CLASSIFICATION AND EVOLUTION OF THE EUMENINE WASP GENUS SYMMORPHUS WESMAEL (HYMENOPTERA: VESPIDAE)

1989 ◽  
Vol 121 (S148) ◽  
pp. 5-168 ◽  
Author(s):  
Jeffrey M. Cumming
Keyword(s):  
Cladistic Analysis ◽  
Rio Grande Valley ◽  
Rio Grande ◽  
Neotropical Region ◽  
Type Locality ◽  
Nesting Behavior ◽  
Phylogenetic Position ◽  
Species Groups ◽  
Three New Species ◽  
New Synonymy

AbstractThe genus Symmorphus Wesmael is distributed throughout the Oriental, Palearctic, and Nearctic regions, as well as the northernmost Neotropical region, and appears to occupy a relatively isolated phylogenetic position within the Eumeninae. Females of Symmorphus, for those species in which nesting behavior is known, mass-provision cells in pre-existing cavities with chrysomelid, curculionid, or microlepidopterous larvae.Thirty-six species are recognized, including three new species and two new subspecies. Keys are provided for the identification of adult specimens, all species are described, and their geographical distributions mapped.Two subgenera, S. (Parasymmorphus) Cumming and Vecht and S. (Symmorphus) Wesmael, are recognized. Symmorphus (Parasymmorphus) includes S. parvilineatus (Cameron), S. momunganensis (Schulthess), and S. negrosensis Cumming and Vecht. Within S. (Symmorphus) three informal species groups are recognized, with the following included species and subspecies: (1) S. hoozanensis group — S. hoozanensis (Schulthess), S. alkimus alkimus Cumming and Vecht, S. alkimus dialeukus ssp.nov. (type-locality Sibolangit, N. Sumatra), S. ambotretus sp.nov. (type-locality Mt. Omei, Sichuan Province, China), and S. canlaonicus sp.nov. (type-locality Mt. Canlaon, Negros Oriental, Philippines); (2) S. declivis group — S. declivis Harttig; (3) S. murarius group — S. murarius (Linnaeus), S. foveolatus Gussakovskij, S. glasunowi Morawitz, S. sichuanensis Lee, S. crassicornis (Panzer), S. captivus (Smith), S. angustatus (Zetterstedt), S. albomarginatus albomarginatus (Saussure), S. albomarginatus midas ssp.nov. (type-locality Bentsen Rio Grande Valley State Park, Texas, USA), S. decens (Kostylev), S. violaceipennis Soika, S. mizuhonis Tsuneki, S. bifasciatus (Linnaeus), S. debilitatus (Saussure), S. cliens Soika, S. connexus (Curtis), S. lucens (Kostylev), S. sublaevis (Kostylev), S. canadensis (Saussure), S. projectus Bohart, S. tukvarensis (Meade-Waldo), S. nipteroides sp.nov. (type-locality N. Yourdou, India), S. fuscipes (Herrich-Schaeffer), S. apiciornatus (Cameron), S. aurantiopictus Soika, S. allobrogus (Saussure), S. cristatus (Saussure), S. gracilis (Brullé), and S. paralleliventris Soika.The following new synonymy is proposed: S. calciatii (Zavattari) = S. glasunowi Morawitz; S. nursei (Bingham), S. hospes (Nurse), and S. hospitans (Dalla Torre) = S. crassicornis (Panzer); S. hakutozanus Tsuneki and S. nansetsurei Tsuneki = S. angustatus (Zetterstedt); S. walshianus (Saussure) and S. meridionalis Viereck = S. albomarginatus albomarginatus (Saussure); S. kurentzovi Kurzenko, S. iiyamai Tsuneki, S. shiroyamai Tsuneki, S. piceanus Tsuneki, and S. sassai Tsuneki = S. mizuhonis Tsuneki; S. sinuatus (Fabricius), S. sinuatus mutinensis (Baldini), S. sparsus Morawitz, S. sinuatissimus Richards, S. mutinensis auster Soika, and S. mutinensis yezoanus Tsuneki = S. bifasciatus (Linnaeus); S. ishikawai Soika = S. lucens (Kostylev); S. seoulensis Tsuneki = S. apiciornatus (Cameron); S. pumilus (Saussure) and S. trisulcatus Cameron = S. cristatus (Saussure); S. elegans libanicus Soika = S. gracilis (Brullé); and S. allobrogus (Saussure) is removed from synonymy with S. bifasciatus (Linnaeus). Odynerus latipennis Smith and Odynerus cilicius Cameron, previously considered to belong within Symmorphus, are excluded from the genus, and S. sounkionis Tsuneki is transferred as Ancistrocerus sounkionis (Tsuneki) comb.nov. Neotypes are designated for S. crassicornis (Panzer) and S. fuscipes (Herrich-Schaeffer), and lectotypes are designated for S. foveolatus Gussakovskij, S. glasunowi Morawitz, S. nursei (Bingham), S. laeviventris (Thomson), S. albomarginatus (Saussure), S. decens (Kostylev), S. sinuatus (Fabricius), S. sinuatus mutinensis (Baldini), S. debilitatus (Saussure), S. lucens (Kostylev), S. sublaevis (Kostylev), S. debilis (Saussure), S. apiciornatus (Cameron), S. philadelphiae (Saussure), and S. elegans (Wesmael).A cladistic analysis of Symmorphus, based on 26 characters, is presented, including a discussion of the relationships of the genus to the remainder of the Eumeninae. Hypotheses outlining the historical zoogeography of Symmorphus are presented in light of current paleogeographic and paleoclimatic reconstructions, by comparing these cladistic relationships with the chorological patterns observed within the genus and other Vespidae.

Taxonomic revision of the Brazilian Atlantic Forest Atractus (Reptilia: Serpentes: Dipsadidae)

Zootaxa ◽  
2010 ◽  
Vol 2364 (1) ◽  
pp. 1 ◽  
Author(s):  
PAULO PASSOS ◽  
RONALDO FERNANDES ◽  
RENATO S. BÉRNILS ◽  
JULIO C. DE MOURA-LEITE
Keyword(s):  
New Species ◽  
Atlantic Forest ◽  
Taxonomic Status ◽  
Taxonomic Revision ◽  
Neotropical Region ◽  
Brazilian Atlantic Forest ◽  
Mountainous Regions ◽  
Species Groups ◽  
Three New Species ◽  
Maxillary Dentition

Dipsadine snakes of the genus Atractus are endemic to the Neotropical region, occurring from Panama to Argentina. Currently, the taxonomic status of most species of the genus is unclear and previous attempts of taxonomic revisions have been local in scale. In this paper we evaluate the taxonomic status of the Brazilian Atlantic Forest species of Atractus based on meristic, morphometric, maxillary dentition, and hemipenis characters. Quantitative and qualitative analyses suggest the recognition of one new species (A. caete sp. nov.) from the state of Alagoas, another (A. francoi sp. nov.) from the mountainous regions of the states of Rio de Janeiro and São Paulo, and the synonymy of A. kangueryensis with A. thalesdelemai. Specimens previously assigned to A. taeniatus in Argentina and Brazil are here considered A. paraguayensis. A key to the Atlantic Forest Atractus is provided and three new species groups are proposed for some cisAndean Atractus, mainly on the basis of hemipenial morphology: the A. emmeli, A. maculatus, and A. pantostictus species groups.

scholarly journals Phylogenetic position of Dendropsophus gaucheri (Lescure and Marty 2000) highlights the need for an in-depth investigation of the phylogenetic relationships of Dendropsophus (Anura: Hylidae)

Zootaxa ◽  
2011 ◽  
Vol 3035 (1) ◽  
pp. 59 ◽  
Author(s):  
ANTOINE FOUQUET ◽  
BRICE P. NOONAN ◽  
MICHEL BLANC ◽  
VICTOR GOYANNES DILL ORRICO
Keyword(s):  
Phylogenetic Relationships ◽  
Species Group ◽  
Genetic Distances ◽  
Type Locality ◽  
Phylogenetic Position ◽  
Guiana Shield ◽  
Mtdna Sequences ◽  
Species Groups

Dendropsophus gaucheri is a recently described species which inhabits open areas of the eastern part of the Guiana Shield and is currently assigned to the D. parviceps species group based on the presence of a subocular cream spot. Herein we investigate its phylogenetic position including material from the type locality and newly documented populations from Suriname and Brazil based on mtDNA sequences. The species, as well as D. riveroi which is assigned to the D. minimus species group, were recovered nested within the D. microcephalus species group which implies the paraphyly of the three Dendropsophus species groups. Such result, along with other evidences, highlights the need for a thorough revision of the genus. The genetic distances among D. gaucheri samples studied are low confirming their conspecificity and suggesting recent connections among populations from open areas currently isolated by rainforest in the lowlands of the Guiana Shield.

scholarly journals Description of three new species of Apatelodidae from the southern neotropical region (Lepidoptera, Bombycoidea)

ZooKeys ◽  
2018 ◽  
Vol 788 ◽  
pp. 3-17
Author(s):  
Daniel Herbin ◽  
Hernán Mario Beccacece
Keyword(s):  
New Species ◽  
Detailed Examination ◽  
Neotropical Region ◽  
Three New Species ◽  
New Synonymy

Three new species of Apatelodidae are described from Argentina, Bolivia, Brazil, and Paraguay: Apatelodesnavarroisp. n., Apatelodeschalupaesp. n., and Apatelodesulfisp. n., and are figured with their genitalia. Detailed examination of primary types leads to the establishment of a new synonymy: A.florisa Schaus, 1929 = A.schreiteri Schaus, 1924, syn. n., and a revised status for another Apatelodidae species previously misplaced in the Bombycidae: Apatelodesbrunnea (Dognin, 1916), comb. n.

A phylogeny of the family Fanniidae Schnabl (Insecta:Diptera:Calyptratae) based on adult morphological characters, with special reference to the Austral species of the genus Fannia

2008 ◽  
Vol 22 (5) ◽  
pp. 563 ◽  
Author(s):  
M. C. Domínguez ◽  
S. A. Roig-Juñent
Keyword(s):  
Cladistic Analysis ◽  
Morphological Characters ◽  
External Morphology ◽  
Phylogenetic Position ◽  
Phylogenetic Hypothesis ◽  
Parsimony Analysis ◽  
Male And Female ◽  
Southern South America ◽  
The Family ◽  
Species Groups

The present study proposed a phylogenetic hypothesis of the family Fanniidae based on a cladistic analysis using characters from adult external morphology and female and male terminalia. The main purpose of this study was to clarify the phylogenetic position of newly described or poorly known species, mostly from southern South America, the Neotropics, Africa, Australia and New Zealand. In total, 151 characters from adult male and female external morphology and terminalia were scored for 78 species of Fanniidae. Ten continuous characters were included and analysed as such. Three genera of Fanniidae and all the species-groups and subgroups proposed for the genus Fannia, except for the admirabilis-group and the setifer-subgroup were included as terminal taxa. An heuristic parsimony analysis under implied weights was performed. The analysis recovered the monophyly of the Fanniidae and the genus Fannia, as well as the monophyly of several species-groups within Fannia. Male and female external morphological characters were, in general, highly homoplasious, whereas characters from male terminalia showed low level of homoplasy and provided resolution at suprageneric nodes and species-groups.

REVISION OF THE SPECIES OF FRONTINIELLA TOWNSEND (DIPTERA: TACHINIDAE)

1993 ◽  
Vol 125 (1) ◽  
pp. 11-45 ◽  
Author(s):  
James E. O’Hara
Keyword(s):  
New Mexico ◽  
Species Distributions ◽  
Neotropical Region ◽  
The Other ◽  
Type Locality ◽  
New Combination ◽  
Annotated List ◽  
Adult Females ◽  
Species Groups ◽  
Nearctic Region

AbstractThe genus Frontiniella Townsend is transferred from the Eryciini to the Goniini, based on the production of microtype eggs in adult females. Frontiniella is known only from the Nearctic Region and the northern extreme of the Neotropical Region. Eufrontina Brooks is newly synonymized with Frontiniella and all species formerly placed in that genus are transferred to Frontiniella. Frontiniella is hypothesized to be monophyletic based on two derived character states: long hairs on the male surstylus, and elongate posterior spiracles on the puparium. Two species groups are recognized, one with six species and the other with four: the parancilla-group with F. parancilla Townsend, F. incarcerata sp.nov. (type-locality Hwy. 366, Graham Co., Arizona, USA), F. spectabilis (Aldrich) (new combination), F. surstylata sp.nov. (type-locality near Olustee, Florida, USA), F. jorgenseni sp.nov. (type-locality 20 miles northwest of Kenna, New Mexico, USA), and F. ethniae (Brooks) (new combination); and the festinans-group with F. festinans (Aldrich and Webber), F. apache sp.nov. (type-locality Sulphur Draw, Chiricahua Mts., Arizona, USA), F. mitis (Curran), and F. regilla (Reinhard). This revision includes a key to the 10 known species of Frontiniella, a description of each species (including puparium where known), maps of species distributions, an annotated list of hosts (including corrections to previously published records and additions from material examined), and illustrations of taxonomically useful features.

scholarly journals Cladistic analysis of the Brazilian troglobitic harvestmen genus Iandumoema Pinto-da-Rocha (Opiliones : Gonyleptidae) with the description of three new species: a brief exercise over the use of troglomorphisms in cladistic analysis

2020 ◽  
Author(s):  
Ludson Neves de Ázara ◽  
Marcos Ryotara Hara ◽  
Rodrigo Lopes Ferreira
Keyword(s):  
New Species ◽  
Cladistic Analysis ◽  
Type Locality ◽  
Morphological Data ◽  
Phylogenetic Hypothesis ◽  
Minas Gerais State ◽  
The Matrix ◽  
Cave Environment ◽  
Three New Species ◽  
Subterranean Environment

From an ecological and evolutionary standpoint, troglobitic organisms are of special interest because they have evolved in, and are restricted to, the subterranean environment. Iandumoema Pinto-da-Rocha, 1997 stands out for being the only Brazilian harvestmen genus with more than one troglobitic species, with three species described from caves in Minas Gerais state. Traditionally, testing the monophyly of troglobitic groups is more challenging than testing groups that do not include troglobites. Many of their shared features might be the result of convergence or parallelism imposed by the cave environment, such as the absence of light, limited and infrequent availability of food resources and low population density, among others. In the case of Iandumoema, this becomes even more difficult because the genus is currently included in the species-rich and polyphyletic subfamily Pachylinae. This study tested the monophyly of this troglobitic genus and proposed the first phylogenetic hypothesis for Iandumoema based on cladistic analysis using morphological data. The analysis included all described species of Iandumoema and three new troglobitic species: I. cuca, sp. nov. (type locality: Itacarambi, Gruta da Água do João Ferreira); I. gollum, sp. nov. (type locality: Presidente Juscelino, Lapa D’Água); and I. stygia, sp. nov. (type locality: Montes Claros, Gruta do Cedro). The matrix comprises 79 characters and 28 terminal taxa: six species of Iandumoema; 14 of Pachylinae; six from other Gonyleptidae subfamilies; one species of Cosmetidae; and one of Metasarcidae. The cladistic analysis resulted in one parsimonious tree (339 steps, consistency index = 0.35, retention index = 0.56). Iandumoema is a monophyletic and well supported genus, nestled among Brazilian ‘Pachylinae’. Three new species are described and an identification key and ecological remarks for all six species of the genus Iandumoema is provided.

Revision of Nanophareus, a mysterious harvestman genus from Chile, with descriptions of three new species (Opiliones: Laniatores: Gonyleptidae)

Zootaxa ◽  
2012 ◽  
Vol 3579 (1) ◽  
pp. 37 ◽  
Author(s):  
MARCOS RYOTARO HARA ◽  
RICARDO PINTO-DA-ROCHA ◽  
ADRIANO BRILHANTE KURY
Keyword(s):  
New Species ◽  
Male Genitalia ◽  
Cladistic Analysis ◽  
Lateral View ◽  
Type Locality ◽  
Data Matrix ◽  
Lateral Margin ◽  
Three New Species ◽  
Tibia Length

The Chilean genus Nanophareus Roewer, 1929 is revised and three new species are described: N. araucanus sp. nov. (typelocality: Parque Nacional La Campana, Valparaíso, Chile); N. bipartitus sp. nov. (type locality: Parque Nacional La Cam-pana, Valparaíso, Chile); N. bosqenublado sp. nov. (type locality: Parque Nacional Fray Jorge, Coquimbo, Chile). Thetype species, N. palpalis Roewer, 1929, is redescribed and a lectotype is designated. A cladistic analysis was performedusing these three new species plus N. palpalis and 14 more laniatorid species, and a data matrix of 72 characters: Sevenfrom the ocularium, 22 from the dorsal scutum, one from the venter, one from the chelicera, eight from the pedipalp, 24from male legs, and nine from male genitalia. Two equally most parsimonious trees were found (L = 210; C.I. = 0.41; R.I.= 0.51). Nanophareus was recovered as nested within a paraphyletic subfamily Pachylinae. The genus Nanophareus wasfound to be monophyletic based on the following exclusive synapomorphies: An external row of enlarged tubercles in-serted among small ones on lateral margin of the dorsal scutum (innapplicable in N. bosqenublado); the ventro-basal mar-gin of pedipalpal tibia curved 90 degrees in lateral view; and retrolateral seta of the pedipalpal tibia with a socket apically bifid (socket and seta longer than pedipalpal tibia length).

THE SPECIES GROUPS OF GONATOCERUS NEES IN NORTH AMERICA WITH A REVISION OF THE SULPHURIPES AND ATER GROUPS (HYMENOPTERA: MYMARIDAE)

1988 ◽  
Vol 120 (S141) ◽  
pp. 3-110 ◽  
Author(s):  
John T. Huber
Keyword(s):  
North America ◽  
Cladistic Analysis ◽  
Junior Synonym ◽  
Species Groups ◽  
New Synonymy ◽  
Group Species

AbstractA provisional cladistic analysis of the species groups of Gonatocerus Nees is presented, based on 24 characters. Six species groups are defined, but only the four Nearctic ones are described. The species groups proposed are the ater, deficiens, litoralis, membraciphagus, straeleni, and sulphuripes groups. The ater group is divided into the morrilli, ater (s.s.), and bucculentus subgroups. Keys are provided to distinguish the species groups, and females of the sulphuripes- and ater-group species. Twenty-four species are recognized in the sulphuripes and ater groups for America north of Mexico. Eight species are recognized in the sulphuripes group, of which californicus, koebelei, maga, mexicanus, rivalis, and utahensis are redescribed. Two species, floridensis and lissonotus, are described as new. One new synonymy is proposed: titillatus is considered a junior synonym of utahensis. Sixteen species are recognized in the ater group, of which ashmeadi, capitatus, dolichocerus, fasciatus, latipennis, morrilli, novifasciatus, ovicenatus, and triguttatus are redescribed. Seven species, bucculentus, enicmophilus, flagellatus, impar, incomptus, inexpectatus, and inflatiscapus, are described as new. Two new synonymies are proposed: maximus is considered a junior synonym of latipennis, and marilandicus a junior synonym of dolichocerus. Lectotypes are designated for ashmeadi, capitatus, koebelei, mexicanus, morrilli, ovicenatus, triguttatus, and utahensis.

Description of three new species of the tropical Asian jumping spider genus Onomastus Simon, 1900 from high altitude cloud forests of Sri Lanka (Araneae: Salticidae)

Zootaxa ◽  
2016 ◽  
Vol 4205 (5) ◽  
pp. 431 ◽  
Author(s):  
SURESH P. BENJAMIN ◽  
NILANI KANESHARATNAM
Keyword(s):  
New Species ◽  
Sri Lanka ◽  
High Altitude ◽  
Cloud Forest ◽  
Cladistic Analysis ◽  
Morphological Characters ◽  
Evergreen Forest ◽  
Phylogenetic Position ◽  
Jumping Spider ◽  
Three New Species

Spiders of the tropical Asian jumping spider genus Onomastus Simon, 1900 are small to medium-sized, delicate, translucent, commonly found inhabitants of Asian evergreen forest foliage. In this paper, three new species of the genus, O. jamestaylori sp. nov. (♂♀), O. corbetensis sp. nov. (♂♀) and O. maskeliya sp. nov. (♂♀) are described from Sri Lanka. The three new species are added to the matrix of a previous study to assess their phylogenetic position. The resulting cladistic analysis, based on 35 morphological characters from 18 taxa (13 Onomastus species and 5 outgroups) supports the monophyly of the genus. Additionally, a monophyletic, well-supported South Asian clade (India, Sri Lanka), which is restricted to the Sri Lanka-Western Ghats biodiversity hotspot, is recovered in most analysis. The three newly described species might be endangered due to their small population size and restricted distribution in high altitude cloud forest. 

Revision of the Mecodema curvidens species group (Coleoptera: Carabidae: Broscini)

Zootaxa ◽  
2011 ◽  
Vol 2829 (1) ◽  
pp. 1 ◽  
Author(s):  
DAVID S. SELDON ◽  
RICHARD A. B. LESCHEN
Keyword(s):  
East Coast ◽  
Cladistic Analysis ◽  
Species Group ◽  
The Other ◽  
Data Matrix ◽  
The North ◽  
Apical Plate ◽  
Apical Lobe ◽  
Species Groups ◽  
New Synonymy

The flightless carabid genus Mecodema Blanchard 1843 is restricted to New Zealand and presently contains 64 species. In this study, we examine species in the newly constituted curvidens group (sulcatum and curvidens are synonymised), reducing the Mecodema species groups to seven. They share two synapomorphies (rounded apical lobe of the aedeagus and lack of microsculpture on the vertex of the head) and are distributed in Northland, south along the east coast of the North Island and the northeast portion of the South Island. Adult specimens of the curvidens and sulcatum groups, along with exemplars of the other Mecodema groups (alternans, costellum, ducale, infimate, laterale and spiniferum), and an outgroup Oregus Putzeys 1868 were examined for cladistic analysis using a data matrix composed of a 63 characters and 21 terminal taxa. The analysis resulted in 18 most parsimonious trees. The following new species in the curvidens group are described: M. aoteanoho sp. n., M. haunoho sp. n., M. manaia sp. n., M. parataiko sp. n., M. ponaiti sp. n., and M. tenaki sp. n. Mecodema exitiosus was wrongly sysnonymised under M. occiputale and we propose a new synonymy M. curvidens (=M. exitiosus). We consider the parts of the male genitalia in detail with special attention to the structures of apical plate once everted from the endophallus.

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