Phylogenetic analysis of Lachesilla Westwood (Psocodea: Psocomorpha: Lachesillidae) and relationships of sister genera of the subfamily Lachesillinae

Lachesilla is the most species-rich psocid genus, with 341 species grouped into 19 species groups (with at least 100 undescribed species), plus an additional undescribed species group. We present the first phylogenetic analysis of Lachesilla and a discussion on the relationships of sister genera of the subfamily Lachesillinae. We codified a matrix of 60 morphological characters (16 multistate and 44 binary) formed by 44 terminals: 37 ingroup species and seven outgroup species. A phylogenetic analysis was performed using Maximum Parsimony (MP) as optimality criterion, under equal weights and implied weights approaches. Monophyly of Lachesilla is not recovered, showing a polyphyletic genus divided in two major clades: clade A, composed of the pedicularia species group + Nadleria, and clade B, including the remaining species groups. The implied weights analysis produced a hypothesis with Hemicaecilius as sister group of clade B, nesting seven subclades mainly with low branch support values (symmetric resampling and bootstrap). These results suggest that all species groups, included in this analysis, can be considered monophyletic clades based on several synapomorphies, despite the andra species group was split up showing that L. punctata + L. dentata cluster together with high branch support values, but L. tehuautlensis and L. falcicula were recovered in different subclades.

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New taxa of the Lygistorrhinidae (Diptera: Sciaroidea) and their implications for a phylogenetic analysis of the family

Zootaxa ◽

10.11646/zootaxa.960.1.1 ◽

2005 ◽

Vol 960 (1) ◽

pp. 1 ◽

Cited By ~ 8

Author(s):

HEIKKI HIPPA ◽

INGEGERD MATTSSON ◽

PEKKA VILKAMAA

Keyword(s):

Phylogenetic Analysis ◽

New Taxa ◽

Cladistic Analysis ◽

Sister Group ◽

Morphological Characters ◽

Undescribed Species ◽

Parsimony Analysis ◽

The Family ◽

Outgroup Species ◽

New Material

New Oriental taxa of the Lygistorrhinidae - Blagorrhina gen. n., with B. blagoderovi sp. n. and B. brevicornis sp. n.; Gracilorrhina gracilis gen. n., sp. n.; and Labellorrhina gen. n., with L. grimaldii sp. n. and L. quantula sp. n. are described, and two undescribed species, known only from females, are characterized. Based on this new material, the family is redefined. The phylogeneticrelationships among the taxa of Lygistorrhinidae were studied by parsimony analysis using 43 morphological characters from the adults of 25 ingroup and one outgroup species. The cladistic analysis produced 14 most parsimonious cladograms. The solution obtained suggests unambiguously the following phylogeny: Palaeognoriste Meunier and “Lygistorrhina” asiatica Senior-White are successively sister groups of the rest of the Lygistorrhinidae; there is a clade Labellorrhina + (Gracil- orrhina + (Blagorrhina + ((Seguyola Matile + (Loyugesa Grimaldi & Blagoderov + Matileola Papp))))) with a monophyletic Lygistorrhina Skuse – Probolaeus Williston lineage as sister group. The phylogeny among the latter group remains largely unresolved.

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The phylogeny and classification of the tribe Halictini, with special reference to the Halictus genus-group (Hymenoptera: Halictidae)

Zoosystematica Rossica ◽

10.31610/zsr/2004.13.1.83 ◽

2004 ◽

Vol 13 (1) ◽

pp. 83-113

Author(s):

Yu.A. Pesenko

Keyword(s):

Phylogenetic Analysis ◽

Male Genitalia ◽

Posterior Margin ◽

Sister Group ◽

Morphological Characters ◽

Species Group ◽

Transverse Ridge ◽

Inner Surface ◽

Paraphyletic Group

The monophyly of the cosmopolitan tribe Halictini, including over 2300 currently recognized species, is supported by at least a single manifested synapomorphy shared by all members of the tribe: metasomal tergum VII of the male is modified; this forms a transverse ridge giving a false apex beneath which the tergum is strongly reflexed to the morphological posterior margin. On the basis of the present phylogenetic analysis, the tribe Halictini is subdivided into five subtribes: Halictina (comprised of 7 genera: Echthralictus, Glossodialictus, Halictus, Homalictus, Patellapis, Seladonia, and Thrincohalictus), Sphecodina (4 genera: Eupetersia, Microsphecodes, Ptilocleptis, and Sphecodes), Thrinchostomina (2 genera: Thrinchostoma and Parathrincostoma), Caenohalictina (9 genera: Agapostemon, Caenohalictus, Dinagapostemon, Habralictus, Mexalictus, Paragapostemon, Pseudagapostemon, Rhinetula, and Ruizantheda), and Gastrohalictina (one large and diverse genus: Lasioglossum s. l.). The subtribe Halictina is a paraphyletic group; the remaining four tribes are strictly monophyletic (holophyletic). The monophyly of the Halictus genus-group, comprising the genera Halictus and Seladonia, is supported by two distinct synapomorphies of the male genitalia: (1) dorsal gonostylus simple (not double), flattened, broad, narrowed proximally, and provided with a clump of very coarse bristles on the inner surface; (2) ventral gonostylus sclerotized, relatively thin and long, directed backward. Only generalized members of the group possess both the character states above. In derived members, the clump of bristles and the ventral gonostylus are often lost independently. The sister group of the Halictus genus-group is the genus Thrincohalictus. This is supported by the following synapomorphy found among the tribe Halictini only in Halictus, Seladonia, and Thrincohalictus: the ventral gonobasal rim in the male genitalia is forming a right posterolateral angle with a short projection directed laterally. Another feature characterizing these three genera is the presence (except in the parasitic subgenus Paraseladonia) of posterior bands of tomentum or dense and much plumose appressed hairs on the metasomal terga. However, this character is shared also with Patellapis subgenera Patellapis and Lomatalictus. The following subgeneric classification of the genera Halictus and Seladonia is suggested. The genus Halictus includes 12 subgenera: Acalcaripes (2 species), Argalictus (8), Halictus s. str. (4), Hexataenites (11), Lampralictus (1), Monilapis (29), Nealictus (2), Odontalictus (2), Platyhalictus (14), Protohalictus (13), Ramalictus (2), and Tytthalictus (4). The genus Seladonia comprises 6 subgenera, including 2 new ones: Mucoreohalictus subg. n. (15), Pachyceble (22), Paraseladonia (1), Placidohalictus subg. n. (5), Seladonia s. str. (36), and Vestitohalictus (16). The subgenera of Halictus and Seladonia are keyed. The phylogenetic tree of the subgenera of Halictus and Seladonia is reconstructed with use of 46 morphological characters of adults. All other genera of the tribe Halictini were taken as outgroup. The genus Halictus is ascertained as a strictly monophyletic group based upon a single postulated synapomorphy: dorsal gonostylus with a triangular hair patch as a distal appendage on inner side. All the subgenera of this genus appear as strictly monophyletic groups, with the exception of Monilapis, which is a paraphyletic one in relation to Acalcaripes. The monophyly of the genus Seladonia is supported by three synapomorphies, including a novelty (unique synapomorphy): male dorsal gonostylus with a deep cleft. All the subgenera of this genus are strictly monophyletic with the exception of Placidohalictus, which is a paraphyletic one in relation to Vestitohalictus and Mucoreohalictus. A synonymical catalogue of species and species-group names in the genera Halictus and Seladonia, including 442 names, is provided as an appendix.

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Revision of the genus Peirates Serville, with a cladistic and biogeographic analysis (Heteroptera: Reduviidae, Peiratinae)

Insect Systematics & Evolution ◽

10.1163/187631297x00150 ◽

1997 ◽

Vol 28 (1) ◽

pp. 39-73 ◽

Cited By ~ 3

Author(s):

MARIA DEL CARMEN Coscaron

Keyword(s):

Phylogenetic Analysis ◽

Indian Ocean ◽

Geographical Distribution ◽

Sister Group ◽

Species Group ◽

Sister Group Relationship ◽

Palearctic Region ◽

Nomina Dubia ◽

Species Groups ◽

Biogeographic Analysis

AbstractThe genus Peirates Serville is revised. A total of 32 species are recognized. Its species are assigned to the P. quadrinotatus, P. singularis and P. lepturoides species groups. The P. quadrinotatus species group includes P. argenteopilosus Schouteden and P. quadrinotatus Fabricius; the P. singularis species group comprises its sole species P. singularis StÅl, and the P. lepturoides species group is formed by P. atromaculatus (StÅl) (= P. buruanus Miller, P. consobrinus Miller, P. excelsus Miller, P. immaculatus Miller, P. sordidus Miller, syn. n.), P. monodi Villiers, P. mundulus StÅl, P. nanus Miller, P. niger Signoret, P. nitidicollis Reuter, P. ochripennis Jeannel, P. perinetensis Villiers, P. strepitans Rambur, P. stridulus (Fabricius), P. tellini Schouteden, P. turpis Walker, P. aurigans Distant, P balteatus Germar, P. castaneipennis Reuter, P. cinctiventris Horváth, P. collarti Schouteden, P. confusus Schouteden (= P. amieti Villiers, syn. n.), P. crudelis sp. n., P. dimidiatus Walker (= P. areatus Miller, syn. n.), P. diola Lindberg, P. hybridus (Scopoli), P. lepturoides (Wolff) (= P. mimulus Miller, P. musculus Miller, P. sparsus Miller, syn. n.), P. macilentus Miller, P. marginiventris Distant, P. maurus StÅl, P. monodi Villiers, P. mundulus StÅl, P. nanus Miller, P. nigerrimus sp. n., P. nitidicollis Reuter, P. ochripennis Jeannel, P. perinetensis Villiers, P strepitans Rambur, P. stridulus (Fabricius), P. tellini Schouteden, and P. turpis Walker. Besides, P. sepulchralis Distant and P. sin Linnavuori are transferred to Brachysandalus StÅl, and P. barbarus Miller, P. bicolor Distant, P bicoloripes Breddin, P. flavipes (Walker), P. montivagus Distant, P. pallens Miller, and P. semifasciatus to Lestomerus Amyot & Serville, and P. umbrosus StÅl to Microsandalus StÅl. P. concinnus Walker cannot be assigned to any existing genus. P. cyaneonitens Breddin was not studied, and P. parvulus (Signoret), P. punctum (Fabricius), P. vittatus Reuter are considered as nomina dubia. Species are described, illustrated, and their geographical distribution is mapped. A key for separating all the species is given. A phylogenetic analysis of the species is presented. In the cladogram, the P. quadrinotatus, P. singularis and P. lepturoides species groups are monophyletic and a sister group relationship exists between the P. quadrinotatus group and the combined P. singularis + P. lepturoides groups. The geographical distribution of the species shows an African-Indopacific generalized track with an Indian Ocean baseline. The genus Peirates is hypothesized to have a tropical Gondwanan origin, with subsequent spread into the Palearctic region.

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Hypotheses from mitochondrial DNA: congruence and conflict between DNA sequences and morphology in Dolichopodinae systematics (Diptera:Dolichopodidae)

Invertebrate Systematics ◽

10.1071/is09040 ◽

2010 ◽

Vol 24 (1) ◽

pp. 32 ◽

Cited By ~ 4

Author(s):

Marc Pollet ◽

Christoph Germann ◽

Samuel Tanner ◽

Marco Valerio Bernasconi

Keyword(s):

Phylogenetic Analysis ◽

Mitochondrial Dna ◽

Molecular Analysis ◽

Dna Sequences ◽

Morphological Characters ◽

Species Group ◽

Data Matrix ◽

Cyt B ◽

Molecular Phylogenetic ◽

Species Groups

The molecular phylogeny of the subfamily Dolichopodinae (Diptera : Dolichopodidae) is reconstructed based on 79 species of 7 dolichopodine genera as ingroup, and 10 non-dolichopodine species from different genera as outgroup. A Bayesian analysis based on a mitochondrial DNA dataset consisting of 1702 characters (COI : 810; 12S : 366; 16S : 526) was carried out. Genital and non-genital morphological characters from a hitherto unpublished data matrix (based on 57 Dolichopodidae species) were used to explain and support the lineages hypothesised by our molecular phylogenetic analysis. The monophyly of the subfamily Dolichopodinae, and of the genera Dolichopus and Gymnopternus, was confirmed. The molecular analysis yielded nine species groups in Dolichopus that were proposed in previous studies using COI and Cyt-b. No evidence was found to support a clade including Dolichopus, Ethiromyia, and Gymnopternus. The genus Hercostomus proved polyphyletic with respect to Poecilobothrus, Sybistroma, and Gymnopternus. The following lineages were represented by strongly supported clades: Hercostomus germanus species group, H. vivax species group, H. nigrilamellatus species group, H. plagiatus species group, H. longiventris species group, H. fulvicaudis species group, and Poecilobothrus, Gymnopternus, Tachytrechus and Sybistroma (including Hercostomus nanus and H. parvilamellatus). Two clades that were previously established on the basis of morphology were confirmed in our phylogenetic analysis: (i) Poecilobothrus and the flower-feeding Hercostomus germanus species group, and (ii) the H. longiventris lineage and Sybistroma. In most cases, the groups identified in the molecular analysis could be supported and explained by morphological characters. Species of the Hercostomus germanus species group, Poecilobothrus, the Hercostomus longiventris species group, and a Sybistroma subclade have a similar microhabitat affinity.

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Taxonomic revision and cladistic analysis of ghost spiders of the genus Tafana Simon, 1903 (Araneae: Dionycha, Anyphaenidae), with the descriptions of twelve new species

European Journal of Taxonomy ◽

10.5852/ejt.2021.742.1291 ◽

2021 ◽

Vol 742 ◽

pp. 1-77

Author(s):

Luiz Fernando M. De Oliveira ◽

Antonio Domingos Brescovit

Keyword(s):

Phylogenetic Analysis ◽

New Species ◽

New Records ◽

Cladistic Analysis ◽

Taxonomic Revision ◽

Sister Group ◽

Species Group ◽

Genital Morphology ◽

Distribution Maps ◽

Species Groups

The diversity of the genus Tafana Simon, 1903 is poorly known in the Neotropical regions. In this work we provide a taxonomic review of the genus as well as a phylogenetic analysis. The ingroup of the analysis is composed of sixteen species of Tafana and the outgroup is composed of five representatives of Anyphaenidae. The sister-group recovered for Tafana is the clade Aysha + Xiruana, being supported by the embolic process on the male bulb. Two species groups within Tafana are herein proposed, the silhavyi group and the riveti group, based on two exclusive synapomorphies in the male bulb. We redescribe Tafana quelchi and present a description of the previously unknown female of Tafana silhavyi, both from Venezuela. In addition, we describe the first adult specimens of Tafana straminea. Twelve new species, along with several previously described species, are described, illustrated and mapped: T. riveti, T. straminea, T. quelchi, T. kunturmarqa sp. nov., T. humahuaca sp. nov., T. pastaza sp. nov., T. nevada sp. nov., T. huatanay sp. nov. and T. ruizi sp. nov. from the riveti species group; T. maracay sp. nov., T. arawak sp. nov., T. chimire sp. nov. and T. pitieri sp. nov. from the silhavyi species group; T. oliviae sp. nov. from Argentina and T. orinoco sp. nov. from Venezuela, neither of which belongs to any species group. We also discuss the genital morphology of the species groups based on the results of the phylogenetic analysis. Furthermore, distribution maps for all species, including new records for T. riveti, T. straminea and T. quelchi, are presented.

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Phylogenetic analysis of the Paulsoni species group (Decapoda : Alpheidae) from the American Pacific, with implications for the phylogenetic classification of the genus Synalpheus

Zootaxa ◽

10.11646/zootaxa.1744.1.2 ◽

2008 ◽

Vol 1744 (1) ◽

pp. 19 ◽

Cited By ~ 3

Author(s):

MARGARITA HERMOSO-SALAZAR ◽

MARY WICKSTEN ◽

JUAN J. MORRONE

Keyword(s):

Phylogenetic Analysis ◽

Cladistic Analysis ◽

Morphological Characters ◽

Species Group ◽

Natural Classification ◽

Phylogenetic Classification ◽

Species Groups

A cladistic analysis of 22 species of Synalpheus, represented primarily by species of the Paulsoni species group from the American Pacific and selected species from the Gambarelloides, Neomeris, Brevicarpus, and Biunguiculatus species groups was undertaken, based on 51 morphological characters. The Paulsoni species group proved to be paraphyletic, because species of the Neomeris, Brevicarpus, and Biunguiculatus species groups nested within it. It is proposed herein that in order to achieve a more natural classification, only two groups should be maintained within Synalpheus: Gambarelloides and Paulsoni, the latter in its broadest sense, treating the remaining species groups as synonyms.

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Phylogenetic relationships within Philodromidae, with a taxonomic revision of Philodromus subgenus Artanes in the western Palearctic (Arachnida:Araneae)

Invertebrate Systematics ◽

10.1071/is08044 ◽

2009 ◽

Vol 23 (2) ◽

pp. 135 ◽

Cited By ~ 10

Author(s):

Christoph Muster

Keyword(s):

Phylogenetic Analysis ◽

Eastern Mediterranean ◽

Taxonomic Revision ◽

Morphological Characters ◽

Species Group ◽

Caspian Region ◽

Western Palearctic ◽

The Family ◽

Palearctic Species ◽

Species Groups

The first quantitative phylogenetic analysis within the spider family Philodromidae (using 65 morphological characters from 40 ingroup taxa) does not corroborate Schick’s division into the tribes Thanatini (Apollophanes O. P.-Cambridge, 1898, Pelloctanes Schick, 1965, Thanatus C. L. Koch, 1837; Tibellus, Simon, 1875) and Philodromini (Ebo Keyserling, 1884, Philodromus Walckenaer, 1826; Rhysodromus Schick, 1965). Instead, Ebo is sister to all other genera in the family, and a fraction of Philodromus (the histrio species-group = Rhysodromus Schick) are contained with the Thanatini. The forgotten genus Artanes Thorell, 1870 constitutes a well-defined distal taxon in a clade that contains the majority of Philodromus species. Here Artanes is considered a subgenus of Philodromus, and includes the margaritatus and the poecilus species-groups. The western Palearctic species of the subgenus are revised. Twelve species are (re-)described, keyed and illustrated: Philodromus blanckei (Wunderlich, 1995) (first description of ♀); P. calidus Lucas, 1846; P. femurostriatus, sp. nov. from the eastern Mediterranean; P. fuscomarginatus (De Geer, 1778); P. johani, sp. nov. from Crete; P. laricium Simon, 1875, removed from synonymy with P. corticinus (C. L. Koch); P. maghrebi, sp. nov. from northern Africa (♀ only); P. margaritatus (Clerck, 1757); P. parietalis Simon, 1875; P. pentheri, sp. nov. from the Caspian region and Albania (♀ only); P. pinetorum, sp. nov. from the Mediterranean; and P. poecilus (Thorell, 1872).

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Revision of the Oriental species of Calleida Latreille (sensu lato). Part 2: the C. discoidalis species group (Coleoptera, Carabidae, Lebiini)

ZooKeys ◽

10.3897/zookeys.806.30051 ◽

2018 ◽

Vol 806 ◽

pp. 87-120

Author(s):

Hongliang Shi ◽

Achille Casale

Keyword(s):

Phylogenetic Analysis ◽

New Species ◽

Morphological Characters ◽

Species Group ◽

Type Locality ◽

Valid Species ◽

Species Relationships ◽

Species Groups

The C.discoidalis species group of the genus Calleida Latreille from Asia (in the sense of Casale and Shi 2018) is revised with six species recognized. Four new species are described: C.piligera Shi & Casale, sp. n. (type locality: Taiwan: Siling, 24.65°N, 121.42°E); C.cochinchinae Casale & Shi, sp. n. (type locality: Vietnam: “Cochinchina”); C.yunnanensis Shi & Casale, sp. n. (type locality: Yunnan: Caiyanghe, 22.60°N, 101.12°E); and C.luzonensis Casale & Shi, sp. n. (type locality: Philippines: Nagtipunan, 16.22°N, 121.60°E). C.fukiensis Jedlička, 1963 is confirmed as an available and valid species name, and C. suensoni Kirschenhofer, 1986 is newly synonymized with it. A phylogenetic analysis of Oriental Calleida species, based on adult morphological characters, is performed. The results show that the monophyly of most species groups in Oriental Calleida is accepted, but the C.discoidalis group appears polyphyletic and comprises three lineages. However, because many species relationships in the cladogram lack significant supporting, presently the C.discoidalis group was remained to use for morphological convenience. Five types of female reproductive tracts were recognized, corresponding to five branches in the cladogram.

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Beyond Wallace: a new lineage of Chrysorthenches (Lepidoptera: Yponomeutoidea: Glyphipterigidae) reveals a journey tracking its host-plants, Podocarpus (Pinopsida: Podocarpaceae)

Zoological Journal of the Linnean Society ◽

10.1093/zoolinnean/zlaa009 ◽

2020 ◽

Vol 190 (2) ◽

pp. 709-736

Author(s):

Jae-Cheon Sohn ◽

Shigeki Kobayashi ◽

Yutaka Yoshiyasu

Keyword(s):

Maximum Likelihood ◽

East Asia ◽

Host Plants ◽

Morphological Characters ◽

Species Group ◽

Geological History ◽

Dna Barcodes ◽

History Of ◽

Species Groups ◽

Maximum Likelihood Phylogeny

Abstract A northward trans-Wallacean radiation is demonstrated for Chrysorthenches, a member of the Orthenches group. Here we review Chrysorthenches and allied genera resulting in a generic transfer of Diathryptica callibrya to Chrysorthenches and two new congeners: C. muraseaeSohn & Kobayashisp. nov. from Japan and C. smaragdinaSohnsp. nov. from Thailand. We review morphological characters of Chrysorthenches and allied genera, and find polyphyly of Diathryptica and the association of the Orthenches-group with Glyphipterigidae. These findings were supported in a maximum likelihood phylogeny of DNA barcodes from ten yponomeutoids. We analysed 30 morphological characters for 12 species of Chrysorthenches, plus one outgroup, via a cladistic approach. The resulting cladogram redefined two pre-existing Chrysorthenches species-groups and identified one novel lineage: the C. callibrya species-group. We review the host associations between Chrysorthenches and Podocarpaceae, based on mapping the working phylogenies. Our review suggests that ancestral Chrysorthenches colonized Podocarpus and later shifted to other podocarp genera. Biogeographical patterns of Chrysorthenches show that they evolved long after the Podocarpaceae radiation. Disjunctive trans-Wallacean distribution of the C. callibrya species-group is possibly related to the tracking of their host-plants and the complicated geological history of the island-arc system connecting Australia and East Asia.

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Comparative morphology and species-groups of the oribatid mite genus Scapheremaeus (Acari: Oribatida: Cymbaeremaeidae), with new species from South Australia

Zootaxa ◽

10.11646/zootaxa.2213.1.1 ◽

2009 ◽

Vol 2213 (1) ◽

pp. 1-46 ◽

Cited By ~ 1

Author(s):

MATTHEW J. COLLOFF

Keyword(s):

New Species ◽

Oribatid Mite ◽

South Australia ◽

Biosphere Reserve ◽

Single Species ◽

Comparative Morphology ◽

Species Group ◽

Undescribed Species ◽

Species Groups ◽

Semi Arid

The morphology of the genus Scapheremaeus Berlese, 1910 is reviewed and characters of taxonomic utility delineated. Based on the morphological review, some 13 species-groups are outlined based on major morphotypes. There are two main categories: i) species that have a complete circumdorsal scissure with plicate microsculpture on the circumnotogastral plate and strongly contrasting microsculpture (foveolae, ridges or tubercles) on the centrodorsal plate (plicate species-groups), and ii) species with the circumdorsal scissure complete, incomplete or absent but with little or no contrast in microsculpture between the central and lateral regions: typically both regions foveolate or reticulate (non- plicate species-groups). A catalogue of world species of Scapheremaeus is provided. Scapheremaeus petrophagus (Banks, 1906) is not a Scapheremaeus but belongs to an undetermined genus in the Ameronothroidea. Cymbaeremaeus cyclops Oudemans, 1915 is recombined to Scapheremaeus. Five new species are described (S. angusi sp. nov., S. cheloniella sp. nov., S. ewani sp. nov., S. lambieae sp. nov., and S. pulleni sp. nov.) from soil and litter habitats in semi-arid Mallee eucalypt vegetation at Bookmark Biosphere Reserve, South Australia. These are the first members of the genus Scapheremaeus to be described from Australia, though undescribed species have been recorded previously. All the new species are morphologically closely-related and belong to a single species-group: Carinatus.

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