Effects of Fertilization On The Growth Dominance of Inland Northwest Forests of The United States
Abstract Large trees have disproportionally large competitive advantage in access to light, which has been proposed to increase growth dominance of large trees (e.g., accounting for a larger proportion of growth than volume of a stand). Tree growth may also be limited by the availability of other resources besides light. Nutrient deficiency, especially of nitrogen, is considered common among temperate forests, including mixed-conifer forests of the Inland Northwest of the United States. Data from a long-term forest nutrition study across four Inland Northwest states were analyzed to evaluate nitrogen × potassium fertilization's effects on growth dominance over an eight-year period following treatment in a region of complex forest vegetation types and site conditions. Our results show that growth dominance varied substantially across similar plots in each fertilization treatment, and its mean values generally were at minuscule magnitudes, negative, and not significantly different from zero. We propose that this lack of a clear pattern in growth dominance was the result of the mixed-species composition where shade-tolerant species remained in lower crown positions, yet their relative growth kept pace with large trees. Limited moisture availability at dry sites may have hampered the development of growth dominance. Growth dominance also was lowered by mortality observed among relatively large trees. The largely negative growth dominance across fertilization treatments indicates that small trees maintained higher relative growth rates than large trees, even if absolute growth and size was concentrated in large trees. In the case fertilization does improve stand growth, a significant part of this improved growth will be lost in density-dependent mortality over time if not captured through biomass removals.