scholarly journals Anatomy and systematics of the diplodocoid Amphicoelias altus supports high sauropod dinosaur diversity in the Upper Jurassic Morrison Formation of the USA

2021 ◽  
Vol 8 (6) ◽  
pp. 210377
Author(s):  
Philip D. Mannion ◽  
Emanuel Tschopp ◽  
John A. Whitlock

Sauropod dinosaurs were an abundant and diverse component of the Upper Jurassic Morrison Formation of the USA, with 24 currently recognized species. However, some authors consider this high diversity to have been ecologically unviable and the validity of some species has been questioned, with suggestions that they represent growth series (ontogimorphs) of other species. Under this scenario, high sauropod diversity in the Late Jurassic of North America is greatly overestimated. One putative ontogimorph is the enigmatic diplodocoid Amphicoelias altus , which has been suggested to be synonymous with Diplodocus . Given that Amphicoelias was named first, it has priority and thus Diplodocus would become its junior synonym. Here, we provide a detailed re-description of A. altus in which we restrict it to the holotype individual and support its validity, based on three autapomorphies. Constraint analyses demonstrate that its phylogenetic position within Diplodocoidea is labile, but it seems unlikely that Amphicoelias is synonymous with Diplodocus . As such, our re-evaluation also leads us to retain Diplodocus as a distinct genus. There is no evidence to support the view that any of the currently recognized Morrison sauropod species are ontogimorphs. Available data indicate that sauropod anatomy did not dramatically alter once individuals approached maturity. Furthermore, subadult sauropod individuals are not prone to stemward slippage in phylogenetic analyses, casting doubt on the possibility that their taxonomic affinities are substantially misinterpreted. An anatomical feature can have both an ontogenetic and phylogenetic signature, but the former does not outweigh the latter when other characters overwhelmingly support the affinities of a taxon. Many Morrison Formation sauropods were spatio-temporally and/or ecologically separated from one another. Combined with the biases that cloud our reading of the fossil record, we contend that the number of sauropod dinosaur species in the Morrison Formation is currently likely to be underestimated, not overestimated.

Geosphere ◽  
2021 ◽  
Author(s):  
John I. Ejembi ◽  
Sally L. Potter-McIntyre ◽  
Glenn R. Sharman ◽  
Tyson M. Smith ◽  
Joel E. Saylor ◽  
...  

Middle to Upper Jurassic strata in the Paradox Basin and Central Colorado trough (CCT; south­western United States) record a pronounced change in sediment dispersal from dominantly aeolian deposition with an Appalachian source (Entrada Sandstone) to dominantly fluvial deposi­tion with a source in the Mogollon and/or Sevier orogenic highlands (Salt Wash Member of the Morrison Formation). An enigmatic abundance of Cambrian (ca. 527–519 Ma) grains at this prove­nance transition in the CCT at Escalante Canyon, Colorado, was recently suggested to reflect a local sediment source from the Ancestral Front Range, despite previous interpretations that local base­ment uplifts were largely buried by Middle to Late Jurassic time. This study aims to delineate spatial and tem­poral patterns in provenance of these Jurassic sandstones containing Cambrian grains within the Paradox Basin and CCT using sandstone petrog­raphy, detrital zircon U-Pb geochronology, and detrital zircon trace elemental and rare-earth ele­mental (REE) geochemistry. We report 7887 new U-Pb detrital zircon analyses from 31 sandstone samples collected within seven transects in west­ern Colorado and eastern Utah. Three clusters of zircon ages are consistently present (1.53–1.3 Ga, 1.3–0.9 Ga, and 500–300 Ma) that are interpreted to reflect sources associated with the Appalachian orogen in southeastern Laurentia (mid-continent, Grenville, Appalachian, and peri-Gondwanan terranes). Ca. 540–500 Ma zircon grains are anom­alously abundant locally in the uppermost Entrada Sandstone and Wanakah Formation but are either lacking or present in small fractions in the overlying Salt Wash and Tidwell Members of the Morrison Formation. A comparison of zircon REE geochem­istry between Cambrian detrital zircon and igneous zircon from potential sources shows that these 540–500 Ma detrital zircon are primarily magmatic. Although variability in both detrital and igneous REE concentrations precludes definitive identifica­tion of provenance, several considerations suggest that distal sources from the Cambrian granitic and rhyolitic provinces of the Southern Oklahoma aulacogen is also likely, in addition to a proximal source identified in the McClure Mountain syenite of the Wet Mountains, Colorado. The abundance of Cambrian grains in samples from the central CCT, particularly in the Entrada Sandstone and Wana­kah Formation, suggests northwesterly sediment transport within the CCT, with sediment sourced from Ancestral Rocky Mountains uplifts of the southern Wet Mountains and/or Amarillo-Wichita Mountains in southwestern Oklahoma. The lack of Cambrian grains within the Paradox Basin sug­gests that the Uncompahgre uplift (southwestern Colorado) acted as a barrier to sediment transport from the CCT.


PeerJ ◽  
2020 ◽  
Vol 8 ◽  
pp. e7803 ◽  
Author(s):  
Daniel J. Chure ◽  
Mark A. Loewen

Allosaurus is one of the best known theropod dinosaurs from the Jurassic and a crucial taxon in phylogenetic analyses. On the basis of an in-depth, firsthand study of the bulk of Allosaurus specimens housed in North American institutions, we describe here a new theropod dinosaur from the Upper Jurassic Morrison Formation of Western North America, Allosaurus jimmadseni sp. nov., based upon a remarkably complete articulated skeleton and skull and a second specimen with an articulated skull and associated skeleton. The present study also assigns several other specimens to this new species, Allosaurus jimmadseni, which is characterized by a number of autapomorphies present on the dermal skull roof and additional characters present in the postcrania. In particular, whereas the ventral margin of the jugal of Allosaurus fragilis has pronounced sigmoidal convexity, the ventral margin is virtually straight in Allosaurus jimmadseni. The paired nasals of Allosaurus jimmadseni possess bilateral, blade-like crests along the lateral margin, forming a pronounced nasolacrimal crest that is absent in Allosaurus fragilis.


PeerJ ◽  
2020 ◽  
Vol 8 ◽  
pp. e9510
Author(s):  
Julia B. McHugh ◽  
Stephanie K. Drumheller ◽  
Anja Riedel ◽  
Miriam Kane

A survey of 2,368 vertebrate fossils from the Upper Jurassic Mygatt-Moore Quarry (MMQ) (Morrison Formation, Brushy Basin Member) in western Colorado revealed 2,161 bone surface modifications on 884 specimens. This is the largest, site-wide bone surface modification survey of any Jurassic locality. Traces made by invertebrate actors were common in the assemblage, second in observed frequency after vertebrate bite marks. Invertebrate traces are found on 16.174% of the total surveyed material and comprise 20.148% of all identified traces. Six distinct invertebrate trace types were identified, including pits and furrows, rosettes, two types of bioglyph scrapes, bore holes and chambers. A minimum of four trace makers are indicated by the types, sizes and morphologies of the traces. Potential trace makers are inferred to be dermestid or clerid beetles, gastropods, an unknown necrophagous insect, and an unknown osteophagus insect. Of these, only gastropods are preserved at the site as body fossils. The remaining potential trace makers are part of the hidden paleodiversity from the North American Late Jurassic Period, revealed only through this ichnologic and taphonomic analysis. Site taphonomy suggests variable, but generally slow burial rates that range from months up to 6 years, while invertebrate traces on exposed elements indicate a minimum residence time of five months for carcasses with even few preserved invertebrate traces. These traces provide insight into the paleoecology, paleoclimate, and site formation of the MMQ, especially with regards to residence times of the skeletal remains on the paleolandscape. Comprehensive taphonomic studies, like this survey, are useful in exploring patterns of paleoecology and site formation, but they are also rare in Mesozoic assemblages. Additional work is required to determine if 16.174% is typical of bulk-collected fossils from Jurassic ecosystems in North America, or if the MMQ represents an unusual locality.


2005 ◽  
Vol 42 (3) ◽  
pp. 307-321 ◽  
Author(s):  
Ursula B Göhlich ◽  
Luis M Chiappe ◽  
James M Clark ◽  
Hans-Dieter Sues

Macelognathus vagans was described by O.C. Marsh in 1884, based on a mandibular symphysis from the Upper Jurassic Morrison Formation of Wyoming. Often considered a dinosaur but later tentatively referred to the Crocodylia, its phylogenetic identity has until now been enigmatic. New material of this species from the Morrison Formation of western Colorado demonstrates its affinities with basal crocodylomorphs commonly grouped together as the Sphenosuchia, which are characterized by a gracile postcranial skeleton with erect limb posture. Macelognathus shares features with Kayentasuchus from the Lower Jurassic Kayenta Formation of Arizona and Hallopus, which may be from the Morrison Formation of eastern Colorado. The new material constitutes the youngest definitive occurrence of a sphenosuchian, previously known from the Late Triassic to the Middle or Late? Jurassic.


2016 ◽  
Vol 53 (1) ◽  
pp. 34-40
Author(s):  
Robin L. Sissons ◽  
Michael W. Caldwell ◽  
Carol A. Evenchick ◽  
Donald B. Brinkman ◽  
Matthew J. Vavrek

Although the Jurassic was a period of high diversity in ichthyosaurs, only a small number of specimens have been recorded from Canada to date. We describe here a new occurrence of an ophthalmosaurid ichthyosaur from a shallow marine depositional environment within the Bowser Basin of northern British Columbia. Based on vertebral diameters and the size of the humerus, the ichthyosaur was relatively large compared to other contemporaneous forms, yet possessed teeth that were small for its body size. As well, the height to length ratio of the preserved vertebrae suggests it may have had a more elongate, less regionalized body shape. Although indeterminate at a generic level, the presence of Late Jurassic ichthyosaurs in nearshore waters of northwestern North America further demonstrates their cosmopolitan distribution.


PeerJ ◽  
2019 ◽  
Vol 7 ◽  
pp. e6348
Author(s):  
Philip D. Mannion

The Jurassic/Cretaceous (J/K) boundary, 145 million years ago, has long been recognised as an extinction event or faunal turnover for sauropod dinosaurs, with many ‘basal’ lineages disappearing. However, recently, a number of ‘extinct’ groups have been recognised in the Early Cretaceous, including diplodocids in Gondwana, and non-titanosauriform macronarians in Laurasia. Turiasauria, a clade of non-neosauropod eusauropods, was originally thought to have been restricted to the Late Jurassic of western Europe. However, its distribution has recently been extended to the Late Jurassic of Tanzania (Tendaguria tanzaniensis), as well as to the Early Cretaceous of the USA (Mierasaurus bobyoungi and Moabosaurus utahensis), demonstrating the survival of another ‘basal’ clade across the J/K boundary. Teeth from the Middle Jurassic–Early Cretaceous of western Europe and North Africa have also tentatively been attributed to turiasaurs, whilst recent phylogenetic analyses recovered Late Jurassic taxa from Argentina and China as further members of Turiasauria. Here, an anterior dorsal centrum and neural arch (both NHMUK 1871) from the Early Cretaceous Wealden Supergroup of the UK are described for the first time. NHMUK 1871 shares several synapomorphies with Turiasauria, especially the turiasaurs Moabosaurus and Tendaguria, including: (1) a strongly dorsoventrally compressed centrum; (2) the retention of prominent epipophyses; and (3) an extremely low, non-bifid neural spine. NHMUK 1871 therefore represents the first postcranial evidence for Turiasauria from European deposits of Early Cretaceous age. Although turiasaurs show clear heterodont dentition, only broad, characteristically ‘heart’-shaped teeth can currently be attributed to Turiasauria with confidence. As such, several putative turiasaur occurrences based on isolated teeth from Europe, as well as the Middle Jurassic and Early Cretaceous of Africa, cannot be confidently referred to Turiasauria. Unequivocal evidence for turiasaurs is therefore restricted to the late Middle Jurassic–Early Cretaceous of western Europe, the Late Jurassic of Tanzania, and the late Early Cretaceous of the USA, although remains from elsewhere might ultimately demonstrate that the group had a near-global distribution.


2020 ◽  
Vol 7 ◽  
pp. 29-67
Author(s):  
John Foster ◽  
Darrin Pagnac ◽  
ReBecca Hunt-Foster

The Little Houston Quarry in the Black Hills of Wyoming contains the most diverse vertebrate fauna in the Morrison Formation (Upper Jurassic) north of Como Bluff and the second-most diverse in the entire formation, after Reed’s Quarry 9. The deposit was an occasionally reactivated abandoned river channel, in interbedded green mudstone and laminated green-gray siltstone above a channel sandstone. The dinosaur material is densely distributed and is disarticulated to articulated, with several associated skeletons. The biota contains charophytes, horsetails, a possible seed fern, possible conifers, gastropods, two types of unionoid bivalves, diplostracans (“conchostracans”), a malacostracan, ray-finned fish, lungfish, a frog, salamanders, two types of turtles, rhynchocephalians, a lizard, choristoderes, two types of crocodyliforms, a pterosaur, Allosaurus and several types of small theropods including Tanycolagreus? and probable dromaeosaurids, numerous Camarasaurus and a diplodocine sauropod, a stegosaur, the neornithischian Nanosaurus, and the mammals Docodon, Amblotherium, and a multituberculate. Among these taxa, one of the unionoid bivalves, an atoposaurid crocodyliform, and the species of Amblotherium, which appear to be new and unique to the locality so far. The Docodon material may represent the first occurrence of D. apoxys outside of its type area in Colorado. Additionally, small, unusual theropod tooth types reported here may represent the first Late Jurassic occurrence of cf. Richardoestesia in North America and a possible abelisauroid, respectively.


Author(s):  
Christine Turner ◽  
Fred Peterson ◽  
D. Chure ◽  
T. Demko ◽  
G. Engelmann

The Morrison Project is a multidisciplinary effort to interpret the ancient ecosystem that was present in the Western Interior of the United States during deposition of the Upper Jurassic Morrison Formation. The project began in June of 1994 and the first two years of research (1994-95, 1995-96) were devoted primarily to identifying problems and gathering information that would form the basis for later interpretations. Efforts during the final year (1996-97) were directed toward resolving remaining problems or conflicting findings and synthesizing the various research endeavors into a reconstruction of the Late Jurassic ecosystem.


2020 ◽  
pp. 1-7
Author(s):  
Penélope Cruzado-Caballero ◽  
Ignacio Díaz-Martínez ◽  
Bruce Rothschild ◽  
Malcolm Bedell ◽  
Xabier Pereda-Suberbiola

2016 ◽  
Vol 154 (4) ◽  
pp. 837-858 ◽  
Author(s):  
ILARIA PAPARELLA ◽  
ERIN E. MAXWELL ◽  
ANGELO CIPRIANI ◽  
SCILLA RONCACÈ ◽  
MICHAEL W. CALDWELL

AbstractThe first ophthalmosaurid ichthyosaur from the Upper Jurassic deposits of the Central–Northern Apennines (Marche, Italy) is here described for the first time. The specimen is relatively complete and is referred to Gengasaurus nicosiai gen. et sp. nov. based on a unique combination of characters, including a peculiar condition of the preaxial accessory facet on the humerus. The faunal association of the ichthyosaur-bearing level indicates a late Kimmeridgian – earliest Tithonian age, and its finding contributes significantly to our knowledge of the diversity of Late Jurassic ichthyosaurs from the Western Tethys. Two shark teeth assigned to the order Hexanchiformes were also recovered in association with the ichthyosaur specimen, suggesting that scavenging of the carcass might have occurred. Gengasaurus can be referred to Ophthalmosauridae based on the reduced extracondylar area of the basioccipital, and the presence of a preaxial digit. It differs from Ophthalmosaurus spp. in several respects, including the shape of the posterior basisphenoid, the shape of the supraoccipital, the anteriorly deflected preaxial facet of the humerus, and a proximodistally shortened ulna. The new taxon actually shares diagnostic characters with both members of the two main lineages recovered in previous phylogenetic analyses, more nested within Ophthalmosauridae. The affinities of Gengasaurus to genera from both the northern and southern hemispheres also suggest that connectivity between pelagic habitats was high during the early Late Jurassic, allowing dispersal of some forms, followed by local, endemic divergence.


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