Some Cacti Produce “Reactioncortex” Rather than Reaction Wood in Curved Sections of Weight-Stressed Shoots

Haseltonia ◽  
2020 ◽  
Vol 2019 (26) ◽  
pp. 86
Author(s):  
James D Mauseth
Keyword(s):  
Reaction Wood

The nature of reaction wood. IV. Variations in cell wall organization of tension wood fibres

1955 ◽  
Vol 3 (2) ◽  
pp. 177 ◽  
Author(s):  
AB Wardrop ◽  
HE Dadswell
Keyword(s):  
Cell Wall ◽  
Tension Wood ◽  
Secondary Wall ◽  
Degree Of Crystallinity ◽  
Tropical Species ◽  
Normal Wood ◽  
Reaction Wood ◽  
Gelatinous Layer ◽  
Wood Fibres ◽  
Cell Wall Organization

The cell wall organization, the cell wall texture, and the degree of lignification of tension wood fibres have been investigated in a wide variety of temperate and tropical species. Following earlier work describing the cell wall structure of tension wood fibres, two additional types of cell wall organization have been observed. In one of these, the inner thick "gelatinous" layer which is typical of tension wood fibres exists in addition to the normal three-layered structure of the secondary wall; in the other only the outer layer of the secondary wall and the thick gelatinous layer are present. In all the tension wood examined the micellar orientation in the inner gelatinous layer has been shown to be nearly axial and the cellulose of this layer found to be in a highly crystalline state. A general argument is presented as to the meaning of differences in the degree, of crystallinity of cellulose. The high degree of crystallinity of cellulose in tension wood as compared with normal wood is attributed to a greater degree of lateral order in the crystalline regions of tension wood, whereas the paracrystalline phase is similar in both cases. The degree of lignification in tension wood fibres has been shown to be extremely variable. However, where the degree of tension wood development is marked as revealed by the thickness of the gelatinous layer the lack of lignification is also most marked. Severity of tension wood formation and lack of lignification have also been correlated with the incidence of irreversible collapse in tension wood. Such collapse can occur even when no whole fibres are present, e.g. in thin cross sections. Microscopic examination of collapsed samples of tension wood has led to the conclusion that the appearance of collapse in specimens containing tendon wood can often be attributed in part to excessive shrinkage associated with the development of fissures between cells, although true collapse does also occur. Possible explanations of the irreversible shrinkage and collapse of tension wood fibres are advanced.

Some Tree Growth – Wood Property Relationships of Eucalypts

1975 ◽  
Vol 5 (3) ◽  
pp. 424-432 ◽  
Author(s):  
J. E. Nicholson ◽  
W. E. Hillis ◽  
N. Ditchburne
Keyword(s):  
Wood Property ◽  
Basic Density ◽  
Wood Properties ◽  
Reaction Wood ◽  
Growth Strain ◽  
Fiber Structure ◽  
Tree Crown ◽  
Stem Form ◽  
The Relationship ◽  
Longitudinal Growth Strain

The relationship between level of longitudinal growth strain and stress, modulus of elasticity, basic density, volumetric shrinkage, fiber classification, and stem form was investigated with 10 Eucalyptusregnans regrowth trees. Close relationships were observed. It is suggested that variations in these properties within trees are closely controlled, possibly to enable optimum positioning of the tree crown in relation to its immediate environment. The concept of fiber structure varying as a response to environment is supported by the observed variation in wood properties within and between trees of this species. The often-reported association between eccentric radial growth and reaction wood was not substantiated in this study.The data indicate that if economically justifiable, it would be possible to segregate trees that are likely to contain material that is hard to season.

scholarly journals Reaction wood anatomy and lignin distribution in Gnetum gnemon branches

2018 ◽  
Vol 64 (6) ◽  
pp. 872-879 ◽  
Author(s):  
Haruna Aiso-Sanada ◽  
Futoshi Ishiguri ◽  
Denny Irawati ◽  
Imam Wahyudi ◽  
Shinso Yokota
Keyword(s):  
Wood Anatomy ◽  
Reaction Wood ◽  
Lignin Distribution ◽  
Gnetum Gnemon

scholarly journals QUALITY OF REACTION WOOD IN Eucalyptus TREES TILTED BY WIND FOR PULP PRODUCTION

CERNE ◽  
2017 ◽  
Vol 23 (3) ◽  
pp. 291-297
Author(s):  
Walter Torezani Neto Boschetti ◽  
Juarez Benigno Paes ◽  
Graziela Baptista Vidaurre ◽  
Marina Donária Chaves Arantes ◽  
João Gabriel Missia da Silva
Keyword(s):  
Chemical Composition ◽  
Chemical Constituents ◽  
Wood Quality ◽  
Basic Density ◽  
Kraft Pulping ◽  
Normal Wood ◽  
Reaction Wood ◽  
Opposite Wood ◽  
Holocellulose Content

ABSTRACT This study aims to evaluate the quality of normal, tension and opposite wood of eucalyptus trees lengthwise, in straight and inclined stems, affected by wind action. It also aims to explain the pulping parameters resultant from the quality of the wood. The trees were grouped into four tilt ranges, ranging from 0 to 50º, and the basic density, chemical composition of the wood, and performance in kraft pulping were assessed. Normal and tension wood had similar basic densities; while for opposite wood, the density was lower, being responsible for a decrease in reaction wood density. The chemical composition of the wood was influenced by the presence of reaction wood in the stem. Tension and opposite wood showed lower levels of extractives and lignin and higher holocellulose content when compared to normal wood, with favorable wood quality for pulping. The increase in holocellulose content and the reduction of lignin and extractives content contributed positively to a more delignified pulp and reduction of the Kappa number. However, after cooking the reaction wood under the same conditions as those of normal wood, reaction wood pulping tends to have a lower screen yields. Due to differences in basic density and chemical constituents between opposite and normal wood, it is recommended not to designate the opposite wood as normal wood.

scholarly journals Resin canals in spruce (Picea abis /L./ Karst.) with the occurrence of reaction wood

2005 ◽  
Vol 53 (1) ◽  
pp. 85-92 ◽  
Author(s):  
Vladimír Gryc ◽  
Petr Horáček
Keyword(s):  
Compression Wood ◽  
3D Models ◽  
Resin Canal ◽  
The Other ◽  
Stem Length ◽  
Reaction Wood ◽  
Spruce Wood ◽  
Other Hand ◽  
Resin Canals

The paper was aimed at the determination of variability of horizontal resin canal dimension in spruce wood in relation to the position in a spruce stem. Significant changes of dimensions in horizontal resin canal along the stem length and radius were found. On the basis obtained of results 3D models (for CW, OW, SWL and SWP zones) describing changes in resin canal dimensions in spruce in relation to the position in a stem were created. In the models, the resin canal dimension decreases with the height of a stem and on the other hand, with an increasing distance from the stem pith the dimension of resin canal increases. The importance of the paper consists in the enlargement of findings about the structure of spruce with compression wood.

scholarly journals Straightening the crooked: intraspecific divergence of stem posture control and associated trade-offs in a model conifer

2021 ◽  
Author(s):  
R Sierra-de-Grado ◽  
V Pando ◽  
J Voltas ◽  
R Zas ◽  
J Majada ◽  
...  
Keyword(s):  
Mechanical Stability ◽  
Common Garden ◽  
Shoot Elongation ◽  
Reaction Wood ◽  
Forest Trees ◽  
Manipulative Experiment ◽  
Intraspecific Divergence ◽  
Trade Offs ◽  
Adaptive Phenotypic Plasticity ◽  
Light Capture

Abstract Although the straightening capacity of the stem is key for light capture and mechanical stability in forest trees, little is known about its adaptive implications. Assuming that stem straightening is costly, trade-offs are expected with competing processes such as growth, maintenance and defences. We established a manipulative experiment in a common garden of Pinus pinaster including provenances typically showing either straight-stemmed or crooked-stemmed phenotypes. We imposed a bending up to 35º on plants aged nine years of both provenance groups and followed the straightening kinetics and shoot elongation after releasing. Eight months later, we destructively assessed biomass partitioning, reaction wood, wood microdensity, xylem reserve carbohydrates and phloem secondary metabolites. The experimental bending and release caused significant, complex changes with a marked difference between straight- and crooked-type plants. The straight-type recovered verticality faster and to a higher degree and developed more compression wood, while displaying a transitory delay in shoot elongation, reducing resource allocation to defences and maintaining the levels of non-structural carbohydrates compared to the crooked type. This combination of responses indicates the existence of intraspecific divergence in the reaction to mechanical stresses which may be related to different adaptive phenotypic plasticity.

Anatomical Characteristics and Ecological Trends in the Xylem and Phloem of Brassicaceae and Resedacae

IAWA Journal ◽  
2006 ◽  
Vol 27 (4) ◽  
pp. 419-442 ◽  
Author(s):  
Fritz Hans Schweingruber
Keyword(s):  
Western Europe ◽  
Life Forms ◽  
Reaction Wood ◽  
Lower Altitude ◽  
Alpine Zone ◽  
Brassicaceae Species ◽  
Ray Cells ◽  
The Alps ◽  
The Hill ◽  
Vestured Pits

The xylem and phloem of Brassicaceae (116 and 82 species respectively) and the xylem of Resedaceae (8 species) from arid, subtropical and temperate regions in Western Europe and North America is described and analysed, compared with taxonomic classifications, and assigned to their ecological range. The xylem of different life forms (herbaceous plants, dwarf shrubs and shrubs) of both families consists of libriform fibres and short, narrow vessels that are 20–50 μm in diameter and have alternate vestured pits and simple perforations. The axial parenchyma is paratracheal and, in most species, the ray cells are exclusively upright or square. Very few Brassicaceae species have helical thickening on the vessel walls, and crystals in fibres. The xylem anatomy of Resedaceae is in general very similar to that of the Brassicaceae. Vestured pits occur only in one species of Resedaceae.Brassicaceae show clear ecological trends: annual rings are usually distinct, except in arid and subtropical lowland zones; semi-ring-porosity decreases from the alpine zone to the hill zone at lower altitude. Plants with numerous narrow vessels are mainly found in the alpine zone. Xylem without rays is mainly present in plants growing in the Alps, both at low and high altitudes. The reaction wood of the Brassicaceae consists primarily of thick-walled fibres, whereas that of the Resedaceae contains gelatinous fibres. The frequency of sclereids in Brassicaceae bark is an indicator of ecological differences: sclereids are rare in plants from the alpine zone and frequent in plants from all other ecotones.

Analyse dendrochronologique d'un glissement de terrain de la région du Lac à l'Eau Claire (Québec nordique)

1985 ◽  
Vol 22 (2) ◽  
pp. 175-182 ◽  
Author(s):  
Christian Bégin ◽  
Louise Filion
Keyword(s):  
Tree Ring ◽  
Forest Regeneration ◽  
Climatic Conditions ◽  
Reaction Wood ◽  
Growth Rings ◽  
Glissement De Terrain ◽  
Landslide Occurrence ◽  
Slow Progression ◽  
Ring Analysis

A landslide in Clearwater Lake has been dated to spring of 1933 from tree-ring analysis (reaction wood, growth suppression, and corrosion scars). From the 52 sampled trees, seven peak periods of movement were registered within the site before landslide occurrence: 1785, 1815, 1827, 1829, 1852–1853, 1871–1872, 1897, and 1926. After a slow progression lasting 200 years, the slope movements accelerated in 1926, as indicated by suppressed growth rings. It is proposed here that the landslide was the outcome of a long-term slope development partly controlled by climate (precipitation). Postdisturbance forest regeneration (between 1950 and 1976) on the newly exposed substrate is also related to climatic conditions.

Negative gravitropism of Ginkgo biloba: growth stress and reaction wood formation

Holzforschung ◽  
2016 ◽  
Vol 70 (3) ◽  
pp. 267-274 ◽  
Author(s):  
Tatsuya Shirai ◽  
Hiroyuki Yamamoto ◽  
Miyuki Matsuo ◽  
Mikuri Inatsugu ◽  
Masato Yoshida ◽  
...  
Keyword(s):  
Ginkgo Biloba ◽  
Lignin Content ◽  
Linear Regression Analysis ◽  
Multiple Linear Regression Analysis ◽  
Wood Formation ◽  
Growth Stress ◽  
Reaction Wood ◽  
Cellulose Content ◽  
S2 Layer ◽  
Negative Gravitropism

Abstract Ginkgo (Ginkgo biloba L.) forms thick, lignified secondary xylem in the cylindrical stem as in Pinales (commonly called conifers), although it has more phylogenetic affinity to Cycadales than to conifers. Ginkgo forms compression wood-like (CW-like) reaction wood (RW) in its inclined stem as it is the case in conifers. However, the distribution of growth stress is not yet investigated in the RW of ginkgo, and thus this tissue resulting from negative gravitropism is still waiting for closer consideration. The present study intended to fill this gap. It has been demonstrated that, indeed, ginkgo forms RW tissue on the lower side of the inclined stem, where the compressive growth stress (CGS) was generated. In the RW, the micorofibril angle in the S2 layer, the air-dried density, and the lignin content increased, whereas the cellulose content decreased. These data are quite similar to those of conifer CWs. The multiple linear regression analysis revealed that the CGS is significantly correlated by the changes in the aforementioned parameters. It can be safely concluded that the negative gravitropism of ginkgo is very similar to that of conifers.

Sign in / Sign up

Export Citation Format

Share Document