A SAMPLING SYSTEM FOR EGGS OF WESTERN BLACKHEADED BUDWORM, ACLERIS GLOVERANA (WALSINGHAM) (LEPIDOPTERA: TORTRICIDAE), ON WESTERN HEMLOCK, TSUGA HETEROPHYLLA (RAF.) SARG.

1990 ◽  
Vol 122 (3) ◽  
pp. 555-562 ◽  
Author(s):  
R.F. Shepherd ◽  
T.G. Gray
Keyword(s):  
Sequential Sampling ◽  
Lower Surface ◽  
Tsuga Heterophylla ◽  
Western Hemlock ◽  
Sampling System ◽  
Relative Contribution ◽  
Egg Density ◽  
Branch Weight ◽  
Branch Size ◽  
Criteria For Selection

AbstractEggs of western blackheaded budworm, Acleris gloverana (Walsingham), are laid on the lower surface of western hemlock, Tsuga heterophylla (Raf.) Sarg., needles. A comparison was made of the following measures of sample branch size as a basis for expressing egg density: fresh branch weight, branch area, total twig length, branch volume, and number of buds. The criteria for selection of these measures were as follows: correlations of branch size with dry needle weight, variances of egg density and their relative contribution to sample size, and ease of measurement. Fresh branch weight was the best choice. A sequential sampling system was developed on this basis and was related to a scale of predicted defoliation. In addition, a transformation was provided for use in data analysis.

DISTRIBUTION OF EGGS OF WESTERN BLACKHEADED BUDWORM, ACLERIS GLOVERANA (WALSINGHAM) (LEPIDOPTERA: TORTRICIDAE), AND OF FOLIAGE OVER THE CROWNS OF WESTERN HEMLOCK, TSUGA HETEROPHYLLA (RAF.) SARG.

1990 ◽  
Vol 122 (3) ◽  
pp. 547-554 ◽  
Author(s):  
R.F. Shepherd ◽  
T.G. Gray
Keyword(s):  
Tsuga Heterophylla ◽  
High Variability ◽  
Weight Basis ◽  
Western Hemlock ◽  
Uneven Distribution ◽  
Crown Area ◽  
Sample Bias ◽  
Branch Weight ◽  
Oviposition Sites

AbstractEggs of western blackheaded budworm, Acleris gloverana (Walsingham), were found on the underside of needles of western hemlock, Tsuga heterophylla (Raf.) Sarg., with higher densities in the mid-crown area toward the outer tips of branches. Fringe and dominant trees were preferred oviposition sites over trees in lower crown classes or shaded positions. Vertical differences in needle density were considerable and led to high variability in insect densities when expressed on a per branch basis. When densities of eggs were based on a fresh branch weight basis there was a significant decrease in sample bias caused by the uneven distribution of foliage over the crowns.

Growth ofPseudotsuga menziesiiandTsuga heterophyllaseedlings along a light gradient: resource allocation and morphological acclimation

1997 ◽  
Vol 75 (9) ◽  
pp. 1424-1435 ◽  
Author(s):  
D. Mailly ◽  
J. P. Kimmins
Keyword(s):  
Leaf Morphology ◽  
Biomass Accumulation ◽  
Tsuga Heterophylla ◽  
Douglas Fir ◽  
Western Hemlock ◽  
Low Light ◽  
Growing Seasons ◽  
Light Levels ◽  
Light Gradient ◽  
Relative Light Intensity

Silvicultural alternatives that differ in the degree of overstory removal may create shady environments that will be problematic for the regeneration of Douglas-fir (Pseudotsuga menziesii (Mirb.) Franco). Gradients of light in the field were used to compare mortality, growth, and leaf morphological acclimation of two conifer species of contrasting shade tolerances: Douglas-fir and western hemlock (Tsuga heterophylla (Raf.) Sarg.). Results after two growing seasons indicated that Douglas-fir mortality occurred mainly at relative light intensity (RLI) below 20%, while western hemlock mortality was evenly distributed along the light gradient. Height, diameter, and biomass of the planted seedlings increased with increasing light for both species but at different rates, and maximum biomass accumulation always occurred in the open. Douglas-fir allocated more resources to stem biomass than western hemlock, which accumulated more foliage biomass. Increases in specific leaf area for Douglas-fir seedlings occurred at RLI ≤ 0.4 and red/far red (R/FR) ratio ≤ 0.6, which appear to be the minimal optimum light levels for growth. Conversely, western hemlock seedlings adjusted their leaf morphology in a more regular pattern, and changes were less pronounced at low light levels. These results, along with early mortality results for Douglas-fir, suggest that the most successful way to artificially regenerate this species may be by allowing at least 20% of RLI for ensuring survival and at least 40% RLI for optimum growth. Key words: light, light quality, leaf morphology, acclimation.

TOXICITY OF SMOKE FROM SPOROPHORES OF THE INDIAN PAINT FUNGUS AND WOOD FROM WESTERN HEMLOCK TO YELLOWFEVER MOSQUITOES

1987 ◽  
Vol 119 (12) ◽  
pp. 1109-1115
Author(s):  
W.P.L. Osborn ◽  
J.H. Borden
Keyword(s):  
British Columbia ◽  
Aedes Aegypti ◽  
Combustion Products ◽  
Tsuga Heterophylla ◽  
Water Soluble ◽  
Western Hemlock ◽  
Aqueous Extracts ◽  
Smoke Water

AbstractTo mitigate the effects of mosquitoes, settlers in the Revelstoke area of British Columbia reportedly burned the sporophores of the Indian paint fungus, Echinodontium tinctorium (Ell. & Ev.) Ell. & Ev., a pathogen of western hemlock, Tsuga heterophylla (Raf.) Sarg., and true firs, Abies spp. Larval and adult yellowfever mosquitoes, Aedes aegypti (L.), were exposed to aqueous extracts of smoke (smoke-waters) from E. tinctorium sporophores, and from western hemlock sapwood and heartwood. Smoke-waters were of approximately equal toxicity to larvae. Fungus smoke-water, but not sapwood or heartwood smoke-waters, lost 50% of its potency in 5 months. Vapors from fungus smoke-water were significantly more toxic to adult mosquitoes than those from sapwood or heartwood. Thus smoke from E. tinctorium sporophores and T. heterophylla wood apparently contain different water-soluble combustion products toxic to A. aegypti.

Genotype × shade effects for western hemlock

2005 ◽  
Vol 35 (6) ◽  
pp. 1496-1501
Author(s):  
G R Johnson ◽  
C Cartwright
Keyword(s):  
Tsuga Heterophylla ◽  
Western Hemlock ◽  
Different Levels

Western hemlock (Tsuga heterophylla (Raf.) Sarg.) families were grown under different levels of shade for 2 or 3 years at two nursery sites to determine whether families performed differently relative to one another in the different shade environments. Differences were found both for levels of shade and families, but no family × shade interaction was found. Results suggest that families selected in full-sun environments (clearcuts or farm fields) may be well suited for use in silvicultural systems where seedlings are planted in understory conditions.

Survival dynamics of mechanically topped Douglas-fir (Pseudotsuga menziesii) and western hemlock (Tsuga heterophylla) snags in Douglas-fir plantations, Oregon, USA

2019 ◽  
Vol 433 ◽  
pp. 105-110
Author(s):  
Matthew E. Hane ◽  
Andrew J. Kroll ◽  
Aaron Springford ◽  
Jack Giovanini ◽  
Mike Rochelle ◽  
...  
Keyword(s):  
Pseudotsuga Menziesii ◽  
Tsuga Heterophylla ◽  
Douglas Fir ◽  
Western Hemlock

STUDIES IN FOREST PATHOLOGY: VII. DECAY IN WESTERN HEMLOCK AND FIR IN THE FRANKLIN RIVER AREA, BRITISH COLUMBIA

1949 ◽  
Vol 27c (6) ◽  
pp. 312-331 ◽  
Author(s):  
D. C. Buckland ◽  
R. E. Foster ◽  
V. J. Nordin
Keyword(s):  
British Columbia ◽  
Tsuga Heterophylla ◽  
Western Hemlock ◽  
Forest Pathology ◽  
Abies Amabilis ◽  
Armillaria Mellea ◽  
Forest Region ◽  
Volume Increment ◽  
Juan De Fuca

An investigation of decay in western hemlock (Tsuga heterophylla (Raf.) Sarg.) and fir (mainly Abies amabilis (Loud.) Forb.) in the Juan de Fuca forest region of British Columbia has shown that the major organisms causing root and butt rots are the same in both species. These are Poria subacida (Peck) Sacc., Fomes annosus (Fr.) Cke., Armillaria mellea Vahl ex Fr., Polyporus sulphureus Bull. ex Fr., and P. circinatus Fr. Those organisms causing trunk rots of western hemlock, in decreasing order of importance, are Fomes pinicola (Sw.) Cke., F. Pini (Thore) Lloyd, Stereum abietinum Pers., Fomes Hartigii (Allesch.) Sacc. and Trav., and Hydnum sp. (H. abietis). These same organisms causing trunk rots of fir, in decreasing order of importance, are Fomes pinicola, Stereum abietinum, Hydnum sp. (H. abietis), Fomes Pini, and Fomes Hartigii. The logs of 963 western hemlock were analyzed in detail. Maximum periodic volume increment was reached between 225 and 275 years of age. Maximum periodic volume increment was reached between 275 and 325 years of age in the 719 fir that were analyzed. Scars were the most frequent avenue of entrance for infection. In 59% of the cases of infection studied the fungus had entered through wounds.

Canopy disturbances over the five-century lifetime of an old-growth Douglas-fir stand in the Pacific Northwest

2002 ◽  
Vol 32 (6) ◽  
pp. 1057-1070 ◽  
Author(s):  
Linda E Winter ◽  
Linda B Brubaker ◽  
Jerry F Franklin ◽  
Eric A Miller ◽  
Donald Q DeWitt
Keyword(s):  
Pacific Northwest ◽  
Variable Density ◽  
Tsuga Heterophylla ◽  
Douglas Fir ◽  
Western Hemlock ◽  
Old Growth ◽  
The Pacific ◽  
History Of ◽  
Horizontal Heterogeneity ◽  
Restoration Strategies

The history of canopy disturbances over the lifetime of an old-growth Douglas-fir (Pseudotsuga menziesii (Mirb.) Franco) stand in the western Cascade Range of southern Washington was reconstructed using tree-ring records of cross-dated samples from a 3.3-ha mapped plot. The reconstruction detected pulses in which many western hemlock (Tsuga heterophylla (Raf.) Sarg.) synchronously experienced abrupt and sustained increases in ringwidth, i.e., "growth-increases", and focused on medium-sized or larger ([Formula: see text]0.8 ha) events. The results show that the stand experienced at least three canopy disturbances that each thinned, but did not clear, the canopy over areas [Formula: see text]0.8 ha, occurring approximately in the late 1500s, the 1760s, and the 1930s. None of these promoted regeneration of the shade-intolerant Douglas-fir, all of which established 1500–1521. The disturbances may have promoted regeneration of western hemlock, but their strongest effect on tree dynamics was to elicit western hemlock growth-increases. Canopy disturbances are known to create patchiness, or horizontal heterogeneity, an important characteristic of old-growth forests. This reconstructed history provides one model for restoration strategies to create horizontal heterogeneity in young Douglas-fir stands, for example, by suggesting sizes of areas to thin in variable-density thinnings.

STUDIES IN FOREST PATHOLOGY: XII. DECAY OF WESTERN HEMLOCK IN THE UPPER COLUMBIA REGION, BRITISH COLUMBIA

1954 ◽  
Vol 32 (1) ◽  
pp. 145-171 ◽  
Author(s):  
R. E. Foster ◽  
H. M. Craig ◽  
G. W. Wallis
Keyword(s):  
Site Index ◽  
Tsuga Heterophylla ◽  
Western Hemlock ◽  
Site Quality ◽  
Advanced Stage ◽  
Multiple Correlation ◽  
Relative Percentage ◽  
Total Incidence ◽  
Average Defect ◽  
Mature Stands

An investigation of western hemlock (Tsuga heterophylla (Raf.) Sarg.) in the Upper Columbia region has shown that decay losses are excessive, amounting to 52 and 74% in terms of cubic and board foot computations respectively. The logs of 833 hemlock on 36 sample plots located in mature timber were analyzed in detail. Twenty-six decay-producing fungi have been isolated. Sixty-two per cent of the total decay was attributed to Echinodontium tinctorium Ell. & Ever. and 25% to Fomes pini (Thore) Lloyd. Considerable variation in the fungi responsible for decay was encountered intraregionally. Stand defect losses varied from 38 to 71%. Some of this variation could be predicted on the basis of the site index determinations of hemlock. Relative to the total incidence and importance of decay, Echinodontium tinctorium decreased and Fomes pini increased in significance with increasing site quality. Sporophores, scars, dead tops, large rotten branches, swollen knots, frost cracks, forked trees, and trunk infections of mistletoe were classed as indicators of significance in relation to hidden defect. Living trees, classed as Suspect or Residual depending on the presence or absence of one or more reliable indicators of defect, were found to differ appreciably in average defect. Within each of the preceding classes decay increased progressively with increasing diameter and decreasing site quality. Immature hemlock was found to be susceptible to appreciable decay and mature stands reached an advanced stage of deterioration at 250 years. Multiple correlation analyses between percentage of decay, site, age, and the relative percentage of Residual trees weighted by volume permitted an estimate of total stand defect within ± 7%.

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