scholarly journals Horseshoe crab trace fossils from the Upper Cretaceous Two Medicine Formation of Montana, USA, and a brief review of the xiphosurid ichnological record

2020 ◽  
Vol 94 (5) ◽  
pp. 887-905
Author(s):  
Masateru Shibata ◽  
David J. Varricchio
Keyword(s):  
Fossil Record ◽  
Upper Cretaceous ◽  
Horseshoe Crab ◽  
Taxonomic Diversity ◽  
Late Triassic ◽  
Upper Carboniferous ◽  
Two Factors ◽  
Horseshoe Crabs ◽  
Freshwater Habitats ◽  
First Occurrence

AbstractA locality in the Upper Cretaceous Two Medicine Formation of Montana preserves abundant and variable horseshoe crab tracks and trails of the ichnotaxon Kouphichnium isp. These specimens span six morphologies differing in track form and trail configuration. These differences likely reflect variations in track-maker locomotion and behavior, substrate consistency, epichnial versus hypichnial preservation, and undertrack versus true tracks. Several tracks preserve the first clear appendage impressions for an extinct horseshoe crab. This discovery adds new information to the fossil horseshoe crab diversity in the Cretaceous Period. Trackway dimensions, such as the external width across the pusher legs or of the prosomal drag mark, provide information on the track-maker size. Most trackways correspond with crabs 9–14 cm wide; the abundance but limited size range of the traces suggests the large assemblage corresponds to a mating aggregation. The trace fossil record of xiphosurids indicates that throughout their history, horseshoe crabs inhabited both marine and nonmarine settings. They were definitively present in freshwater habitats from the lower Carboniferous through at least the Paleogene. Horseshoe crab trace abundance is highest from the upper Carboniferous through the Jurassic and likely reflects two factors: true upper Carboniferous taxonomic diversity and a preponderance of suitable sites for trackway preservation in the Late Triassic and Jurassic. Cretaceous traces are uncommon, and this Two Medicine locality is the first occurrence of horseshoe crab traces in the Late Cretaceous worldwide. Overall, track abundance and diversity would seem to correspond well with the reported horseshoe crab body fossil record.

Taxonomic diversity estimation using rarefaction

Paleobiology ◽  
1975 ◽  
Vol 1 (4) ◽  
pp. 333-342 ◽  
Author(s):  
David M. Raup
Keyword(s):  
Species Richness ◽  
Species Diversity ◽  
Sample Size ◽  
Fossil Record ◽  
World Wide ◽  
Taxonomic Diversity ◽  
Early Jurassic ◽  
Late Triassic ◽  
Geologic Time ◽  
Large Samples

Benthic ecologists have successfully applied rarefaction techniques to the problem of compensating for the effect of sample size on apparent species diversity (= species richness). The same method can be used in studies of diversity at higher taxonomic levels (families and orders) in the fossil record where samples represent world-wide distributions of species or genera over long periods of geologic time.Application of rarefaction to several large samples of post-Paleozoic echinoids (totaling 7,911 species) confirms the utility of the method. Rarefaction shows that the observed increase in the number of echinoid families since the Paleozoic is real in the sense that it cannot be explained solely by the increase in numbers of preserved species. There has been no statistically significant increase in the number of families since mid-Cretaceous, however. At the order level, echinoid diversity may have been nearly constant since late Triassic or early Jurassic.

scholarly journals Gladius-bearing coleoids from the Upper Cretaceous Lebanese Lagerstätten: diversity, morphology, and phylogenetic implications

2015 ◽  
Vol 89 (1) ◽  
pp. 148-167 ◽  
Author(s):  
Romain Jattiot ◽  
Arnaud Brayard ◽  
Emmanuel Fara ◽  
Sylvain Charbonnier
Keyword(s):  
New Species ◽  
Fossil Record ◽  
Upper Cretaceous ◽  
Soft Part ◽  
Taxonomic Diversity ◽  
Phylogenetic Implications ◽  
Cretaceous Period ◽  
First Time ◽  
One New Species

AbstractGladius-bearing coleoids are rare in the fossil record. For the Cretaceous period, these cephalopods are mainly recorded in a few Lagerstätten in Lebanon (Haqel, Hajoula, En Nammoura, and Sahel Aalma). Here, we study 16 specimens of gladius-bearing coleoids from these Upper Cretaceous Lebanese Lagerstätten to investigate their taxonomic diversity. Besides two species that were already reported (Dorateuthis syriacaandGlyphiteuthis libanotica), one new species is identified in the Cenomanian site of Hajoula:Rachiteuthis acutalin. sp., as well as another form ofGlyphiteuthisfrom En Nammoura. Several studied specimens exhibit well-preserved soft-part characters. Among them, we document for the first time two transverse rows of sessile suckers inD.syriacaand we confirm the absence of tentacles, as well as the presence of a crop in this species. This strongly supports the phylogenetic proximity ofD.syriacawith modern vampyropods rather than with modern decabrachians. In turn, the similarity in gladius morphology between this taxon and modern squids is regarded as convergent.

scholarly journals Late Cretaceous domatia reveal the antiquity of plant–mite mutualisms in flowering plants

2019 ◽  
Vol 15 (11) ◽  
pp. 20190657 ◽  
Author(s):  
S. Augusta Maccracken ◽  
Ian M. Miller ◽  
Conrad C. Labandeira
Keyword(s):  
Late Cretaceous ◽  
Fossil Record ◽  
Evolutionary History ◽  
Upper Cretaceous ◽  
Flowering Plants ◽  
Predaceous Mites ◽  
First Occurrence ◽  
History Of ◽  
Living Species ◽  
Cenozoic Era

Mite houses, or acarodomatia, are found on the leaves of over 2000 living species of flowering plants today. These structures facilitate tri-trophic interactions between the host plant, its fungi or herbivore adversaries, and fungivorous or predaceous mites by providing shelter for the mite consumers. Previously, the oldest acarodomatia were described on a Cenozoic Era fossil leaf dating to 49 Myr in age. Here, we report the first occurrence of Mesozoic Era acarodomatia in the fossil record from leaves discovered in the Upper Cretaceous Kaiparowits Formation (76.6–74.5 Ma) in southern UT, USA. This discovery extends the origin of acarodomatia by greater than 25 Myr, and the antiquity of this plant–mite mutualism provides important constraints for the evolutionary history of acarodomatia on angiosperms.

scholarly journals Xiphosurid from the Tournaisian (Carboniferous) of Scotland confirms deep origin of Limuloidea

2019 ◽  
Vol 9 (1) ◽  
Author(s):  
Russell D. C. Bicknell ◽  
Stephen Pates
Keyword(s):  
Fossil Record ◽  
Evolutionary History ◽  
Horseshoe Crab ◽  
New Taxon ◽  
Geometric Morphometric ◽  
Lower Carboniferous ◽  
Morphometric Analyses ◽  
Lower Ordovician ◽  
Horseshoe Crabs ◽  
History Of

AbstractHorseshoe crabs are archetypal marine chelicerates with a fossil record extending from the Lower Ordovician to today. The major horseshoe crab groups are thought to have arisen in the middle to late Palaeozoic. Here we present the oldest known limuloid from the lower Carboniferous (Tournaisian stage, c. 350 million years ago) of Scotland: Albalimulus bottoni gen. et sp. nov. A comprehensive phylogenetic analysis supports the placement of A. bottoni as a representative of the extant family Limulidae and 100 million years older than any other limulid taxon. The use of geometric morphometric analyses corroborate the erection of the new taxon and illustrates the exploitation of morphospace by xiphosurids. This new taxon highlights the complex evolutionary history of xiphosurids and the importance of documenting these unique Palaeozoic individuals.

SIZE FREQUENCY DISTRIBUTION BIASES IN HORSESHOE CRAB MOLT ASSEMBLAGES: IMPLICATIONS FOR THE FOSSIL RECORD

2016 ◽  
Author(s):  
Shelly J. Wernette ◽  
◽  
Scott Evans ◽  
Christine Hall ◽  
Mary L. Droser ◽  
...  
Keyword(s):  
Frequency Distribution ◽  
Fossil Record ◽  
Horseshoe Crab ◽  
Size Frequency Distribution ◽  
Size Frequency

The nasal cavity of two traversodontid cynodonts (Eucynodontia, Gomphodontia) from the Upper Triassic of Brazil

2021 ◽  
pp. 1-16
Author(s):  
Arymathéia Santos Franco ◽  
Rodrigo Temp Müller ◽  
Agustín G. Martinelli ◽  
Carolina A. Hoffmann ◽  
Leonardo Kerber
Keyword(s):  
Nasal Cavity ◽  
Fossil Record ◽  
Nasal Septum ◽  
Internal Surface ◽  
Nasolacrimal Duct ◽  
Upper Triassic ◽  
Late Triassic ◽  
Southern Brazil ◽  
Dorsal Region ◽  
Left And Right

Abstract Traversodontidae is a group of Triassic herbivorous/omnivorous cynodonts that represents the most diversified lineage within Cynognathia. In southern Brazil, a rich fossil record of late Middle/mid-Late Triassic cynodonts has been documented, with Exaeretodon riograndensis Abdala, Barberena, and Dornelles, 2002 and Siriusgnathus niemeyerorum Pavanatto et al., 2018 representing two abundant and well-documented traversodontids. The present study provides a comparative analysis of the morphology of the nasal cavity, nasal recesses, nasolacrimal duct, and maxillary canals of both species using computed tomography, highlighting the changes that occurred in parallel to the origin of mammaliaforms. Our results show that there were no ossified turbinals or a cribriform plate delimiting the posterior end of the nasal cavity, suggesting these structures were probably cartilaginous as in nonmammaliaform cynodonts. Both species show lateral ridges on the internal surface of the roof of the nasal cavity, but the median ridge for the attachment of a nasal septum is absent. Exaeretodon riograndensis and S. niemeyerorum show recesses on the dorsal region of the nasal cavity, which increase the volume of the nasal cavity, potentially enhancing the olfactory chamber and contributing to the sense of smell. On the lateral sides of the nasal cavity, the analyzed taxa show a well-developed maxillary recess. Although E. riograndensis and S. niemeyerorum have roughly similar nasal cavities, in the former taxon, the space between the left and right dorsal recesses of the nasal cavity is uniform along its entire extension, whereas this space narrows posteriorly in S. niemeyerorum. Finally, the nasolacrimal duct of S. niemeyerorum is more inclined anteroposteriorly than in E. riograndensis.

Anatomical, morphometric, and stratigraphic analyses of theropod biodiversity in the Upper Cretaceous (Campanian) Dinosaur Park Formation

2021 ◽  
pp. 1-15
Author(s):  
Thomas M. Cullen ◽  
Lindsay Zanno ◽  
Derek W. Larson ◽  
Erinn Todd ◽  
Philip J. Currie ◽  
...  
Keyword(s):  
High Resolution ◽  
North America ◽  
Fossil Record ◽  
Upper Cretaceous ◽  
Environmental Changes ◽  
Morphometric Analyses ◽  
Variable Distribution ◽  
Taxic Diversity ◽  
Dinosaur Park Formation ◽  
Broad Scale

The Dinosaur Park Formation (DPF) of Alberta, Canada, has produced one of the most diverse dinosaur faunas, with the record favouring large-bodied taxa, in terms of number and completeness of skeletons. Although small theropods are well documented in the assemblage, taxonomic assessments are frequently based on isolated, fragmentary skeletal elements. Here we reassess DPF theropod biodiversity using morphological comparisons, high-resolution biostratigraphy, and morphometric analyses, with a focus on specimens/taxa originally described from isolated material. In addition to clarifying taxic diversity, we test whether DPF theropods preserve faunal zonation/turnover patterns similar to those previously documented for megaherbivores. Frontal bones referred to a therizinosaur (cf. Erlikosaurus), representing among the only skeletal record of the group from the Campanian–Maastrichtian (83–66 Ma) fossil record of North America, plot most closely to troodontids in morphospace, distinct from non-DPF therizinosaurs, a placement supported by a suite of troodontid anatomical frontal characters. Postcranial material referred to cf. Erlikosaurus in North America is also reviewed and found most similar in morphology to caenagnathids, rather than therizinosaurs. Among troodontids, we document considerable morphospace and biostratigraphic overlap between Stenonychosaurus and the recently described Latenivenatrix, as well as a variable distribution of putatively autapomorphic characters, calling the validity of the latter taxon into question. Biostratigraphically, there are no broad-scale patterns of faunal zonation similar to those previously documented in ornithischians from the DPF, with many theropods ranging throughout much of the formation and overlapping extensively, possibly reflecting a lack of sensitivity to environmental changes, or other cryptic ecological or evolutionary factors.

scholarly journals First rebbachisaurid sauropod dinosaur from Asia

PLoS ONE ◽  
2021 ◽  
Vol 16 (2) ◽  
pp. e0246620
Author(s):  
Alexander Averianov ◽  
Hans-Dieter Sues
Keyword(s):  
Phylogenetic Analysis ◽  
Fossil Record ◽  
Upper Cretaceous ◽  
Neural Spine ◽  
Lateral Side ◽  
New Taxon ◽  
Caudal Vertebra

Dzharatitanis kingi gen. et sp. nov. is based on an isolated anterior caudal vertebra (USNM 538127) from the Upper Cretaceous (Turonian) Bissekty Formation at Dzharakuduk, Uzbekistan. Phylogenetic analysis places the new taxon within the diplodocoid clade Rebbachisauridae. This is the first rebbachisaurid reported from Asia and one of the youngest rebbachisaurids in the known fossil record. The caudal is characterized by a slightly opisthocoelous centrum, ‘wing-like’ transverse processes with large but shallow PRCDF and POCDF, and the absence of a hyposphenal ridge and of TPRL and TPOL. The neural spine has high SPRL, SPDL, SPOL, and POSL and is pneumatized. The apex of neural spine is transversely expanded and bears triangular lateral processes. The new taxon shares with Demandasaurus and the Wessex rebbachisaurid a high SPDL on the lateral side of the neural spine, separated from SPRL and SPOL. This possibly suggests derivation of Dzharatitanis from European rebbachisaurids. This is the second sauropod group identified in the assemblage of non-avian dinosaurs from the Bissekty Formation, in addition to a previously identified indeterminate titanosaurian.

scholarly journals The first occurrence of the enigmatic archosaur Crosbysaurus (Heckert 2004) from the Chinle Formation of Southern Utah

2014 ◽  
Author(s):  
Robert J Gay ◽  
Isabella St. Aude
Keyword(s):  
New Mexico ◽  
Close Association ◽  
First Record ◽  
Geographic Range ◽  
Type Locality ◽  
Late Triassic ◽  
Southern Area ◽  
Chinle Formation ◽  
First Occurrence ◽  
Unique Compound

Originally identified as an ornithisichian dinosaur, Crosbysaurus has been found in New Mexico, Arizona, and the type locality in Texas. The genus has been reassessed by other workers in light of revelations about the postcrania of another putative Triassic ornithischian, Revueltosaurus. The understanding of Triassic dental faunas has become more complicated by the extreme convergence between pseudosuchian archosaurus and ornithichian dinosaur dental morphologies. We report here on a new specimen of Crosbysaurus from the Petrified Forest Member of the Chinle Formation at Comb Ridge in southeastern Utah. This new specimen is assigned to Crosbysaurus on the basis of the unique compound posterior denticles, mediolateral width, and curvature. While this specimen, MNA V10666, does not help resolve the affinities of Crosbysaurus it does represent an approximately 250 kilometer extension of the geographic range of this taxon. This is the first record of this taxon in Utah and as such it represents the northernmost known record of Crosbysaurus. This indicates that Crosbysaurus was not limited to the southern area of Chinle/Dockum deposition but instead was widespread across the paleoriver systems of the Late Triassic in western Pangea. The specimen we report on here was found in close association with a typical Late Triassic Chinle fauna, including phytosaurs, metoposaurs, and dinosauromorphs.

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