<p>The quantification of the net carbon flux from land use and land cover changes (f<sub>LULCC</sub>) is essential to understand the global carbon cycle, and consequently, to support climate change mitigation. However, large-scale f<sub>LULCC</sub> is not directly measurable, and can only be inferred by models, such as semi-empirical bookkeeping models, and process-based dynamic global vegetation models (DGVMs). By definition, f<sub>LULCC</sub> estimates between these two model types are not directly comparable. For example, transient DGVM-based f<sub>LULCC</sub> of the annual global carbon budget includes the so-called Loss of Additional Sink Capacity (LASC). The latter accounts for environmental impacts on the land carbon storage capacities of managed land compared to potential vegetation which is not included in bookkeeping models. Additionally, estimates of transient DGVM-based f<sub>LULCC</sub> differ from bookkeeping model estimates, since they depend on arbitrarily chosen simulation time periods and the timing of land use and land cover changes within the historic period (which includes different accumulation periods for legacy effects). However, DGVMs enable a f<sub>LULCC</sub> approximation independent of the timing of land use and land cover changes and their legacy effects by simulations run under constant pre-industrial or present-day environmental forcings.</p><p>In this study, we analyze these different DGVM-derived f<sub>LULCC</sub> definitions, under transiently changing environmental conditions and fixed pre-industrial and fixed present-day conditions, within 18 regions for twelve DGVMs and quantify their differences as well as climate- and CO<sub>2</sub>-induced components. The multi model mean under transient conditions reveals a global f<sub>LULCC</sub> of 2.0&#177;0.6 PgC yr<sup>-1</sup> for 2009-2018, with ~40% stemming from the LASC (0.8&#177;0.3 PgC yr<sup>-1</sup>). Within the industrial period (1850 onward), cumulative f<sub>LULCC</sub> reached 189&#177;56 PgC with 40&#177;15 PgC from the LASC.</p><p>Regional hotspots of high LASC values exist in the USA, China, Brazil, Equatorial Africa and Southeast Asia, which we mainly relate to deforestation for cropland. Distinct negative LASC estimates were observed in Europe (early reforestation) and from 2000 onward in the Ukraine (recultivation of post-Soviet abandoned agricultural land). Negative LASC estimates indicate that fLULCC estimates in these regions are lower in transient DGVM simulations compared to bookkeeping-approaches. By unraveling the spatio-temporal variability of the different DGVM-derived f<sub>LULCC</sub> estimates, our study calls for a harmonized attribution of model-derived f<sub>LULCC</sub>. We propose an approach that bridges bookkeeping and DGVM approaches for f<sub>LULCC</sub> estimation by adopting a mean DGVM-ensemble LASC for a defined reference period.</p>