Impacts of drought on forest growth and regeneration following fire in southwestern Yukon, Canada

2005 ◽  
Vol 35 (9) ◽  
pp. 2141-2150 ◽  
Author(s):  
EH (Ted) Hogg ◽  
Ross W Wein

The valleys of southwestern Yukon have a continental climate with average annual precipitation of <300 mm. In 1958, fires burned large areas of mature mixedwood forests dominated by white spruce (Picea glauca (Moench) Voss) in the valleys near Whitehorse. Since then, the burned areas have shown poor regeneration of spruce, but have been colonized by scattered clones of trembling aspen (Populus tremuloides Michx.) interspersed by grassland. The objective of the study was to examine the influence of climatic variation on forest growth and regeneration in the 1958 burn and the adjacent unburned forests. Tree-ring analysis was conducted on 50 aspen and 54 white spruce in 12 mature stands where these species were codominant, and on 147 regenerating aspen in the 1958 Takhini burn. The mature stands were uneven-aged and the patterns of growth variation for the aspen and spruce between 1944 and 2000 were similar. Growth of both species was most strongly related to variation in precipitation. The regenerating aspen had a wide age-class distribution (1959–2000) and their growth was also positively related to precipitation. The results indicate that these forests have been slow to regenerate after fire, and are vulnerable if the climate becomes drier under future global change.

1999 ◽  
Vol 75 (3) ◽  
pp. 505-513 ◽  
Author(s):  
Rongzhou Man ◽  
Victor J. Lieffers

In boreal mixedwood forests, aspen (Populus tremuloides) and white spruce (Picea glauca) commonly grow in mixture. These species may avoid competition through differential shade tolerance, physical separation of canopies, phenological differences, successional separation, and differences in soil resource utilization. Aspen may also be able to positively affect the growth of white spruce by improving litter decomposition and nutrient cycling rates, controlling grass and shrub competition, ameliorating environmental extremes, and reducing pest attack. These positive relationships likely make mixed-species stands more productive than pure stands of the same species. The evidence regarding the productivity of pure versus mixed aspen/white spruce stands in natural unmanaged forests is examined in this paper. Key words: Tree mixture; productivity; boreal mixedwoods; aspen; white spruce


2006 ◽  
Vol 36 (6) ◽  
pp. 1597-1609 ◽  
Author(s):  
Vernon S Peters ◽  
S Ellen Macdonald ◽  
Mark RT Dale

The timing of white spruce regeneration in aspen (Populus tremuloides Michx.) – white spruce (Picea glauca (Moench) Voss) boreal mixedwood stands is an important factor in stand development. We examined boreal mixedwood stands representing a 59-year period of time since fire and determined (1) whether and when a delayed regeneration period of white spruce occurred, (2) whether the relative abundance of initial (<20 years) versus delayed (≥20 years postfire) regeneration is related to seed availability at the time of the fire, and (3) what are the important regeneration substrates for initial versus delayed regeneration. Initial regeneration occurred primarily on mineral soil or humus, while delayed regeneration established primarily on logs and peaked 38–44 years after fire. Of the 20 stands investigated, seven were dominated by initial regeneration, six were dominated by delayed regeneration, and seven were even mixtures of both. The dominance of a site by initial or delayed regeneration could not be simply explained by burn timing relative to mast years or distance to seed source; our results suggested that fire severity and the competitive influence of initial regeneration on delayed regeneration were important at fine scales. Based on our results we describe several possible postfire successional pathways for boreal mixedwood forests.


2009 ◽  
Vol 85 (4) ◽  
pp. 631-638 ◽  
Author(s):  
Alison D Lennie ◽  
Simon M Landhäusser ◽  
Victor J Lieffers ◽  
Derek Sidders

Trembling aspen regeneration was studied in 2 types of partial harvest systems designed to harvest mature aspen but protect immature spruce and encourage natural aspen regeneration. Two partial harvest systems, where the residual aspen was either left in strips or was dispersed uniformly, were compared to traditional clearcuts. After the first and second year since harvest, aspen sucker density and growth was similar between the 2 partial harvests, but was much lower than in the clearcuts. However, in the partial cuts the regeneration density was very much dependent on the location relative to residual trees. The density of regeneration was inversely related to the basal area of residual aspen; however, sucker height was inversely related to the basal area of the residual spruce. Although there were adequate numbers of suckers after partial harvest, their viability and contribution to the long-term productivity of these mixedwood stands is not clear. Key words: silvicultural systems, forest management, residual canopy, white spruce, Populus tremuloides, Picea glauca, traffic


1996 ◽  
Vol 72 (3) ◽  
pp. 286-292 ◽  
Author(s):  
V. J. Lieffers ◽  
J. D. Stewart ◽  
R. B. Macmillan ◽  
D. Macpherson ◽  
K. Branter

Boreal mixedwood forests of aspen (Populus tremuloides) and white spruce (Picea glauca) are found on mesic sites in the western boreal forest. In the natural development of mixedwood stands, aspen is usually the first species to dominate the site. However, depending upon spruce seed sources and seedbeds, spruce can establish immediately after disturbance or in the next several decades. In most cases, spruce grow in the understory of deciduous species during its early development. If there are no spruce seed sources, aspen may be the sole tree species for a long period. In most circumstances, however, the longer-lived and taller white spruce eventually becomes the dominant species. If stands remain undisturbed for long periods, they will likely become uneven-aged mixtures of spruce and balsam fir (Abies balsamea). We propose silvicultural systems that will develop stands of a range of compositions, structures and value. As a starting point, we identify eight different mixed-wood compositions that might be identified in stand inventories, and propose various silvicultural treatments, including underplanting of white spruce, understory protection, shelterwood, and uneven-aged management. Fundamental changes in land tenure and silvicultural regulations, and improvements in estimation of growth and yield will be required before this range of management of mixed-woods can be implemented. Key words: aspen, white spruce, shelterwood, Populus tremuloides, Picea glauca, succession, ecosystem management


2007 ◽  
Vol 22 (3) ◽  
pp. 163-170 ◽  
Author(s):  
Ryan J. Klos ◽  
G. Geoff Wang ◽  
Qing-Lai Dang ◽  
Ed W. East

Abstract Kozak's variable exponent taper equation was fitted for balsam poplar (Populus balsamifera L.), trembling aspen (Populus tremuloides Michx.), white spruce (Picea glauca [Moench] Voss), black spruce (Picea mariana [Mill.] B.S.P.), and jack pine (Pinus banksiana Lamb.) in Manitoba. Stem taper variability between two ecozones (i.e., Boreal Shield and Boreal Plains) were tested using the F-test. Regional differences were observed for trembling aspen, white spruce, and jack pine, and for those species, separate ecozone-specific taper equations were developed. However, the gross total volume estimates using the ecozone-specific equations were different from those of the provincial equations by only 2 percent. Although the regional difference in stem form was marginal within a province, a difference of approximately 7 percent of gross total volume estimation was found when our provincial taper equations were compared with those developed in Alberta and Saskatchewan. These results suggest that stem form variation increases with spatial scale and that a single taper equation for each species may be sufficient for each province.


2014 ◽  
Vol 44 (2) ◽  
pp. 162-171 ◽  
Author(s):  
Derek F. Sattler ◽  
Philip G. Comeau ◽  
Alexis Achim

Radial patterns of modulus of elasticity (MOE) were examined for white spruce (Picea glauca (Moench) Voss) and trembling aspen (Populus tremuoides Michx.) from 19 mature, uneven-aged stands in the boreal mixedwood region of northern Alberta, Canada. The main objectives were to (1) evaluate the relationship between pith-to-bark changes in MOE and cambial age or distance from pith; (2) develop species-specific models to predict pith-to-bark changes in MOE; and (3) to test the influences of radial growth, relative vertical height, and tree slenderness (tree height/DBH) on MOE. For both species, cambial age was selected as the best explanatory variable with which to build pith-to-bark models of MOE. For white spruce and trembling aspen, the final nonlinear mixed-effect models indicated that an augmented rate of increase in MOE occurred with increasing vertical position within the tree. For white spruce trees, radial growth and slenderness were found to positively influence maximum estimated MOE. For trembling aspen, there was no apparent effect of vertical position or radial growth on maximum MOE. The results shed light on potential drivers of radial patterns of MOE and will be useful in guiding silvicultural prescriptions.


2006 ◽  
Vol 36 (7) ◽  
pp. 1818-1833 ◽  
Author(s):  
Daniel A MacIsaac ◽  
Philip G Comeau ◽  
S Ellen Macdonald

This study assessed the dynamics of gap development in postharvest regeneration in five stands in northwestern Alberta dominated by trembling aspen (Populus tremuloides Michx.). The pattern of gap development over time was determined from analysis of air photographs taken preharvest and 1, 4, 10, and 12 years postharvest. The area of each stand covered by gaps increased after harvest because of the addition of harvest-related gaps over and above those that had been present prior to harvest. The blocks we studied had a combined gap area of up to 29% of stand area 12 years postharvest. We measured regeneration characteristics, microsite, soil, light, and browse conditions in 30 aspen regeneration gaps (gaps in regeneration that were not gaps preharvest and were not due to obvious harvest-related disturbance) 14 years following harvest. Although deciduous trees within postharvest regeneration gaps were the same age as those outside (i.e., in a fully stocked matrix of newly established even-aged aspen stems), they were often suppressed, with significantly lower density and growth. Within the 14-year-old postharvest regenerating aspen stands, aspen height varied from 1 to 11 m; this substantial variability appeared to be largely due to the influence of browsing. There was little evidence of ongoing regeneration within postharvest regeneration gaps, indicating that these gaps will probably persist over time. This may impact future deciduous stocking and volume. It is unknown what may have initiated the formation of these gaps, although results suggest that they are not due to edaphic conditions or disease in the preharvest stands. There is evidence that bluejoint (Calamagrostis canadensis (Michx.) Beauv.) cover and browsing are important factors in the maintenance of postharvest regeneration gaps. The spatial heterogeneity resulting from gaps could be advantageous, however, either as part of ecosystem-based management emulating natural disturbance or as a template for mixedwood management, where white spruce (Picea glauca (Moench) Voss) are established in gaps.


2002 ◽  
Vol 32 (6) ◽  
pp. 1071-1079 ◽  
Author(s):  
Clive Welham ◽  
Brad Seely ◽  
Hamish Kimmins

The ecosystem model FORECAST was used to simulate the yield potential in Saskatchewan mixedwoods of the two-pass harvesting system. The simulated two-pass stand consisted of an overstory population of pure trembling aspen (Populus tremuloides Michx.) with a white spruce (Picea glauca (Moench) Voss) understory. The aspen was removed at year 60, and yields of the understory spruce and resprouting aspen were simulated for 80 years thereafter. The two-pass simulations were compared with two simulated conventional harvesting systems. The first system consisted of a single final harvest at year 140. In the second system, a clearcut was conducted at year 60. White spruce was then planted in the subsequent year at 400, 600, or 800 stems/ha, and aspen also permitted to resprout. Growth was then simulated for a further 80 years. FORECAST projections indicated that the two-pass system might be effective for releasing the white spruce understory, achieving at least a twofold gain in spruce volume relative to conventional methods. Furthermore, total volumes exceeded those derived from the unmanaged stand, while second rotation yields of aspen declined with spruce understory density. These simulations suggest the two-pass harvesting system has strong potential as a tool for mixedwood management.


2002 ◽  
Vol 80 (6) ◽  
pp. 684-689 ◽  
Author(s):  
Simon M Landhäusser ◽  
Tawfik M Muhsin ◽  
Janusz J Zwiazek

Low soil temperatures, common during the growing season in northern forests, have the potential to impede plant growth. In this study, water uptake, water relations, and growth characteristics were examined in aspen (Populus tremuloides) and white spruce (Picea glauca) seedlings that were inoculated with ectomycorrhizal fungi and grown at 20°C daytime air temperatures and low soil temperatures of 4°C and 8°C. Mycorrhizal associations had little effect on root and shoot biomass at both soil temperatures. Root hydraulic conductance (Kr) was higher in both mycorrhizal plant species compared to nonmycorrhizal plants, but there was no soil temperature effect on Kr in either species. Mycorrhizae also increased shoot water potential (Ψw) in Populus tremuloides but had no effect on Ψw in Picea glauca. The increases in Kr and Ψw were not reflected by changes in stomatal conductance (gs) and transpiration rates (E), suggesting that the reduction of water flow in seedlings exposed to low soil temperature was not likely the factor limiting gs in both plant species.Key words: boreal forest, root hydraulic conductance, root growth, stomatal conductance, water uptake.


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