five new species and two new subspecies are described. Notes on the biology of the species have been collated from rearing records, label data, and the relevant literature. With one exception all existing types have been examined and, in addition, a number of types designated.A major feature of this work is a study of the genitalia of the moths — never before undertaken extensively on the North American Nepticulidae. These structures give valuable diagnostic features and are of fundamental importance in the provision of modern descriptions and keys. As genitalia characters are thought to be the best taxonomic indicators of relationships between species in the Nepticulidae, the species-groups proposed have been mainly defined on these structures.The confusion over the nomenclature of the family name is discussed. Taxonomic histories of the genera are given and one generic synonymy is made.Taxonomic problems are discussed in relation to distribution and food-plant choice. The host-plant associations of the family are reviewed, with reference to the mining habits of the larvae. A priori use of host-plant data in the taxonomy of the Nepticulidae is found to be limited. However, the proposed species-groups illustrate a general conclusion that related species tend to feed on plants that are themselves taxonomically related.

A phylogeny of the family Fanniidae Schnabl (Insecta:Diptera:Calyptratae) based on adult morphological characters, with special reference to the Austral species of the genus Fannia

2008 ◽  
Vol 22 (5) ◽  
pp. 563 ◽  
Author(s):  
M. C. Domínguez ◽  
S. A. Roig-Juñent
Keyword(s):  
Cladistic Analysis ◽  
Morphological Characters ◽  
External Morphology ◽  
Phylogenetic Position ◽  
Phylogenetic Hypothesis ◽  
Parsimony Analysis ◽  
Male And Female ◽  
Southern South America ◽  
The Family ◽  
Species Groups

The present study proposed a phylogenetic hypothesis of the family Fanniidae based on a cladistic analysis using characters from adult external morphology and female and male terminalia. The main purpose of this study was to clarify the phylogenetic position of newly described or poorly known species, mostly from southern South America, the Neotropics, Africa, Australia and New Zealand. In total, 151 characters from adult male and female external morphology and terminalia were scored for 78 species of Fanniidae. Ten continuous characters were included and analysed as such. Three genera of Fanniidae and all the species-groups and subgroups proposed for the genus Fannia, except for the admirabilis-group and the setifer-subgroup were included as terminal taxa. An heuristic parsimony analysis under implied weights was performed. The analysis recovered the monophyly of the Fanniidae and the genus Fannia, as well as the monophyly of several species-groups within Fannia. Male and female external morphological characters were, in general, highly homoplasious, whereas characters from male terminalia showed low level of homoplasy and provided resolution at suprageneric nodes and species-groups.

Supraspecific classification.

2022 ◽  
pp. 11-12
Author(s):  
Richard A. I. Drew ◽  
Meredith C. Romig
Keyword(s):  
Host Plant ◽  
Morphological Characters ◽  
Plant Biology ◽  
Evolutionary Origins ◽  
Species Groups

Abstract The definitions of genera and subgenera used in the classification of the Dacini have been in a continual state of change for over a century. The early definitions were based on often homoplasious morphological characters, some examples for the Oriental and Australian regions being Tryon (1927), Perkins (1937), Hardy (1951), May (1951) and Drew (1972). More recently revised subgeneric definitions for most species groups were published by Drew and Hancock (2016) and Hancock and Drew (2006, 2015, 2016, 2017a,b,c,d,e, 2018a,b,c, 2019), based on detailed analyses of dacine biogeography, host plant biology and morphology. This chapter discusses the evolutionary origins of the Dacini, the host plant and its influence on speciation in the Dacini.

Systematics of reptile-associated scale mites of the genus Pterygosoma (Acariformes: Pterygosomatidae) derived from external morphology

Zootaxa ◽  
2019 ◽  
Vol 4603 (3) ◽  
pp. 401
Author(s):  
MONIKA FAJFER
Keyword(s):  
North Africa ◽  
New Genus ◽  
Arabian Peninsula ◽  
Morphological Characters ◽  
Mite Species ◽  
South American ◽  
The Family ◽  
Outgroup Species ◽  
Tree Lizards ◽  
Species Groups

The phylogeny of the mite genus Pterygosoma Peters, 1849 (Acariformes: Pterygosomatidae), permanent parasites of lizards of the families Agamidae, Gerrhosauridae and Liolaemidae (Reptilia: Squamata), was inferred with maximum parsimony, successive weighting and implied weighting approaches based on 182 characters of 48 ingroup and 12 outgroup species. All undertaken analyses questioned the monophyly of the genus. Based on this research, Pterygosoma includes 56 mite species associated with agamas from Africa and Asia (Sauria: Agamidae). Within the genus seven natural species groups were found: mutabilis, inermis, melanum, caucasica, fimbriata, singularis and foliosetis; 13 species of the genus were not assigned to any of the groups due to their unique morphological characters. For the ligare group of the subgenus Pterygosoma s. str. (7 species) associated with the South American tree lizards from the family Liolaemidae, a new genus, Neopterygosoma gen. nov., is established. The subgenus Gerrhosaurobia Lawrence, 1959 (3 species) associated with the African plated lizards of the family Gerrhosauridae is elevated to the genus rank. Diagnoses for the three genera are presented. Historical associations are reconstructed by fitting the obtained mite tree into the host topology on the family level. Results of coevolutionary analysis highlight the coincidence of both trees. The studies suggest that the ancestor of the genus Pterygosoma switched from the hosts of Gerrhosaurobia i.e. from the plated lizards (Gerrhosauridae) to the agamas (Agamidae), and the genus Pterygosoma has its biogeographic origin in North Africa, and colonized Asia via the Arabian Peninsula, which is congruent with the historical biogeography of its hosts.

Morphotaxonomic revision of Simulium (Gomphostilbia) (Diptera: Simuliidae) in the Oriental Region

Zootaxa ◽  
2012 ◽  
Vol 3577 (1) ◽  
pp. 1 ◽  
Author(s):  
HIROYUKI TAKAOKA
Keyword(s):  
Geographical Distribution ◽  
Classification Scheme ◽  
Cladistic Analysis ◽  
Morphological Characters ◽  
Species Group ◽  
Morphological Identification ◽  
Oriental Region ◽  
Preliminary Attempt ◽  
The Third ◽  
Species Groups

Simulium (Gomphostilbia) Enderlein, the third largest (206 named species included) in the genus Simulium Latreille s. l., is one of the two most abundant and diverse subgenera in the Oriental Region. To provide a classification scheme to facilitate morphological identification of the species within this subgenus, its diagnostic characters are redefined, and nine known species-groups within it are reviewed. Based on putative lineages explored by using certain adult morphological characters, seven more species-groups are proposed: asakoae, darjeelingense, epistum, gombakense, heldsbachense, hemicyclium and palauense species-groups, while the trirugosum species-group is merged in the varicorne species-group. Subgroups are also introduced to represent apparently different lineages within certain species-groups based on certain pupal morphological characters: two in the banauense species-group, seven in the batoense species-group redefined, four in the ceylonicum species-group redefined, four in the epistum species-group, two in the hemicyclium species-group, two in the sherwoodi species-group and four in the varicorne species-group redefined. A new checklist of species of the subgenus Gomphostilbia, and a key to all 15 species-groups within it are provided. The eastward expansion of the geographical distribution of the subgenus Gomphostilbia is inferred on the basis of the more frequent occurrence of apomorphic characters of certain adult and pupal morphological features in insular species-groups than in continental species-groups. A preliminary attempt using a cladistic analysis of morphological characters shows that among 10 subgenera examined, Gomphostilbia has a sister-taxon relationship with the Australasian subgenus Morops Enderlein, and this clade, together with the Central-Western Pacific subgenus Inseliellum Rubtsov, is positioned closest to the most derived clade formed by Daviesellum Takaoka & Adler and Simulium Latreille s. str.

Revision, phylogeny and natural history of Cotithene Voss (Coleoptera: Curculionidae)

Zootaxa ◽  
2008 ◽  
Vol 1782 (1) ◽  
pp. 1 ◽  
Author(s):  
NICO M. FRANZ
Keyword(s):  
Costa Rica ◽  
Natural History ◽  
Morphological Traits ◽  
Anterior Margin ◽  
Narrow Range ◽  
Cladistic Analysis ◽  
Morphological Characters ◽  
Closely Related Species ◽  
Host Shifts ◽  
History Of

Cotithene Voss, a previously monotypic genus of Neotropical derelomine flower weevils (Curculionidae: Derelomini), is revised, with provision of a key to the species, cladistic analysis and notes on its natural history. The following six new species are described: C. anaphalanta (Costa Rica), C. dicranopygia (Costa Rica), C. leptorhamphis (Costa Rica, Panama), C. melanoptera (Venezuela), C. stratiotricha (Costa Rica) and C. trigaea (Costa Rica). The monophyly of Cotithene is supported by the characters of a dorsomedially expanded, carinate rostrum, ventrally angulate head, long and anteriorly directed setation on the anterior margin of the prosternum and an apicodorsally expanded aedeagus with paired sclerites in the male, and subcontiguous to separated procoxal cavities in the female. Particularly the males of several species have intriguing and allometrically scaled modifications on the head (triangular projections, long setae) and pronotum (expansion, tumescences), which possibly play a role in male-to-male conflicts. Cotithene species are specialized to visit and reproduce on a narrow range of typically closely related species of Cyclanthaceae. The adults do not function as pollinators, and the herbivorous larvae develop in the fruiting organs of their hosts, frequently triggering the abortion of infructescences. An analysis of 12 taxa (5 outgroup, 7 ingroup) and 32 morphological characters yielded a single most parsimonious cladogram (L = 38, CI = 89, RI = 93) with the topology (C. dicranopygia, (C. stratiotricha, ((C. leptorhamphis, C. trigaea), (C. globulicollis Voss, (C. anaphalanta, C. melanoptera))))). The evolution of morphological traits and host shifts is examined in light of the proposed phylogeny.

Pedrowygomyia, a new Neotropical genus of Prosimuliini (Diptera: Simuliidae): Gigantodax s. lat. split into two genera

1998 ◽  
Vol 29 (2) ◽  
pp. 161-167 ◽  
Author(s):  
Daniel Rafael Miranda-Esquivel ◽  
Sixto Coscarón
Keyword(s):  
New Genus ◽  
Cladistic Analysis ◽  
Morphological Characters ◽  
Species Group ◽  
New Combinations ◽  
Species Groups

AbstractA cladistic analysis of the species groups recognized in Gigantodax Enderlein, using morphological characters, shows that Gigantodax s. lat. is paraphyletic. From a phylogenetic standing Gigantodax must be split into two monophyletic entities: Gigantodax s. str. and the G. cortesi species group. The new genus Pedrowygomyia is proposed for the G. cortesi species group. It presently contains four species, P. cortesi, P. jatunchuspi, P. punapi and P. chacabamba (all new combinations).

Corrigendum to: Systematics and distribution of world hyptiogastrine wasps (Hymenoptera : Gasteruptiidae)

2002 ◽  
Vol 16 (6) ◽  
pp. 957 ◽  
Author(s):  
J. T. Jennings ◽  
A. D. Austin
Keyword(s):  
New Species ◽  
New Zealand ◽  
South America ◽  
New Caledonia ◽  
Cladistic Analysis ◽  
Taxonomic Revision ◽  
Morphological Characters ◽  
The Family ◽  
New Hebrides ◽  
New Britain

This study examines the phylogeny, taxonomy, distribution and biology of the gasteruptiid subfamily Hyptiogastrinae and, at the same time, presents an overview of the family. Following a cladistic analysis of 35 discrete morphological characters, two monophyletic genera are recognised, Hyptiogaster Kieffer and Pseudofoenus s. l. Kieffer. As a consequence, the genera Aulacofoenus Kieffer, Crassifoenus Crosskey, and Eufoenus Szépligeti are synonymised with Pseudofoenus. A total of 88 species are recognised for the subfamily, 10 species of Hyptiogaster, which are restricted to mainland Australia, and 78 species of Pseudofoenus, 40 of which are described as new. Pseudofoenus has a restricted Gondwanan distribution and is found in Australia including Tasmania (65 spp.), New Guinea and New Britain (5 spp.), the south-west Pacific (New Caledonia, New Hebrides and Fiji – 2 spp.), New Zealand (4 spp.) and South America (2 spp.). No new species have been recorded from either New Zealand or South America. For Pseudofoenus, information on the distribution of each species, their biology (if known) and an identification key are presented.Following a taxonomic revision, the following new species are described: P. baileyi, sp. nov., P. baitetaensis, sp. nov., P. beverlyae, sp. nov., P. caperatus, sp. nov., P. cardaleae, sp. nov., P. carrabinensis, sp. nov., P. claireae, sp. nov., P. collessi, sp. nov., P. coorowensis, sp. nov., P. crosskeyi, sp. nov., P. douglasorum, sp. nov., P. eliseae, sp. nov., P. ericae, sp. nov., P. eustonensis, sp. nov., P. feckneri, sp. nov., P. gressitti, sp. nov., P. gullanae, sp. nov., P. hackeri, sp. nov., P. imbricatus, sp. nov., P. iqbali, sp. nov., P. kadowi, sp. nov., P. karimuiensis, sp. nov., P. kelleri, sp. nov., P. leinsterensis, sp. nov., P. macdonaldi, sp. nov., P. malkini, sp. nov., P. marshalli, sp. nov., P. masneri, sp. nov., P. mitchellae, sp. nov., P. morganensis, sp. nov., P. nalbarraensis, sp. nov., P. pumilis, sp. nov., P. schmidti, sp. nov., P. stevensi, sp. nov., P. tasmaniensis, sp. nov., P. taylori, sp. nov., P. umboiensis, sp. nov., P. walkeri, sp. nov. and P. zborowskii, sp. nov. The synonymy of Aulacofoenus, Crassifoenus and Eufoenus with Pseudofoenus result in the following new combinations: from Aulacofoenus: P. bungeyi (Jennings & Austin), comb. nov., P. deletangi (Schletterer), comb. nov., P. fallax (Schletterer), comb. nov., P. fletcheri (Jennings & Austin), comb. nov., P. goonooensis (Jennings & Austin), comb. nov., P. infumatus (Schletterer), comb. nov., P. kurmondi (Jennings & Austin), comb. nov., P. loxleyi (Jennings & Austin), comb. nov., P. marionae (Jennings & Austin), comb. nov., P. perenjorii (Jennings & Austin), comb. nov., P. swani (Jennings & Austin), comb. nov., P. thoracicus (Guérin Menéville), comb. nov., P. whiani (Jennings & Austin), comb. nov. and P. wubinensis (Jennings & Austin), comb. nov.; from Crassifoenus: P. houstoni (Jennings & Austin), comb. nov., P. grossitarsis (Kieffer), comb. nov and P. macronyx (Schletterer), comb. nov.; and from Eufoenus: P. antennalis (Schletterer), comb. nov., P. australis (Westwood), comb. nov., P. crassitarsis (Kieffer), comb. nov., P. darwini (Westwood), comb. nov., P. extraneus (Turner), comb. nov., P. ferrugineus (Crosskey), comb. nov., P. floricolus (Turner), comb. nov., P. inaequalis (Turner), comb. nov., P. melanopleurus (Crosskey), comb. nov., P. minimus (Turner), comb. nov., P. nitidiusculus (Turner), comb. nov., P. patellatus (Westwood), comb. nov., P. pilosus (Kieffer), comb. nov., P. reticulatus (Crosskey), comb. nov., P. rieki (Crosskey), comb. nov., P. ritae (Cheesman), comb. nov. and P. spinitarsis (Westwood), comb. nov. Pseudofoenus microcephalus (Crosskey), comb. nov. is transferred from Hyptiogaster and Eufoenus flavinervis (Kieffer) remains incertae sedis.

scholarly journals Phylogenetic relationships within the Pylochelidae (Decapoda: Anomura: Paguroidea): A cladistic analysis based on morphological characters

Zootaxa ◽  
2009 ◽  
Vol 2022 (1) ◽  
pp. 1-14 ◽  
Author(s):  
RAFAEL LEMAITRE ◽  
PATSY A. MCLAUGHLIN ◽  
ULF SORHANNUS
Keyword(s):  
Hermit Crab ◽  
Phylogenetic Relationships ◽  
Cladistic Analysis ◽  
Morphological Characters ◽  
Data Matrix ◽  
Parsimony Analysis ◽  
Consensus Tree ◽  
Strict Consensus Tree ◽  
The Family

Phylogenetic relationships within the “symmetrical” hermit crab family Pylochelidae were analyzed for 41 of the 45 species and subspecies currently considered valid. In the analyses, 78 morphological characters comprised the data matrix and the outgroup consisted of Thalassina anomala, a member of the Thalassinidae, and Munida quadrispina, a member of the Galatheidae. A poorly resolved strict consensus tree was obtained from a heuristic parsimony analysis of unweighted and unordered characters, which showed the family Pylochelidae and the subfamilies Pylochelinae and Pomatochelinae to be monophyletic taxa – the latter two groups had the highest Bremer support values. Additionally, while the subgenus Pylocheles (Pylocheles) was strongly supported, the subgenera Xylocheles, and Bathycheles were not. More fully resolved trees were obtained when using implied weighting, which recognized the monotypic subfamilies Parapylochelinae, Cancellochelinae and Mixtopagurinae. The subfamily Trizochelinae was found to have four distinct clades and several ambiguously placed taxa.

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