Distribution of epicormic branches and foliage on Douglas-fir as influenced by adjacent canopy gaps

2018 ◽  
Vol 48 (11) ◽  
pp. 1320-1330
Author(s):  
John W. Punches ◽  
Klaus J. Puettmann

The influence of adjacent canopy gaps on spatial distribution of epicormic branches and delayed foliage (originating from dormant buds) was investigated in 65-year-old coastal Douglas-fir (Pseudotsuga menziesii var. menziesii (Mirb.) Franco). Sample trees were selected across a broad range of local densities (adjacent canopy gap sizes) from a repeatedly thinned stand in which gaps had been created 12 years prior to our study. Lengths and stem locations of original and epicormic branches were measured within the south-facing crown quadrant, along with extents to which branches were occupied by sequential (produced in association with terminal bud elongation) and (or) delayed foliage. Epicormic branches, while prevalent throughout crowns, contributed only 10% of total branch length and 2% of total foliage mass. In contrast, delayed foliage occupied over 75% of total branch length, accounted for nearly 39% of total foliage mass, and often overlapped with sequential foliage. Canopy gap size did not influence original or epicormic branch length or location. On original branches, larger gaps may have modestly negatively influenced the relative extent of sequential foliage on branches and (or) slightly positively influenced delayed foliage mass. Delayed foliage appears to contribute substantially to Douglas-fir crown maintenance at this tree age, but canopy gap size had a minor influence, at least in the short term.

Web Ecology ◽  
2002 ◽  
Vol 3 (1) ◽  
pp. 1-5 ◽  
Author(s):  
D. Salvador-Van Eysenrode ◽  
F. Kockelbergh ◽  
J. Bogaert ◽  
I. Impens ◽  
P. Van Hecke

Abstract. Canopy gaps, i.e. openings in the forest cover caused by the fall of structural elements, are considered to be important for the maintenance of diversity and for the forest cycle. A gap can be considered as a young forest patch in the forest matrix, composed of interior surrounded by an edge, both enclosed by a perimeter. Much of the attention has been focused on the gap interior. However, at gap edges the spectrum of regeneration opportunities for plants may be larger than in the interior. Although definitions of gap are still discussed, any definition can describe it in an acceptable way, if justified, but defining edges is complicated and appropriate descriptors should be used. A method to determine gap interior and edge, using light as a descriptor, is presented with an example of gaps from a beech forest (Fagus sylvatica) in Belgium. Also, the relevance and implications of gap edges for plant diversity and calculation of forest turnover is discussed.


1988 ◽  
Vol 18 (2) ◽  
pp. 271-275 ◽  
Author(s):  
Barbara A. Smit

To identify Populustrichocarpa plants with contrasting levels of resistance to flooding, seedlings from five diverse riparian sites were evaluated for growth and survival under flooding conditions. All seedlings survived 6 or 8 weeks of flooding. Total branch length and leaf number were reduced in all flooded plants relative to nonflooded controls. There was also a marked reduction in individual leaf area and increased stomatal resistance of flooded plants compared with nonflooded controls. Growth of flooded and nonflooded plants was highly variable within populations and no significant trends were found among populations. Therefore differential responses to flooding can be selected for within any of the seed collection sites. Plants that were rated as particularly resistant or susceptible fo flooding were selected for further study.


1985 ◽  
Vol 117 (4) ◽  
pp. 395-399 ◽  
Author(s):  
Daniel J. Robison ◽  
Lawrence P. Abrahamson ◽  
Miroslaw M. Czapowskyj ◽  
Edwin H. White ◽  
Douglas C. Allen

AbstractOptimum size of a sample unit and within-branch distribution of overwintering spruce budworm were determined for black spruce in northern Maine. No significant differences in sample reliability were found between whole-branch and 45-cm branch-tip samples. Larval distribution on branches varied with total branch length and a model was developed to estimate the whole-branch population from a 45-cm branch tip. Use of a 45-cm branch-tip sample unit is recommended because it is biologically and statistically valid and reduces sampling costs.


2014 ◽  
Vol 46 (2) ◽  
pp. 496-515 ◽  
Author(s):  
Alexander Gnedin ◽  
Alexander Iksanov ◽  
Alexander Marynych ◽  
Martin Möhle

We show that the total number of collisions in the exchangeable coalescent process driven by the beta (1, b) measure converges in distribution to a 1-stable law, as the initial number of particles goes to ∞. The stable limit law is also shown for the total branch length of the coalescent tree. These results were known previously for the instance b = 1, which corresponds to the Bolthausen-Sznitman coalescent. The approach we take is based on estimating the quality of a renewal approximation to the coalescent in terms of a suitable Wasserstein distance. Application of the method to beta (a, b)-coalescents with 0 < a < 1 leads to a simplified derivation of the known (2 - a)-stable limit. We furthermore derive asymptotic expansions for the moments of the number of collisions and of the total branch length for the beta (1, b)-coalescent by exploiting the method of sequential approximations.


1954 ◽  
Vol 32 (5) ◽  
pp. 630-653 ◽  
Author(s):  
G. P. Thomas ◽  
R. W. Thomas

An investigation of Douglas fir, Pseudotsuga taxifolia (Poir.) Britton, in coastal British Columbia has shown that decay losses in old-growth stands are low, amounting to 2.6 and 7.3% of the gross volume of living and combined living and dead trees respectively. Twenty-five decay-producing fungi were isolated from living trees and 29 from dead trees. The most important of these were Fomes pini (Thore) Lloyd, Polyporus schweinitzii Fries, and Fomes pinicola (Sw.) Cke. Certain irregularities in the occurrence of decay-producing fungi and the amounts of decay associated with them were traced to the influences of site, tree age or size, latitude, and stand history. Root infections were the most frequent but branch-stub infections caused the greatest average amount of decay. An examination of tree abnormalities showed that a select group of them, sporophores and swollen knots of Fomes pini in particular, are useful indicators of hidden decay. A separate analysis of dead trees has shown that they occasion ally form a high proportion of Douglas fir stands and that much of the wood in such trees is sound.


2016 ◽  
Vol 363 ◽  
pp. 31-38 ◽  
Author(s):  
Dominik Seidel ◽  
Kenneth J. Ruzicka ◽  
Klaus Puettmann
Keyword(s):  

1991 ◽  
Vol 21 (6) ◽  
pp. 821-828 ◽  
Author(s):  
Robert G. Wagner ◽  
Steven R. Radosevich

Neighborhood models describing the effect of interspecific competition on the height and stem diameter of 4- to 9-year-old saplings of Douglas-fir (Pseudotsugamenziesii (Mirb.) Franco) were developed from site-preparation experiments in the Oregon Coast Range. The influence of abundance measures, height, distance, arid spatial arrangement of nonconiferous woody plants surrounding individual saplings was examined. Optimum neighborhood heights and radii were defined. The best interspecific competition index for predicting Douglas-fir height and stem diameter was total percent cover for all woody species within a 2.1-m radius. Visual estimates of neighbor cover were superior to objective measures of crown area. The cover of woody species equalling or exceeding the height of the tree provided the best prediction for tree height. Woody species cover equalling or exceeding one-half the height of the tree provided the best index for predicting stem diameter. Accounting for the spatial arrangement of neighboring woody plants did not improve the competition index. Interaction between the competition index and tree age indicated that the negative effect of interspecific competition on Douglas-fir size increased with time. The age-adjusted competition index accounted for 11% of the variation in height and 19% of the variation in stem diameter. Douglas-fir stem diameter was more sensitive to neighboring woody plants than was height.


1985 ◽  
Vol 15 (2) ◽  
pp. 354-364 ◽  
Author(s):  
J. N. Owens ◽  
J. E. Webber ◽  
S. D. Ross ◽  
R. P. Pharis

The relative importance of cell division and cell elongation to shoot elongation and the anatomical changes in vegetative terminal apices were assessed for 9- and 10-year-old seedlings of Douglas-fir (Pseudotsugamenziesii (Mirb.) Franco) in response to two effective cone-induction treatments, gibberellin A4/7 (GA4/7) and root-pruning (RP). Root-pruning was done in mid-April at the start of vegetative bud swelling and GA treatments were begun at vegetative bud flushing in mid-May and continued until early July. Shoot elongation before flushing resulted primarily from cell divisions and was not affected by the RP treatment. Shoot elongation after flushing resulted primarily from cell expansion which was reduced by RP treatments. Root-pruning significantly slowed mitotic activity, apical growth, and development of vegetative terminal buds from mid-June through mid-July. Apical growth then resumed during leaf initiation and the final number of leaf primordia initiated was not affected. This resulted in a delay of 2 to 4 weeks in the transition from bud-scale to leaf initiation. Retarded terminal vegetative apices anatomically resembled latent axillary apices but were never completely inhibited. GA + RP had the same effect as RP. GA4/7 alone had no effect on shoot or apical development. These results show that RP and GA + RP significantly retard shoot elongation and terminal bud development but still allow normal development of vegetative terminal buds. Retardation of bud development by a few weeks shifts the critical morphogenetic phase of transition from bud scale to leaf initiation to a later time when endogenous and environmental conditions may differ from the normal.


Fractals ◽  
1998 ◽  
Vol 06 (01) ◽  
pp. 81-86 ◽  
Author(s):  
Makoto Katori ◽  
Shinya Kizaki ◽  
Youichi Terui ◽  
Takuya Kubo

Importance of the influence of neighboring canopy gaps upon new gap creation has been clarified by the ecological study of a neotropical forest on Barro Colorado Island (BCI), Panama. A stochastic lattice model for the forest dynamics with interacting canopy gap expansion was introduced by Kubo et al. We give a theorem showing a condition that this model can be regarded as a stochastic Ising model, and that its stationary state is exactly given by a Gibbs state. Using this theorem, we obtain a Gibbs state which remarkably well approximates the real gap-size distribution in BCI.


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