Platypodidae under scrutiny

2000 ◽  
Vol 14 (6) ◽  
pp. 771 ◽  
Author(s):  
Guillermo Kuschel ◽  
Richard A. B. Leschen ◽  
Elwood C. Zimmerman

The historical status of the family Platypodidae is reviewed and the family is revised. Results of a cladistic analysis based on 35 terminal taxa and 80 adult morphological characters show that the current placement of Platypodidae makes the subfamily Scolytinae paraphyletic. Moreover, several important genera included in Scolytinae are shown to be members of Cossoninae (i.e. the placement of Protoplatypus Wood and Phylloplatypus Kato in Cossoninae is confirmed). Based on these results, the status of Platypodidae as a family and subfamily is rejected, Scolytinae thereby becoming a monophyletic taxon. Araucarius groups in Scolytinae instead of Cossoninae in the analysis on a single step only, but it is suggested that it be retained in Cossoninae until this subfamily is submitted to a similar phylogenetic study. Three genera and four species of Cossoninae are described as new: Dobionus Kuschel, gen. nov.: type species D. araucarinus Kuschel, sp. nov. (with the inclusion of D. brachyrhinus (Montrouzier)); Coptonus Kuschel, gen. nov.: type species C. fijianus Kuschel, sp. nov. (with the inclusion of C. papuanus Kuschel, sp. nov.) and Dissostomus Kuschel, gen. nov.: type species D. hornabrooki Kuschel, sp. nov.

1991 ◽  
Vol 123 (S160) ◽  
pp. 3-341 ◽  
Author(s):  
Jean-François Landry

AbstractGenera and previously described species of Nearctic Scythrididae are revised for the first time, based on the study of adult structures. About 90 percent of the Nearctic fauna known in collections consists of undescribed species. The supraspecific taxa treated in this work encompass less than half of the Nearctic species diversity. Only six new species are described, all within the largest and structurally most diverse genus. The status of all nominal species is revised. Valid species are redescribed and their features illustrated. General problems in the systematics of the Scythrididae are discussed. A description of adult features of the family Scythrididae is providad. Extra-limital genera are briefly reviewed. A key to the Nearctic genera and informal supraspecific lineages is provided.Six genera, including three new, are treated: Areniscythris Powell, 1976, Arotrura Walsingham, 1888, Asymmetrura gen. nov., Neoscythris gen. nov., Rhamphura gen. nov., and Scythris s. str. Hübner, [1825]. Areniscythris includes a single described species, Areniscythris brachypteris Powell, but is defined more broadly to account for a number of undescribed species. Arotrura is divided into nine informal species groups with the following included species: Arotrura atascosa sp. nov., Arotrura balli sp. nov., Arotrura divaricata (Braun) comb, nov., Arotrura eburnea Walsingham, Arotrura formidabilis sp. nov., Arotrura hymenata sp. nov., Arotrura longissima sp. nov., Arotrura oxyplecta (Meyrick) comb, nov., Arotrura powelli sp. nov., and Arotrura sponsella (Busck) comb. nov. Asymmetrura includes: Asymmetrura albilineata (Walsingham) comb. nov., Asymmetrura graminivorella (Braun) comb. nov., Asymmetrura impositella (Zeller) comb. nov. and type species, Asymmetrura matutella (Clemens) comb, nov., Asymmetrura reducta (Braun) comb, nov., and Asymmetrura scintillifera (Braun) comb. nov. Neoscythris includes: Neoscythris confinis (Braun) comb, nov., Neoscythris euthia (Walsingham) comb. nov., Neoscythris fissirostris (Meyrick) comb. nov. and type species, and Neoscythris planipenella (Chambers) comb. nov. Rhamphura includes: Rhamphura altisierrae (Keifer) comb, nov., Rhamphura ochristriata (Walsingham) comb. nov. and type species, Rhamphura perspicillella (Walsingham) comb. nov., Rhamphura suffusa (Walsingham) comb. nov., and the extra-limital Rhamphura immunis (Meyrick) comb. nov. from Peru. Scythris s. str. includes: Scythris immaculatella (Chambers) rev. stat., Scythris limbella (Fabricius), Scythris mixaula Meyrick, Scythris trivinctella (Zeller), and Scythris ypsilon Braun. A further eight species are phylogenetically distinct from Scythris s. str. but provisionally are only assigned to five informal monophyletic lineages until their cladistic relationships are more firmly established. These are: the Scythris basilaris lineage including Scythris basilaris (Zeller), Scythris eboracensis (Zeller), and Scythris fuscicomella (Clemens); the Scythris interrupta lineage including Scythris interrupta Braun; the Scythris inspersella lineage including Scythris inspersella (Hübner) and Scythris noricella (Zeller); the Scythris anthracina lineage including Scythris anthracina Braun; and the Scythris charon lineage including Scythris charon Meyrick. Three species are incertae sedis: Scythris inornatella (Chambers) comb, nov., Scythrispilosella (Zeller), and Scythris piratica Meyrick.Coleophora albacostella Chambers and Coleophora inornatella Chambers are transferred from the Coleophoridae. Scythris arizoniella (Kearfott) is transferred to the Coleophoridae [Coleophora arizoniella (Kearfott) comb. nov.].The following new synonymy is proposed: Colinita Busck, 1907 = Arotrura Walsingham, 1888; Gelechia aterrimella Walker, 1864 and Scythris epilobiella McDunnough, 1942 = Scythris inspersella [Hübner, (1817)]; Scythris magnatella Busck, 1904 = Scythris noricella (Zeller, 1843); Scythris pacifica McDunnough, 1927 = Scythris immaculatella (Chambers, 1875); Coleophora albacostella Chambers, 1875 and Scythris hemidictyas Meyrick, 1928 = Neoscythris planipenella (Chambers, 1875).A cladistic definition of the family is presented for the first time. The monophyly of the Scythrididae is supported by the following synapomorphies: very narrow ductus bursae, broad ductus seminalis anastomosed with the oviduct and the corpus bursae, lack of signum, unique shape of the apophyses of the metathoracic furca, tarsomeres 1–4 with two subapical spurs, aedeagus ankylosed, and origin of forewing veins R4 and R5 on a common stalk with R4 extended to the costa and R5 to the termen. Relationships of the Scythrididae within the Gelechioidea are discussed. Based on the cladistic analysis of 52 structural characters, phylogenetic relationships of supraspecific taxa are inferred. Two cladograms, one for the genera and one for the species groups of Arotrura, are presented and used in deriving the classification.


2011 ◽  
Vol 20 (1) ◽  
pp. 161-173
Author(s):  
A.P. Kassatkina

Resuming published and own data, a revision of classification of Chaetognatha is presented. The family Sagittidae Claus & Grobben, 1905 is given a rank of subclass, Sagittiones, characterised, in particular, by the presence of two pairs of sac-like gelatinous structures or two pairs of fins. Besides the order Aphragmophora Tokioka, 1965, it contains the new order Biphragmosagittiformes ord. nov., which is a unique group of Chaetognatha with an unusual combination of morphological characters: the transverse muscles present in both the trunk and the tail sections of the body; the seminal vesicles simple, without internal complex compartments; the presence of two pairs of lateral fins. The only family assigned to the new order, Biphragmosagittidae fam. nov., contains two genera. Diagnoses of the two new genera, Biphragmosagitta gen. nov. (type species B. tarasovi sp. nov. and B. angusticephala sp. nov.) and Biphragmofastigata gen. nov. (type species B. fastigata sp. nov.), detailed descriptions and pictures of the three new species are presented.


Zootaxa ◽  
2010 ◽  
Vol 2408 (1) ◽  
pp. 1 ◽  
Author(s):  
C. DELAND ◽  
C. B. CAMERON ◽  
K. P. RAO ◽  
W. E. RITTER ◽  
T. H. BULLOCK

The family Harrimaniidae (Hemichordata: Enteropneusta) is revised on the basis of morphological characters. The number of harrimaniid genera is increased to nine by the addition of Horstia n. gen., Mesoglossus n. gen., Ritteria n. gen. and Saxipendium, a genus previously assigned to the monospecific family Saxipendiidae. The number of species is increased to 34, resulting from the description of five new species from the eastern Pacific — Horstia kincaidi, Mesoglossus intermedius, M. macginitiei, Protoglossus mackiei and Ritteria ambigua. A description is supplied for a sixth harrimaniid species, Stereobalanus willeyi Ritter & Davis, 1904, which previously had the status of a nomen nudum. Four harrimaniids previously assigned to the genus Saccoglossus are transfered to the genus Mesoglossus — M. bournei, M. caraibicus, M. gurneyi and M. pygmaeus, while Saccoglossus borealis is reassigned to the genus Harrimania. Notes on habitat and zoogeography are included for the seven foregoing species and a table of diagnostic characters for existing and new species and a dichotomous key to the enteropneust families and harrimaniid genera are provided. Finally, a phylogenetic hypothesis concerning the Harrimaniidae is postulated, with discussion on the evolution of the group.


1997 ◽  
Vol 11 (1) ◽  
pp. 1 ◽  
Author(s):  
A. D. Austin ◽  
and S. A. Field

The morphology of the sclerotised components of the ovipositor system is comprehensively surveyed for scelionid and platygastrid wasps, with information being assessed for 120 genera and 220 species. A diagnosis for the ovipositor system is presented for most genera to complement existing generic descriptions. Two previously described and mechanically different forms of the ovipositor system are recognised: (1) the Ceratobaeus-type that is extended and retracted by antagonistic muscles and (2) the Scelio-type that is operated by changes in hydrostatic pressure, where the ovipositor is extended at the end of an elongate telescopic tube derived from expanded intersegmental membrane between metasomal segments 6 and 7. Comparison of these forms with the supposed ground plan for the Scelionidae strongly indicates that the Scelio-type is apomorphic, that it defines a monophyletic group associated with orthopteran host eggs, and that it comprises the tribes Scelionini, Calliscelionini, most Psilanteridini, Aradophagini, Neoscelionini, Platyscelionini, Doddiellini and four genera misplaced within the Sparasionini and Baryconini (Archaeoteleia Masner, Bracalba Dodd, Chromoteleia Ashmead and Oxyscelio Kieffer), as well as Sceliacanthella Dodd. Until a more robust classification of the superfamily is forthcoming, it is proposed that this group be informally referred to as the 'Scelionini sensu lato'. Further, seven genera (Habroteleia Kieffer, Palpoteleia Kieffer, Anteris Foerster, Fusicornia Risbec, Leptoteleia Kieffer, Opisthacantha Ashmead and Styloteleia Kieffer) are misplaced in the Calliscelionini and Psilanteridini because they possess the Ceratobaeus-type system. Nixonia Masner, Sparasion Latreille and Sceliomorpha Ashmead (Sparasionini) are considered to have the most primitive ovipositor system because they possess a Ceratobaeus-type system, and sub-basally fused lateral and latero-ventral apodemes, the latter being loosely attached to sternite 6. Sparasion and Sceliomorpha also have very short lateral apodemes and this, in conjunction with the form of the apodemes, can be considered to be the ground plan for the superfamily. The Platygastridae all possess a modified ovipositor system but, nonetheless, one that in most cases is extended and retracted by musculature (i.e. Ceratobaeus-type). In particular, the system in most platygastrids is typified by having metasomal tergite 8 and associated cerci missing, the lateral apodemes short and forming a U-shape, and the ovipositor assembly generally robust. Only one of approximately 30 genera examined, Acerotella Masner, has very elongate apodemes, as in the Scelionidae. Many platygastrids also have a pair of latero-ventral apodemes, a presumed plesiomorphic character, rather than a single medial apodeme on stemite 6, which is the case for many Scelionidae. The most highly modified system is found in Isostasius Foerster and some Synopeas (Sactogaster) Foerster, where the ovipositor assembly is coiled vertically or partly so and the apodemes are greatly reduced. Generally, characters associated with the ovipositor system do not provide any independent support for the most recent higher-level classification of platygastrids, although they show substantial potential for more accurate definition of genera. A preliminary cladistic analysis of 14 ovipositor characters supports the monophyly of five clades that correspond to the Scelionini s. l., the Scelionidae (minus the Sparasionini sensu stricto), the Sparasionini s. str., the Platygastridae, and the Sparasioriini s. str. + Platygastridae. Overall, results from this study will provide baseline information on the ovipositor system as a prelude to a more complete phylogenetic analysis of the superfamily including external morphological characters. Although no new classification for the Scelionidae and Platygastridae is proposed, their higher-level taxonomy is reviewed and discussed and cases identified where, on the basis of ovipositor morphology, taxa (tribes and/or genera) apparently form monophyletic groups, and where taxa are misplaced. Finally, the status of the major higher-level groups within the superfamily is discussed, as is the available evidence to support their monophyly.


2020 ◽  
Vol 86 (1) ◽  
pp. 1-26
Author(s):  
S T Williams ◽  
Y Kano ◽  
A Warén ◽  
D G Herbert

ABSTRACT The assignment of species to the vetigastropod genus Solariella Wood, 1842, and therefore the family Solariellidae Powell, 1951, is complicated by the fact that the type species (Solariella maculata Wood, 1842) is a fossil described from the Upper Pliocene. Assignment of species to genera has proved difficult in the past, and the type genus has sometimes acted as a ‘wastebasket’ for species that cannot easily be referred to another genus. In the light of a new systematic framework provided by two recent publications presenting the first molecular phylogenetic data for the group, we reassess the shell characters that are most useful for delimiting genera. Shell characters were previously thought to be of limited taxonomic value above the species level, but this is far from the case. Although overall shell shape is not a reliable character, our work shows that shell characters, along with radular and anatomical characters, are useful for assigning species to genera. Sculpture of the early teleoconch (the region immediately following the protoconch) and the columella are particularly useful characters that have not been used regularly in the past to distinguish genera. However, even with the combination of all morphological characters used in this study (shell, radular and eye), a few species are still difficult to assign to genera and in such cases molecular systematic data are essential. In the present study, we discuss 13 genera—12 of which were recovered as well-supported clades in recent molecular systematic studies—and provide morphological characters to distinguish them. We describe several new taxa: Chonospeira n. gen. (referred to as ‘clade B’ in previous molecular systematic studies), Phragmomphalina n. gen. (Bathymophila in part in molecular systematic studies) and Phragmomphalina vilvensi n. sp. (type species of Phragmomphalina n. gen.). We synonymize Hazuregyra Shikama, 1962 with Minolia A. Adams, 1860, Minolia subangulata Kuroda & Habe, 1952 with Minolia punctata A. Adams, 1860 and M. gemmulata Kuroda & Habe, 1971 with M. shimajiriensis (MacNeil, 1960). We also present the following new combinations: Bathymophila bairdii (Dall, 1889), B. dawsoni (Marshall, 1979), B. regalis (Marshall, 1999), B. wanganellica (Marshall, 1999), B. ziczac (Kuroda & Habe in Kuroda, Habe & Oyama, 1971), Chonospeira nuda (Dall, 1896), C. iridescens (Habe, 1961), C. ostreion (Vilvens, 2009), C. strobilos (Vilvens, 2009), Elaphriella corona (Lee & Wu, 2001), E. diplax (Marshall, 1999), E. meridiana (Marshall, 1999), E. olivaceostrigata (Schepman, 1908), E. opalina (Shikama & Hayashi, 1977), Ilanga norfolkensis (Marshall, 1999), I. ptykte (Vilvens, 2009), I. zaccaloides (Vilvens, 2009), Minolia shimajiriensis (MacNeil, 1960), M. watanabei (Shikama, 1962), Phragmomphalina alabida (Marshall, 1979), P. diadema (Marshall, 1999), P. tenuiseptum (Marshall, 1999), Spectamen euteium (Vilvens, 2009), S. basilicum (Marshall, 1999), S. exiguum (Marshall, 1999) and S. flavidum (Marshall, 1999).


1999 ◽  
Vol 73 (1) ◽  
pp. 159-160 ◽  
Author(s):  
I. G. Sohn ◽  
F. M. Swain

Molostovskaja (1979, p. 54) described without illustrations the family Darwinuloididae Molostovskaja, 1979. The following year, Molostovskaja (1980, p. 33) described the family again as new [Darwinuloididae fam. nov.]. She included the genus Whipplella Holland, 1934, in the family and illustrated Darwinuloides svijazhicus (Sharapova 1948) (Molostovskaja, 1980, p. 28, figs. 5a, b, 8). Schneider (1948, p. 29, pl. 2, figs, 1a, b) originally described and illustrated this species as “Darwinula svijazhicus Sharapova n. mns.” Two years later, Molostovskaja (1982, p. 158) illustrated cross-sections of the adductor muscle attachment scar patterns in the families of the Darwinulacea Brady and Norman, 1889. Later Molostovskaja (1990, p. 166) erected the superfamily Darwinuloidacea Molostovskaja, 1979 to include Whipplella Holland, 1934, Vymella Kalis and Mischina in Mischina and Kalis, 1975, and questionably Pruvostina Scott and Summerson, 1943. In the same paper, she illustrated Whipplella sp. (pl. 72, figs. 6a, b) [from Sohn, 1977], Vymella dobrinini (Kashevarova, 1961) (pl. 72, figs. 7a-c), Darwinuloides sentjakensis (Sharapova in Schneider, 1948) (pl. 73, figs. 5, 6), and D. svijazhicus (Sharapova in Schneider, 1948) (pl. 74, figs. 14a, b).


1990 ◽  
Vol 68 (12) ◽  
pp. 2649-2687 ◽  
Author(s):  
F. Rafi ◽  
Diana R. Laubitz

The distribution of the northeastern North Pacific Idoteidae reflects the general eurytopy in the shallow marine environment of the component species. The family is represented by four genera and 20 species in the region between Juan de Fuca Strait, Washington, and Prince William Sound, Alaska. Collections studied from this area contained 16 species of idoteids, of which 3 (Idotea (Pentidotea) recta, Synidotea cornuta, and S. minuta) are new; the new species are fully illustrated and described. The status of nine additional species recorded in the literature from the area is discussed, and four of them are diagnosed and included in the keys. The morphological characters of the four genera were examined in detail and their systematic importance is discussed. All the appendages surveyed showed differences at least at the generic level.


1998 ◽  
Vol 72 (4) ◽  
pp. 698-718 ◽  
Author(s):  
Jonathan M. Adrain

Cladistic analysis of the trilobite subfamily Acanthoparyphinae Whittington and Evitt, 1954, yields an explicit hypothesis of relationship for the group. All Silurian species together form a robustly supported monophylum including the genera Hyrokybe Lane, 1972, Parayoungia Chatterton and Perry, 1984, and Youngia Lindström, 1885. Sister to this is the Ordovician type species of Acanthoparypha Whittington and Evitt, 1954. Remaining species that have historically been assigned to either Acanthoparypha or Pandaspinapyga Esker and Levin, 1964, form a rather labile paraphylum. Nevertheless, the entire group thus identified is definitely monophyletic, and supported by several prominent synapomorphic character-states.The basal structure and basal node of the subfamily are more difficult to assess. The relationships of the genera Hammannopyge Přibyl, Vaněk, and Pek, 1985, Holia Bradley, 1930, and Nieszkowskia Schmidt, 1881, need to be addressed within the wider context of the family as a whole. The traditional assignment of Holia to the acanthoparyphines is followed.Wenlock acanthoparyphines from the Cape Phillips Formation of the central Canadian Arctic islands include several species of Hyrokybe and Parayoungia. They are similar to, and in one case conspecific with, coeval forms to the southwest in the southern Mackenzie Mountains.Five species are new: Holia glabra, Hyrokybe lightfooti, Hyrokybe youngi, Hyrokybe mitchellae, and Parayoungia mclaughlini. At least four other potentially new species are reported in open nomenclature.


2002 ◽  
Vol 16 (6) ◽  
pp. 957 ◽  
Author(s):  
J. T. Jennings ◽  
A. D. Austin

This study examines the phylogeny, taxonomy, distribution and biology of the gasteruptiid subfamily Hyptiogastrinae and, at the same time, presents an overview of the family. Following a cladistic analysis of 35 discrete morphological characters, two monophyletic genera are recognised, Hyptiogaster Kieffer and Pseudofoenus s. l. Kieffer. As a consequence, the genera Aulacofoenus Kieffer, Crassifoenus Crosskey, and Eufoenus Szépligeti are synonymised with Pseudofoenus. A total of 88 species are recognised for the subfamily, 10 species of Hyptiogaster, which are restricted to mainland Australia, and 78 species of Pseudofoenus, 40 of which are described as new. Pseudofoenus has a restricted Gondwanan distribution and is found in Australia including Tasmania (65 spp.), New Guinea and New Britain (5 spp.), the south-west Pacific (New Caledonia, New Hebrides and Fiji – 2 spp.), New Zealand (4 spp.) and South America (2 spp.). No new species have been recorded from either New Zealand or South America. For Pseudofoenus, information on the distribution of each species, their biology (if known) and an identification key are presented.Following a taxonomic revision, the following new species are described: P. baileyi, sp. nov., P. baitetaensis, sp. nov., P. beverlyae, sp. nov., P. caperatus, sp. nov., P. cardaleae, sp. nov., P. carrabinensis, sp. nov., P. claireae, sp. nov., P. collessi, sp. nov., P. coorowensis, sp. nov., P. crosskeyi, sp. nov., P. douglasorum, sp. nov., P. eliseae, sp. nov., P. ericae, sp. nov., P. eustonensis, sp. nov., P. feckneri, sp. nov., P. gressitti, sp. nov., P. gullanae, sp. nov., P. hackeri, sp. nov., P. imbricatus, sp. nov., P. iqbali, sp. nov., P. kadowi, sp. nov., P. karimuiensis, sp. nov., P. kelleri, sp. nov., P. leinsterensis, sp. nov., P. macdonaldi, sp. nov., P. malkini, sp. nov., P. marshalli, sp. nov., P. masneri, sp. nov., P. mitchellae, sp. nov., P. morganensis, sp. nov., P. nalbarraensis, sp. nov., P. pumilis, sp. nov., P. schmidti, sp. nov., P. stevensi, sp. nov., P. tasmaniensis, sp. nov., P. taylori, sp. nov., P. umboiensis, sp. nov., P. walkeri, sp. nov. and P. zborowskii, sp. nov. The synonymy of Aulacofoenus, Crassifoenus and Eufoenus with Pseudofoenus result in the following new combinations: from Aulacofoenus: P. bungeyi (Jennings & Austin), comb. nov., P. deletangi (Schletterer), comb. nov., P. fallax (Schletterer), comb. nov., P. fletcheri (Jennings & Austin), comb. nov., P. goonooensis (Jennings & Austin), comb. nov., P. infumatus (Schletterer), comb. nov., P. kurmondi (Jennings & Austin), comb. nov., P. loxleyi (Jennings & Austin), comb. nov., P. marionae (Jennings & Austin), comb. nov., P. perenjorii (Jennings & Austin), comb. nov., P. swani (Jennings & Austin), comb. nov., P. thoracicus (Guérin Menéville), comb. nov., P. whiani (Jennings & Austin), comb. nov. and P. wubinensis (Jennings & Austin), comb. nov.; from Crassifoenus: P. houstoni (Jennings & Austin), comb. nov., P. grossitarsis (Kieffer), comb. nov and P. macronyx (Schletterer), comb. nov.; and from Eufoenus: P. antennalis (Schletterer), comb. nov., P. australis (Westwood), comb. nov., P. crassitarsis (Kieffer), comb. nov., P. darwini (Westwood), comb. nov., P. extraneus (Turner), comb. nov., P. ferrugineus (Crosskey), comb. nov., P. floricolus (Turner), comb. nov., P. inaequalis (Turner), comb. nov., P. melanopleurus (Crosskey), comb. nov., P. minimus (Turner), comb. nov., P. nitidiusculus (Turner), comb. nov., P. patellatus (Westwood), comb. nov., P. pilosus (Kieffer), comb. nov., P. reticulatus (Crosskey), comb. nov., P. rieki (Crosskey), comb. nov., P. ritae (Cheesman), comb. nov. and P. spinitarsis (Westwood), comb. nov. Pseudofoenus microcephalus (Crosskey), comb. nov. is transferred from Hyptiogaster and Eufoenus flavinervis (Kieffer) remains incertae sedis.


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