Relationships and phylogenetic revision of Filistatinella spiders (Araneae : Filistatidae)

2017 ◽  
Vol 31 (6) ◽  
pp. 665 ◽  
Author(s):  
Ivan L. F. Magalhaes ◽  
Martín J. Ramírez

Filistatids represent an antique lineage of araneomorph spiders which are most diverse in arid and semiarid regions of the globe. Phylogenetic relationships among its genera are still largely unexplored, and previous studies disagree on the position of the North American Filistatinella Gertsch & Ivie, 1936, which could either be the sister group of all other Prithinae, or deeply nested in the subfamily. We present a new phylogenetic hypothesis based on morphological data, which supports the position of Filistatinella at the base of Prithinae. We also argue that the central Asian Pholcoides Roewer, 1960, hitherto considered incertae sedis in the subfamily, represents the putative sister group of Filistatinella. The latter genus is revised, and we describe its fine morphology in detail using optical and scanning electron microscopy. We redescribe the three previously known species, F. crassipalpis (Gertsch, 1935), F. domestica Desales-Lara, 2012 and F. palaciosi Jiménez & Palacios-Cardiel, 2012. Seven new species are named: F. kahloae, sp. nov. and F. chilindrina, sp. nov. from Mexico; F. pistrix, sp. nov., F. tohono, sp. nov., F. howdyall, sp. nov. and F. hermosa, sp. nov. from south-western USA; and F. spatulata, sp. nov. from the border between the two countries. The phylogenetic relationships among these 10 species are assessed, revealing the monophyly of the genus. http://zoobank.org/urn:lsid:zoobank.org:pub:71820858-545C-43EC-98E1-F9BF490AA3F1

Zootaxa ◽  
2012 ◽  
Vol 3315 (1) ◽  
pp. 1 ◽  
Author(s):  
FERNANDO LOBO ◽  
CRISTIAN ABDALA ◽  
SOLEDAD VALDECANTOS

With 36 species and at least nine potentially independent lineages (not formally described yet) occurring mostly in theAndes and adjacent Patagonia and Puna plateau areas, Phymaturus lizards represent one of the most endemic vertebrategroups of the arid southwestern region of South America. Phylogenetic relationships among species of Phymaturus areinferred using mainly a morphological data set of 206 characters. Also available sequences of mitochondrial DNA for sev-en terminals were added for a total evidence analysis. Most information is included in the discrete characters block; mostcharacters involve color pattern, osteology and squamation. Continuous characters were taken from body proportions,squamation and skeletons. Among morphological data, binary polymorphic characters were analyzed applying the scaledcoding criteria. Continuous characters were entered in the analysis using standardized ranges, a method that allows a sim-ple optimization to estimate distances/costs avoiding the arbitrary coding as discrete characters. For our parsimony anal-yses we chose the implied weights method, which underweights homoplastic characters. Several runs were madeanalyzing all the information combined and also separating morphological from molecular datasets. Binary polymor-phisms were also analyzed as missing data. All characters affected by sexual dimorphism were analyzed separating thesexes; female information was more congruent with the total evidence analysis. Characters involving continuous and poly-morphic information are relevant for searching and building phylogenetic hypotheses in Phymaturus. There exists signif-icant congruence between the molecular information analyzed in this study and previous published analyses. Within bothmain clades of Phymaturus, northern subgroups are those more recently originated during the genus diversification. Spe-cies belonging to the puna subclade of the palluma group are arranged in two natural groups, one distributed in the north(Catamarca and La Rioja provinces), and the other in the south, La Rioja and San Juan provinces. Within the patagonicusgroup, the majority of the species are arranged in a south-central Chubut clade, eastern-central Chubut clade, central Rio Negro clade and a Payunia clade.


2010 ◽  
Vol 79 (3) ◽  
pp. 93-106 ◽  
Author(s):  
Juliana Sterli

The origin and evolution of the crown-group of turtles (Cryptodira + Pleurodira) is one of the most interesting topics in turtle evolution, second perhaps only to the phylogenetic position of turtles among amniotes. The present contribution focuses on the former problem, exploring the phylogenetic relationships of extant and extinct turtles based on the most comprehensive phylogenetic dataset of morphological and molecular data analyzed to date. Parsimony analyses were conducted for different partitions of data (molecular and morphological) and for the combined dataset. In the present analysis, separate analyses of the molecular data always retrieve Pleurodira allied to Trionychia. Separate analysis of the morphological dataset, by contrast, depicts a more traditional arrangement of taxa, with Pleurodira as the sister group of Cryptodira, being Chelonioidea the most basal cryptodiran clade. The simultaneous analysis of all available data retrieves all major extant clades as monophyletic, except for Cryptodira given that Pleurodira is retrieved as the sister group of Trionychia. The paraphyly of Cryptodira is an unorthodox result, and is mainly caused by the combination of two factors. First, the molecular signal allies Pleurodira and Trionychia. Second, the morphological data with extinct taxa locates the position of the root of crown-group Testudines in the branch leading to Chelonioidea. This study highlights major but poorly explored topics of turtle evolution: the alternate position of Pleurodira and the root of crown turtles. The diversification of crown turtles is characterized by the presence of long external branches and short internal branches (with low support for the internal nodes separating the major clades of crown turtles), suggesting a rapid radiation of this clade. This rapid radiation is also supported by the fossil record, because soon after the appearance of the oldest crown-group turtles (Middle-Late Jurassic of Asia) the number and diversity of turtles increases remarkably. This evolutionary scenario of a rapid diversification of modern turtles into the major modern lineages is likely the reason for the difficulty in determining the interrelationships and the position of the root of crown-group turtles.


Zootaxa ◽  
2004 ◽  
Vol 629 (1) ◽  
pp. 1 ◽  
Author(s):  
MARIAM LEKVEISHVILI ◽  
HANS KLOMPEN

Phylogenetic relationships among the families in the infraorder Sejina and the position of Sejina relative to other infraorders of Mesostigmata are re-examined based on molecular and morphological data. Data sets included DNA sequence data for complete 18S, EF-1 , partial CO1genes, and 69 morphological characters. The two families of Heterozerconina consistently group within Sejina, and we propose to synonymize Heterozerconina with Sejina (Sejina s.l). Microgyniina is not the closest relative of Sejina. Rather, Sejina s.l. most often groups with Gamasina. Uropodellidae and Ichthyostomatogasteridae are sister groups and this lineage forms the sister group to Discozerconidae plus Heterozerconidae. Overall, we recognize 5 families within Sejina: Uropodellidae, Ichthyostomatogasteridae, Sejidae, Discozerconidae, and Heterozerconidae.


Hacquetia ◽  
2019 ◽  
Vol 18 (2) ◽  
pp. 337-347 ◽  
Author(s):  
Mehrshid Riahi ◽  
Farrokh Ghahremaninejad

Abstract Molecular data have been increasingly used to study the phylogenetic relationships among many taxa, including scrophs. Sometimes they have provided phylogenetic reconstructions that are in conflict with morphological data leading to a re-evaluation of long-standing evolutionary hypotheses. In this paper, we review reports of the recent knowledge of the phylogenetic relationships within Scrophularieae (2011–2017). The results of these analyses led to the following conclusions. (1) Species of Scrophularia have undergone one or more Miocene migration events occurred from eastern Asia to the North America with subsequent long dispersal and diversification in three main directions. (2) Allopolyploid and aneuploid hybrid speciation between Scrophularia species can occur, so hybridization and polyploidy have an important role for history of diversification. (3) The ancestral staminode type for the genus Scrophularia seems to be a large staminode. (4) Monophyly of the genus Verbascum with respect to the genus Scrophularia is strongly supported. (5) Oreosolen, is not monophyletic, because all accessions of Oreosolen were nested within Scrophularia. We discuss methods of data collection and analysis, and we describe the areas of conflict and agreement between molecular phylogenies.


2011 ◽  
Vol 25 (4) ◽  
pp. 334 ◽  
Author(s):  
Joel Ledford ◽  
Pierre Paquin ◽  
James Cokendolpher ◽  
Josh Campbell ◽  
Charles Griswold

A phylogenetic analysis of the spider genus Neoleptoneta Brignoli, 1972 is presented based on molecular sequence variation from three genes (mitochondrial cytochrome c oxidase subunit I, nuclear histone H3 and nuclear 28S rDNA) and including exemplars for all North American leptonetid genera except the ecribellate archoleptonetine Darkoneta. Analysis of concatenated data and independent genes using Bayesian, maximum likelihood and parsimony methods failed to recover Neoleptoneta as monophyletic. The genera Archoleptoneta, Appaleptoneta and Calileptoneta are monophyletic and a sister group relationship is supported between Appaleptoneta and Calileptoneta. Morphological data based on a survey of leptonetid genera using scanning electron and compound light microscopy are discussed and traced on the molecular phylogeny. Images for each North American leptonetine genus are provided, including comparison with Asian and European outgroups. Images of the incertae sedis species Leptoneta brunnea Gertsch, 1974 and Leptoneta sandra Gertsch, 1974 are provided and their generic placement is re-evaluated. Ancestral state reconstruction is used to assess patterns of cave evolution and shows that most species are descended from troglophilic ancestors and that troglobites have evolved at least nine times independently within the North American Leptonetidae. Neoleptoneta is relimited to include seven species restricted to central Mexico including N. bonita (Gertsch, 1974), N. capilla (Gertsch, 1971), N. delicata (Gertsch, 1971), N. limpida (Gertsch, 1974), N. rainesi (Gertsch, 1971) and N. reclusa (Gertsch, 1971) and to include Leptoneta brunnea, giving the new combination N. brunnea (Gertsch, 1974). The remaining species described in Neoleptoneta are placed in three new genera: (1) Chisoneta, gen. nov. from south-western Texas and Nuevo Leon, Mexico, including the four species C. chisosea (Gertsch, 1974), C. isolata (Gertsch, 1971), C. modica (Gertsch, 1974) and C. pecki (Gertsch, 1971), new combinations; (2) Ozarkia, gen. nov. from Arizona and New Mexico north-east to Arkansas, Alabama and Georgia, including the nine species O. alabama (Gertsch, 1974), O. apachea (Gertsch, 1974), O. archeri (Gertsch, 1974), O. arkansa (Gertsch, 1974), O. blanda (Gertsch, 1974), O. georgia (Gertsch, 1974), O. ivei (Gertsch, 1974), O. novaegalleciae (Brignoli, 1979) and O. serena (Gertsch, 1974), new combinations; and (3) Tayshaneta, gen. nov. from Texas south to Coahuila, Mexico, with the eleven species T. anopica (Gertsch, 1974), T. bullis (Cokendolpher, 2004), T. coeca (Chamberlin & Ivie, 1942), T. concinna (Gertsch, 1974), T. devia (Gertsch, 1974), T. furtiva (Gertsch, 1974), T. microps (Gertsch, 1974), T. myopica (Gertsch, 1974), T. paraconcinna (Cokendolpher & Reddell, 2001), T. uvaldea (Gertsch, 1974) and T. valverdae (Gertsch, 1974), new combinations. Leptoneta sandra Gertsch, 1974 cannot be placed in any North American, European or Asian genus and is thus transferred to the new genus Montanineta, gen. nov., giving the new combination Montanineta sandra (Gertsch, 1974).


2005 ◽  
Vol 19 (2) ◽  
pp. 105 ◽  
Author(s):  
Mark E. Siddall ◽  
Rebecca B. Budinoff ◽  
Elizabeth Borda

The phylogenetic relationships of Glossiphoniidae, a leech family characterised by its high degree of parental care, were investigated with the combined use of morphological data and three molecular datasets. There was strong support for monophyly of most accepted genera in the group, many of which are consistent with eyespot morphology. The genera Desserobdella Barta & Sawyer, 1990 and Oligobdella Moore, 1918 are suppressed as junior synonyms of Placobdella Blanchard, 1893 and thus recognising each of Placobdella picta (Verrill, 1872) Moore, 1906, Placobdella phalera (Graf, 1899) Moore, 1906, and Placobdella biannulata (Moore, 1900), comb. nov. The species Glossiphonia elegans (Verrill, 1872) Castle, 1900 and Helobdella modesta (Verrill, 1872), comb. nov. are resurrected for the North American counterparts to European non-sanguivorous species. Glossphonia baicalensis (Stschegolew, 1922), comb. nov. is removed from the genus Torix Blanchard 1898 and Alboglossiphonia quadrata (Moore, 1949) Sawyer, 1986 is removed from the genus Hemiclepsis Vejdovsky, 1884. The biogeographic implications of the phylogenetic hypothesis are evaluated in the context of what is already known for vertebrate hosts and Tertiary continental arrangements.


2002 ◽  
Vol 33 (4) ◽  
pp. 421-444 ◽  
Author(s):  
Kenneth P. Collins ◽  
Brian M. Wiegmann

AbstractThe phylogenetic relationships within the Eremoneura (Empidoidea + Cyclorrhapha) have been controversial. The monophyly of the Empidoidea, as well as the position and rank of higher-level empidoid clades remains unresolved despite numerous analyses using morphological data. In addition, the origin of the Cyclorrhapha and their relationship to the Empidoidea continues to be debated. We present the results of a molecular phylogenetic analysis using nucleotide sequences collected from 28S ribosomal DNA (rDNA) and elongation factor-1α (EF-1α) genes. All currently recognized empidoid families and subfamilies, many lower cyclorrhaphan families (including Opetiidae), and several asiloid outgroups are represented in this study. Unweighted and weighted parsimony, as well as maximum likelihood analyses were applied to individual data partitions and a combined data set. Our results support the monophyly of both Empidoidea and Cyclorrhapha (including Opetia), as well as their sister-group relationship. Within Empidoidea we find support for the following: 1) Chvála's (1983) proposal to divide Empidoidea into five families; 2) Atelestidae as the basal empidoid lineage; and 3) monophyly of Microphoridae + Dolichopodidae.


2002 ◽  
Vol 33 (4) ◽  
pp. 445-456 ◽  
Author(s):  
Brian M. Wiegmann ◽  
Kenneth P. Collins

AbstractCyclorrhaphan Diptera are an extremely successful clade of ecologically and phylogenenetically important flies. Despite their significance the relationships among lower cyclorrhaphans ('Aschiza') remain controversial in spite of several morphologically based phylogenetic analyses. We sequenced a 2.7-kb fragment of 28S rDNA for taxa representing all lower cyclorrhaphan families (except Ironomyiidae), four schizophoran families, and seven empidoid out-group taxa. Phylogenetic analysis of these data strongly supports a monophyletic Cyclorrhapha (including the enigmatic taxon Opetia nigra) that is divided into two clades - a well-supported Eumuscomorpha (Syrphidae + Pipunculidae + Schizophora), and a weakly-supported Platypezoidea (all non-Eumuscomorpha). Consequently, the former grouping known as Aschiza, which included syrphids and pipunculids, is not a valid monophyletic clade. Within Platypezoidea, most of our analyses place Lonchopteridae as sister group to Opetiidae, and strongly support the monophyly of Sciadoceridae + Phoridae. Among the Eumuscomorpha we do not recover the monophyly of Syrphoidea (Syrphidae + Pipunculidae). Instead, all analyses place Pipunculidae as the sister group to Schizophora. This novel finding has never been proposed based on morphological data and will require more data (both molecular and morphological) and taxa to confirm.


Zootaxa ◽  
2019 ◽  
Vol 4551 (2) ◽  
pp. 101 ◽  
Author(s):  
K. J. DAVID ◽  
S. RAMANI

The tribe Dacini comprising four genera, namely Bactrocera Macquart, Dacus Fabricius, Monacrostichus Bezzi and Zeugodacus Hendel, is a derived lineage in Tephritidae. It is one of the most economically important tribes in Tephritidae harbouring several species of quarantine concern across the world. We describe two new species of Bactrocera Macquart, B. (Parazeugodacus) conica David & Ramani, sp. n. & B. (B.) prabhui David, sp. n. from India. Postabdominal structures of males and/or females of 23 species of Bactrocera, 16 species of Zeugodacus and 8 species of Dacus from India are illustrated and described for the first time, which revealed similarities between Dacus and Zeugodacus with respect to epandrial shape and praeputium patterning. Bactrocera is unique in possessing oval shaped epandrium and an unpatterned praeputium. An analysis of phylogenetic relationships between three genera of the tribe Dacini from India based on morphological characters has been attempted for the first time. Cladistic analysis employing 51 characters of 62 species in Dacini, with seven species as outgroups revealed the monophyly of Dacini, Bactrocera and Dacus with supporting nonhomplasious synapomorphies. Ichneumonopsis Hardy, often included in the Gastrozonini, does not possess any synapomorphies of Dacini, eventhough it appeared at the base of the Dacini clade. Zeugodacus was retrieved as a monophyletic sister-group to Dacus based solely on a single homoplasious host plant character, with weak statistcal support. 


Paleobiology ◽  
1986 ◽  
Vol 12 (4) ◽  
pp. 383-399 ◽  
Author(s):  
Joel Cracraft

Numerical cladistic analysis of 73 cranial and postcranial characters has resulted in a highly corroborated hypothesis describing the phylogenetic pattern of early avian evolution. Using “non-avian theropod” dinosaurs as a comparative outgroup and root for the tree, the analysis confirmed Archaeopteryx to be the sister-group of all remaining avian taxa, or Ornithurae. This latter taxon is subdivided into two lineages, the Hesperornithiformes and the Carinatae. The carinates, in turn, were also resolved into two sister-groups, the Ichthyornithiformes and the modern birds, or Neornithes. This paper provides morphological data corroborating the divergence of the two basal clades of the Neornithes: the Palaeognathae (tinamous and ratites) and Neognathae (all other modern birds). The phylogenetic relationships of four important Cretaceous taxa were also investigated, but these fossil taxa were too fragmentary to determine their phylogenetic position unambiguously. Alexornis and Ambiortus are both carinates, but their relationships cannot be resolved in greater detail. The relationships of the Enantiornithes may lie within the Carinatae or these two taxa may be sister-groups. Gobipteryx is a neornithine and possibly the sister-group of the Palaeognathae.This analysis indicates that major patterns of morphological change took place at the time of origin of the ancestors of the Ornithurae and the Carinatae. Ornithurine innovations included major changes throughout the skeleton, whereas those of the carinates, while substantial, were primarily restricted to the pectoral girdle and forelimb. The phylogenetic results, in conjunction with the known ages of fossil taxa, indicate that the early lineages of birds very likely arose in the Jurassic. The early cladistic events within the neornithine lineage are also more ancient than generally recognized, and may well extend back to the early Cretaceous.


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