SYSTEMATICS OF NEARCTIC SCYTHRIDIDAE (LEPIDOPTERA: GELECHIOIDEA): PHYLOGENY AND CLASSIFICATION OF SUPRASPECIFIC TAXA, WITH A REVIEW OF DESCRIBED SPECIES

1991 ◽  
Vol 123 (S160) ◽  
pp. 3-341 ◽  
Author(s):  
Jean-François Landry

AbstractGenera and previously described species of Nearctic Scythrididae are revised for the first time, based on the study of adult structures. About 90 percent of the Nearctic fauna known in collections consists of undescribed species. The supraspecific taxa treated in this work encompass less than half of the Nearctic species diversity. Only six new species are described, all within the largest and structurally most diverse genus. The status of all nominal species is revised. Valid species are redescribed and their features illustrated. General problems in the systematics of the Scythrididae are discussed. A description of adult features of the family Scythrididae is providad. Extra-limital genera are briefly reviewed. A key to the Nearctic genera and informal supraspecific lineages is provided.Six genera, including three new, are treated: Areniscythris Powell, 1976, Arotrura Walsingham, 1888, Asymmetrura gen. nov., Neoscythris gen. nov., Rhamphura gen. nov., and Scythris s. str. Hübner, [1825]. Areniscythris includes a single described species, Areniscythris brachypteris Powell, but is defined more broadly to account for a number of undescribed species. Arotrura is divided into nine informal species groups with the following included species: Arotrura atascosa sp. nov., Arotrura balli sp. nov., Arotrura divaricata (Braun) comb, nov., Arotrura eburnea Walsingham, Arotrura formidabilis sp. nov., Arotrura hymenata sp. nov., Arotrura longissima sp. nov., Arotrura oxyplecta (Meyrick) comb, nov., Arotrura powelli sp. nov., and Arotrura sponsella (Busck) comb. nov. Asymmetrura includes: Asymmetrura albilineata (Walsingham) comb. nov., Asymmetrura graminivorella (Braun) comb. nov., Asymmetrura impositella (Zeller) comb. nov. and type species, Asymmetrura matutella (Clemens) comb, nov., Asymmetrura reducta (Braun) comb, nov., and Asymmetrura scintillifera (Braun) comb. nov. Neoscythris includes: Neoscythris confinis (Braun) comb, nov., Neoscythris euthia (Walsingham) comb. nov., Neoscythris fissirostris (Meyrick) comb. nov. and type species, and Neoscythris planipenella (Chambers) comb. nov. Rhamphura includes: Rhamphura altisierrae (Keifer) comb, nov., Rhamphura ochristriata (Walsingham) comb. nov. and type species, Rhamphura perspicillella (Walsingham) comb. nov., Rhamphura suffusa (Walsingham) comb. nov., and the extra-limital Rhamphura immunis (Meyrick) comb. nov. from Peru. Scythris s. str. includes: Scythris immaculatella (Chambers) rev. stat., Scythris limbella (Fabricius), Scythris mixaula Meyrick, Scythris trivinctella (Zeller), and Scythris ypsilon Braun. A further eight species are phylogenetically distinct from Scythris s. str. but provisionally are only assigned to five informal monophyletic lineages until their cladistic relationships are more firmly established. These are: the Scythris basilaris lineage including Scythris basilaris (Zeller), Scythris eboracensis (Zeller), and Scythris fuscicomella (Clemens); the Scythris interrupta lineage including Scythris interrupta Braun; the Scythris inspersella lineage including Scythris inspersella (Hübner) and Scythris noricella (Zeller); the Scythris anthracina lineage including Scythris anthracina Braun; and the Scythris charon lineage including Scythris charon Meyrick. Three species are incertae sedis: Scythris inornatella (Chambers) comb, nov., Scythrispilosella (Zeller), and Scythris piratica Meyrick.Coleophora albacostella Chambers and Coleophora inornatella Chambers are transferred from the Coleophoridae. Scythris arizoniella (Kearfott) is transferred to the Coleophoridae [Coleophora arizoniella (Kearfott) comb. nov.].The following new synonymy is proposed: Colinita Busck, 1907 = Arotrura Walsingham, 1888; Gelechia aterrimella Walker, 1864 and Scythris epilobiella McDunnough, 1942 = Scythris inspersella [Hübner, (1817)]; Scythris magnatella Busck, 1904 = Scythris noricella (Zeller, 1843); Scythris pacifica McDunnough, 1927 = Scythris immaculatella (Chambers, 1875); Coleophora albacostella Chambers, 1875 and Scythris hemidictyas Meyrick, 1928 = Neoscythris planipenella (Chambers, 1875).A cladistic definition of the family is presented for the first time. The monophyly of the Scythrididae is supported by the following synapomorphies: very narrow ductus bursae, broad ductus seminalis anastomosed with the oviduct and the corpus bursae, lack of signum, unique shape of the apophyses of the metathoracic furca, tarsomeres 1–4 with two subapical spurs, aedeagus ankylosed, and origin of forewing veins R4 and R5 on a common stalk with R4 extended to the costa and R5 to the termen. Relationships of the Scythrididae within the Gelechioidea are discussed. Based on the cladistic analysis of 52 structural characters, phylogenetic relationships of supraspecific taxa are inferred. Two cladograms, one for the genera and one for the species groups of Arotrura, are presented and used in deriving the classification.

1975 ◽  
Vol 107 (1) ◽  
pp. 1-43 ◽  
Author(s):  
Evert E. Lindquist

AbstractConceptual and nomenclatural problems of Digamasellus Berlese, 1905 and Dendrolaelaps Halbert, 1915 are reviewed. It is shown that Digamasellus punctum (Berlese, 1904) is conspecific with D. perpusillus Berlese, 1905, the type-species of Digamasellus Berlese, 1905 (new synonymy). Hence, the genus in which punctum is included must take the name Digamasellus.A new case is made for recognizing Digamasellus and Dendrolaelaps as distinct genera in the Digamasellidae. Two other genera of Digamasellidae are recognized, Dendroseius Karg, 1965 and a broadened concept of Longoseius Chant, 1961. Two new subgenera are proposed, Dendrolaelaspis in the genus Dendrolaelaps, and Longoseiulus in the genus Longoseius. Diagnoses, descriptions, and a key to these genera and subgenera of Digamasellidae, along with a comprehensive description of the family, are presented. The phylogenetic relationship of the Digamasellidae in the Rhodacaroidea, and some thoughts on phylogeny within the Digamasellidae are given.A second species of Digamasellus sensu stricto, D. australis, is described as new, and the female and male adults of the type-species of Longoseius, L. (L.) cuniculus Chant, are described for the first time. New combinations include: Dendrolaelaps (Dendrolaelaspis) orientalis (Bhattacharyya), Longoseius (Longoseiulus) longulus (Hirschmann), L. (L.) ornatus (Hirschmann), L. (L.) aberrans (Hirschmann), and L. (L.) brachypoda (Hurlbutt).The absence of the protonymphal seta, md, from the telotarsi of legs II to IV is noted as a singular deficiency in the leg setation of Longoseius cuniculus Chant. This seta is not known to be absent in any other species among the families of Gamasina.A paper published by Hirschmann while the present work was in press is considered in an addendum to this paper. The subgeneric name Dendrolaelaps (Tridendrolaelaps) Hirschmann, 1974 is an objective junior synonym of Digamasellus Berlese, 1905, and the latter also has priority over Dendrolaelaps Halbert, 1915 so long as both names are applied within the same genus. A lectotype is designated for the nominal species Digamasellus punctum (Berlese). The status of the subgenus Dendrolaelaps (Multidendrolaelaps) Hirschmann, 1974 is problematic, pending a more comprehensive diagnosis providing data sufficient to indicate whether this is a monophyletic group. The quadrisetus group is newly proposed for some of the species placed by Hirschmann in the armatus group of Multidendrolaelaps. Digamasellus badenhorsti (Ryke) is tentatively considered as the second known species of Dendroseius Karg. Hirschmann's opinion, that Longoseius Chant warrants no more than a species-group in Dendrolaelaps, is disputed.


Zootaxa ◽  
2019 ◽  
Vol 4685 (1) ◽  
pp. 1-67 ◽  
Author(s):  
IRENE LOBATO-VILA ◽  
JULI PUJADE-VILLAR

A taxonomic revision of the tribe Ceroptresini (Hymenoptera: Cynipidae) is conducted for the first time. Prior to this study, the total number of valid species of Ceroptres, the only genus within Ceroptresini to date, was 23. As a result of this revision, 15 Ceroptres species are retained as valid and one species, Amblynotus ensiger Walsh, 1864, is desynonymized from Ceroptres petiolicola (Osten-Sacken, 1861), being considered here as a valid Ceroptres species: C. ensiger (Walsh, 1864) status verified and comb. nov. An additional five new species are described from Mexico: Ceroptres junquerasi Lobato-Vila & Pujade-Villar sp. nov.; C. lenis Lobato-Vila & Pujade-Villar sp. nov.; C. mexicanus Lobato-Vila & Pujade-Villar sp. nov.; C. nigricrus Lobato-Vila & Pujade-Villar sp. nov.; C. quadratifacies Lobato-Vila & Pujade-Villar sp. nov., increasing the total number of valid Ceroptres species to 21. Ceroptres masudai Abe, 1997 is synonymized with C. kovalevi Belizin, 1973. Ceroptres niger Fullaway, 1911 is transferred to Andricus (Andricus confusus Lobato-Vila & Pujade-Villar comb. nov. and nom. nov.). Five species (Amblynotus inermis Walsh, 1864; Cynips quercusarbos Fitch, 1859; Cynips querficus Fitch, 1859; Cynips quercuspisum Fitch, 1859; and Cynips quercustuber Fitch, 1859) are not considered as valid Ceroptres. The status of Ceroptres quereicola (Shinji, 1938), previously classified as an unplaced species, is commented on. In addition, a Nearctic species from the USA, Ceroptres politus Ashmead, 1896, is here proposed as the type species of a new genus within Ceroptresini: Buffingtonella Lobato-Vila & Pujade-Villar gen. nov. Redescriptions, biological and distribution data, illustrations and keys to genera and species within Ceroptresini are provided. The diagnostic morphological traits of Ceroptresini, Ceroptres and the new genus are discussed. 


Zootaxa ◽  
2007 ◽  
Vol 1418 (1) ◽  
pp. 1-628 ◽  
Author(s):  
CARL J. FERRARIS

A checklist of Recent and fossil catfishes (Order Siluriformes) is presented, summarizing taxonomic literature published through 2005. From 4624 nominal species group names and 810 genus group names, 3093 species are recognized as valid, and are distributed among 478 genera and 36 families. Distributional summaries are provided for each species, and nomenclatural synonymies, including relevant information on all name-bearing types, are included for all taxa. One new name is proposed herein: Clariallabes teugelsi, as a replacement for Clarias (Allabenchelys) dumerili longibarbis David & Poll, 1937, which is preoccupied by Clarias longibarbis Worthington, 1933, but has been treated as a valid species of Clariallabes by Teugels. Acrochordonichthys melanogaster Bleeker, 1854, is designated as type species of Acrochordonichthys Bleeker, 1857, inasmuch as no earlier valid designation has been found. A new genus Pseudobagarius, is proposed for the “pseudobagarius group” of species formerly placed in Akysis. The status of 228 species group names remains unresolved and 31 names based on otoliths ascribed to catfishes are listed but not placed into the checklist. The current emphasis given to catfish taxonomy at present is likely to result in a dramatic increase in the total number of valid taxa as well as major changes in the membership of some of the higher level taxa recognized here.


1980 ◽  
Vol 11 (4) ◽  
pp. 428-453 ◽  
Author(s):  
Göran Nordlander

AbstractThe genus Leptopilina Förster is revised on a world basis. The identity of 31 nominal species, including type species of four other genera, is discussed and 15 lectotypes are designated. Valid species of Leptopilina are L. longipes (Hartig) [type species], L. clavipes (Hartig) n. comb., L. heterotoma (Thomson) n. comb., L. fimbriata (Kieffer) n. comb., L. rufipes (Cameron) n. comb., L. atraticeps (Kieffer) n. comb., L. mahensis (Kieffer) n. comb., L. boulardi (Barbotin et al.) n. comb., and L. cupulifera (Kieffer) n. comb. Leptopilina heterotoma is the valid name for the well-known Drosophila parasitoid currently referred to as Pseudeucoila bochei (Weld). Leptopilina is not synonymous with Ganaspis Förster. Tryhliographa Förster is a senior synonym of Episoclu Förster n. syn. and of Pseudeucoila Ashmead (synonymy confirmed). Leptopilina is described and compared with related genera (Cothonaspis, Rhoptromeris, Odonteucoila). L. victoriae n. sp. is described from the Seychelles. The five Leptopilina occurring in Europe are redescribed on modern material and a key is given to them.


1978 ◽  
Vol 56 (9) ◽  
pp. 1925-1961 ◽  
Author(s):  
Alex E. Peden ◽  
M. Eric Anderson

Six of 11 nominal species of Lycodapus, L. australis Norman, L. derjugini Andriashev, L. dermatinus Gilbert, L. fierasfer Gilbert, L. mandibularis Gilbert, and L. parviceps Gilbert, are recognized. Original descriptions and the disintegrated type do not permit adequate diagnosis of L. extensus Gilbert, which we treat as a nomen dubium. The holotype of L. microchir Schmidt must be reexamined before the status of this nominal species can be assessed. We describe L. endemoscotus n.sp. and L. pachysoma n.sp. off the west coast of North America and another population of L. pachysoma from the Antarctic. The species of Lycodapus form two natural groupings. Long gill rakers and a tendency toward fewer preoperculomandibular head pores characterize L. australis, L. derjugini, L. dermatinus, and L. fierasfer. Short gill rakers, single interorbital pore, and more preoperculomandibular pores typify the remaining species of which L. endemoscotus, L. mandibularis, L. pachysoma, and another undescribed species from the Antarctic possess broad, stout rakers. Very reduced rakers indicate L. parviceps as the most differentiated of the short-rakered species. Descriptions, spot distribution maps, synonymies, and illustrations are provided for all named valid species.


ZooKeys ◽  
2021 ◽  
Vol 1025 ◽  
pp. 91-137
Author(s):  
Marília Pessoa-Silva ◽  
Marcos Ryotaro Hara ◽  
Ricardo Pinto-da-Rocha

Species of the genus Sadocus Sørensen, 1886 are conspicuous gonyleptids that occur in Chile and Argentina. Here, the genus is revised for the first time and the cladistic analysis based on morphological characters does not corroborate its monophyly unless a phylogenetically unrelated species is excluded (explained further on). A new classification is proposed for the seven species left in the genus and considered valid, of the 13 nominal species previously recognized. Two out of the seven valid species are considered as species inquirendae: Sadocus allermayeri (Mello-Leitão, 1945) [= Carampangue allermayeri Mello-Leitão, 1945] and Sadocus nigronotatus (Mello-Leitão, 1943) [= Carampangue nigronotatum Mello-Leitão, 1943]. The following synonymies are proposed: Sadocus bicornis (Gervais, 1849) [original combination = Gonyleptes bicornis Gervais, 1849] is a junior synonym of Sadocus asperatus (Gervais, 1847) [= Gonyleptes asperatus Gervais, 1847]; Sadocus conspicillatus Roewer, 1913, Sadocus exceptionalis (Mello-Leitão, 1946) [= Araucanoleptes exceptionalis Mello-Leitão, 1946] and Sadocus guttatus Sørensen, 1902 are junior synonyms of the valid name Sadocus polyacanthus (Gervais, 1847) [= Gonyleptes polyacanthus Gervais, 1847]; and Sadocus calcar (Roewer, 1913) [= Lycomedes calcar Roewer, 1913] is a junior synonym of the valid name Gonyleptes horridus Kirby, 1819. Sadocus brasiliensis Soares & Soares, 1949 is not congeneric with Argentinean/Chilean species of the genus according to the cladistic analysis and is here synonymized with Discocyrtus catharinensis (Mello-Leitão, 1923 [= Sadocus catharinensis Mello-Leitão, 1923]).


2000 ◽  
Vol 14 (6) ◽  
pp. 771 ◽  
Author(s):  
Guillermo Kuschel ◽  
Richard A. B. Leschen ◽  
Elwood C. Zimmerman

The historical status of the family Platypodidae is reviewed and the family is revised. Results of a cladistic analysis based on 35 terminal taxa and 80 adult morphological characters show that the current placement of Platypodidae makes the subfamily Scolytinae paraphyletic. Moreover, several important genera included in Scolytinae are shown to be members of Cossoninae (i.e. the placement of Protoplatypus Wood and Phylloplatypus Kato in Cossoninae is confirmed). Based on these results, the status of Platypodidae as a family and subfamily is rejected, Scolytinae thereby becoming a monophyletic taxon. Araucarius groups in Scolytinae instead of Cossoninae in the analysis on a single step only, but it is suggested that it be retained in Cossoninae until this subfamily is submitted to a similar phylogenetic study. Three genera and four species of Cossoninae are described as new: Dobionus Kuschel, gen. nov.: type species D. araucarinus Kuschel, sp. nov. (with the inclusion of D. brachyrhinus (Montrouzier)); Coptonus Kuschel, gen. nov.: type species C. fijianus Kuschel, sp. nov. (with the inclusion of C. papuanus Kuschel, sp. nov.) and Dissostomus Kuschel, gen. nov.: type species D. hornabrooki Kuschel, sp. nov.


1997 ◽  
Vol 28 (3) ◽  
pp. 271-305 ◽  
Author(s):  
Daniel Burckhardt ◽  
Pavel Lauterer

AbstractThe species of an ill-defined assemblage centred around Aphalara exilis are revised. Two species, viz. A. siamensis and A. taiwanensis, are described as new and three species are synonymised (A. soosi with A. exilis, A. kunashirensis with A. itadori, and A. augusta with A. fasciata). A neotype is designated for Tettigonia exilis, a lectotype is designated for Aphalara rumicis Mally, and Aphalara ulicis is removed from synonymy with Aphalara calthae. The species are diagnosed, illustrated and keyed. The revision of types of A. polygoni, the type species of Aphalara, showed that it had been misinterpreted and that it corresponds to A. rumicicola Klimaszewski which is synonymised with it. A. polygoni auct. nec Foerster is described as A. freji sp. n. A list is given of the 37 currently recognised valid species of Aphalara with their distributions and host plants, and of 4 nomina dubia. A cladistic analysis using PAUP yielded 5 most parsimonious trees suggesting that Aphalara contains six monophyletic species groups (the itadori (2 spp.), siamensis (1 sp.), exilis (7 spp.), rumicis (2 spp.), maculipennis (4 spp.) and calthae (21 spp.) groups). Biogeographical aspects are analysed in terms of area relationships (BPA), ancestral areas (methods of Bremer and Ronquist) and dispersal (cladistic subordinateness). Four species groups are restricted to the palaearctic, one to the nearctic and only one occurs in both. It is concluded that vicariance events may have occurred in the palaearctic but that the presence in the nearctic is probably due to dispersal. The east palaearctic is likely to have been part of the ancestral area of Aphalara. Three of the species groups are monophagous on Polygonum, two on Rumex and one is polyphagous on Polygonaceae (Polygonum, Rumex), Brassicaceae, Caryophyllaceae, Primulaceae and Ranunculaceae. Polygonum is the likely ancestral host association of Aphalara; Rumex has been colonised independently at least three times.


1977 ◽  
Vol 109 (S103) ◽  
pp. 1-66 ◽  
Author(s):  
J. F. McAlpine

AbstractThe systematic position of the Piophilidae sens. lat. within a group of nine schizophorous families with tephritid-like ovipositors (Lonchaeidae, Otitidae, Platystomatidae, Pyrgotidae, Tephritidae, Tachiniscidae, Richardiidae, Pallopteridae, and Piophilidae) is elucidated. It is shown to be a sister-group of the Pallopteridae, and these two families together with the Richardiidae comprise a monophyletic suprafamily unit within the larger group of families. The evolutionary changes that occurred within the Piophilidae are analyzed and the supposed phylogeny of its component taxa is portrayed. The family is redefined to include neottiophilids and thyreophorids and is classified into two subfamilies, Neottiophilinae and Piophilinae; the latter is divided into two tribes, Mycetaulini and Piophilini (with subtribes Piophilina and Thyreophorina). Twenty-three genera are recognized and taxonomically defined; this includes description of two new genera, Neopiophila and Parapiophila. Clusina Curran is synonymized withProtopiophila Duda (new synonymy), and four nominal species are placed in synonymy for the first time, i.e. Piophila anomala Malloch and Piophila setosa Melander and Spuler = Parapiophila vulgaris (Fallén) (new synonymy), Piophila flavifacies Brunetti = P. casei (Linnaeus) (new synonymy), and Mycetaulus pulchellus Banks = Mycetaulus longipennis Loew (new synonymy). Six new species, Actenoptera avalona (Newfoundland), Neopiophila setaluna (Northwest Territories), Protopiophila atrichosa (Peru and British Honduras), Protopiophila pallida (Peru and British Guiana), Prochyliza azteca (Mexico), and Prochyliza inca (Peru) are described. The following 20 new combinations are made: Mycetaulus lituratus (Melander and Spuler), Allopiophila testacea (Melander), Protopiophila nigriventris (Curran), Prochyliza lundbecki (Duda), nigricornis (Meigen), nigricoxa (Melander and Spuler) and nigrimana (Meigen), Arctopiophila arctica (Holmgren), Parapiophila atrifrons (Melander and Spuler), calceata (Duda), coerulescens (Zetterstedt), dudai (Frey), flavipes (Holmgren), lonchaeoides (Zetterstedt), nitidissima (Melander and Spuler), pectiniventris (Duda), penicillata (Steyskal), vulgaris (Fallén), and xanthopoda (Melander and Spuler). In all, 67 valid species are placed, their geographic distributions are outlined, and the zoogeographic implications are discussed. A lectotype is designated for Piophila flavitarsis Meigen = Madiza glabra Fallén (Milichiidae).A key to subfamilies, tribes, and genera is provided, and keys to world species are given where needed. An annotated world list of all names referred to the family (sens. lat.) is provided. The paper includes 58 figures, two tables, and 122 literature references.


2008 ◽  
Vol 22 (1) ◽  
pp. 55 ◽  
Author(s):  
Owen D. Seeman

Chrysomelobia Regenfuss and Parobia Seeman & Nahrung are sexually transmitted mites, the former being parasites of chrysomelid beetles in the Americas, Europe and Africa, the latter a radiation on the Paropsini (Chrysomelidae) in Australia. A cladistic analysis of Chrysomelobia and Parobia shows the type species of Chrysomelobia, C. mahunkai Regenfuss, is placed within Parobia. Therefore, Parobia is synonymised with Chrysomelobia, which has priority. I describe another nine new species of Chrysomelobia from Australian paropsine beetles: C. aquariolus, sp. nov., C. orthagoriscus, sp. nov. and C. cubile, sp. nov. from Paropsis porosa Erichson; C. lipsettae, sp. nov. from Dicranosterna pedestris (Chapuis); C. armstrongi, sp. nov. from Paropsisterna sp. nr minerva (Blackburn); C. pagurus, sp. nov. from Paropsis charybdis Stål; and C. nahrungae, sp. nov., C. vafer, sp. nov. and C. verecundus, sp. nov. from three species of Trachymela. Chrysomelobia captivus (Seeman & Nahrung) is recorded from Paropsisterna nobilitata (Erichson) for the first time. Most Australian members of Chrysomelobia are host-specific and beetles may host up to three mite species. Within species-groups of mites, species that share the same host genus are closely related. However, mite species that share the same host species are usually from different species-groups of mites. Although these patterns suggest co-speciation at terminal nodes and host transfers at deeper nodes, little can be made of these patterns without a host phylogeny. Keys to Chrysomelobia adults are provided.


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