Middle Miocene (Badenian) macroinvertebrates from Pécs-Danitzpuszta (Mecsek Mts, SW Hungary)

This paper examines Badenian (middle Miocene) macroinvertebrates – corals and molluscs – from the Pécs-Danitzpuszta sand pit (Mecsek Mts, SW Hungary) in order to extend our knowledge on Miocene normal marine deposits of the Mecsek region. Corals occur reworked in the upper Miocene sand that was deposited in the brackish Lake Pannon, and presumably originate either from the middle Badenian Pécsszabolcs or the upper Badenian Rákos Member of the Lajta Formation. A total of seven taxa were identified. These taxa suggest subtropical conditions and a lack of coral reefs in the Badenian. Molluscs were found in situ in the upper Badenian Szilágy Clay Marl Member of the Baden Formation and the Rákos Member of the Lajta Formation. They dominantly consist of bivalves and represent benthic assemblages typical of the middle Miocene Central Paratethys.

Calcareous nannofossils from the middle/upper Miocene succession of Pécs-Danitzpuszta, southern Hungary: cosmopolitan Paratethys and endemic Lake Pannon assemblages

Quantitative analyses on calcareous nannofossils were carried out on 109 middle/late Miocene (Sarmatian/ Pannonian) samples from the section at Pécs-Danitzpuszta sand pit (Hungary). The lower part of the section, which can be assigned to the Sarmatian, contains normal marine low-diversity assemblages dominated by Calcidiscus leptoporus, Reticulofenestra pseudoumbilicus, Sphenolithus moriformis and Syracosphaera spp. accompanied by didemnid ascidian spicules (Perforocalcinela fusiformis). The middle/late Miocene (Sarmatian/Pannonian) boundary is characterized by the last occurrences of normal marine calcareous nannofossils. The upper part of the section (Pannonian) can be subdivided into intervals characterized by monospecific endemic nannofossils Isolithus spp. and ascidians, respectively. A short interval with common endemic coccoliths belonging to the family Noelaerhabdaceae (Bekelithella echinata, Noelaerhabdus bozinovicae, N. jerkovici, Praenoelaerhabdus banatensis) in the upper part of the profile was also documented. The drastic change in nannofossil assemblages at the Sarmatian/Pannonian boundary is a result of paleoenvironmental stress caused by the isolation of the Central Paratethys from the Eastern Paratethys.

A diverse Miocene fish assemblage (Chondrichthyes and Osteichthyes) from the Pécs-Danitzpuszta sand pit (Mecsek Mts, Hungary)

Chondrichthyans and osteichthyans are widely reported from marine sediments of the Central Paratethys, not only by sporadic occurrences, but also by complex, diverse fish assemblages. Here we present a rich fish fauna from the upper Miocene (Pannonian, Tortonian) lacustrine sediments exposed in the Pécs-Danitzpuszta sand pit, in the SW Pannonian Basin. Altogether 22 227 specimens were investigated, and they could be classified into 17 chondrichthyan and 16 osteichthyan taxa. Among the chondrichthyans Odontaspididae (55.51%) and Myliobatidae (14.4%) are the most abundant, while the bony fishes are dominated by the Sparidae (77.07%). The limonitic, yellow, coarse-grained, gravelly sands yielded an extremely large amount of isolated fish fossils. A considerable part of the vertebrate material of the sands is likely reworked from older, middle Miocene (Badenian and Sarmatian, i.e. Langhian and Serravallian) sediments. The late Miocene calcareous marls underlying the sands also yielded a variety of fish remains. From these remains, associated and articulated latid bones are coeval with the sediment and suggest freshwater and brackish conditions in the area, in accord with other biotic data. Other, isolated fossils, namely teeth, otoliths, cycloid scales and jaw elements of Gadidae, Gobiidae, Sparidae and Latidae could be reworked and thus have a very limited paleoecological significance. Based on habitat preferences of extant relatives, all chondrichthyans and most osteichthyan taxa found in the sands must be of Badenian origin. The abundant remains allow for some conclusions on the environmental conditions during the Badenian. They refer to a shallow, coastal environment with tropical-subtropical climate with connection to more open water habitats. Remains of some osteichthyan taxa were found in coprolites, showing that these taxa were part of the food chain as prey items. Sirenian ribs and odontocete limb bones bearing tooth marks refer to trophic relations between marine mammals and large sized macropredatory sharks. The Pécs-Danitzpuszta record of shark-attributed bite marks on bones of marine mammals is the second of the Badenian of the Central Paratethys. Acipenserids and latids of the sands, linked to brackish and/or freshwater environments, could originate from any Miocene stratigraphic units.

Various marginal marine environments in the Central Paratethys: Late Badenian and Sarmatian (middle Miocene) marine and non-marine microfossils from Pécs-Danitzpuszta, southern Hungary

The middle Miocene foraminifera and ostracod record of the Central Paratethys usually reflects stable normal marine epositional environments for the Badenian and more patchy, less stable restricted marine environments for the Sarmatian. A 17 m thick outcrop at Pécs-Danitzpuszta, Mecsek Mts, SW Hungary exposed an upper Badenian to Pannonian succession where foraminifers and ostracods document significant environmental changes. The basal layers of the section contain micro- and macrofossils indicating normal marine, shallow, warm, well-oxygenated habitat with relatively high-energy conditions and algal vegetation on the bottom, and represent the upper Badenian (13.82 to 12.65 Ma). The marine deposits are followed by coarse sandstone, breccia and siltstone layers barren of microfossils but containing rhizoliths. The sediments were probably subaerially exposed for some time. The following marine inundation, marked by the appearance of clays and limestones as well as fossils, was dated to the late Sarmatian (ca. 12 to 11.6 Ma) on the basis of the restricted marine microfossil assemblages from the upper part of the succession (Porosononion granosum Zone, Aurila notata Zone). This community is characterized by exclusively eurytopic forms indicating an unstable and vegetated marginal marine environment with fluctuations in salinity, as well as oxygen and food availability. Within the 5 m thick upper Sarmatian marine interval, a unique fresh- to oligohaline fauna characterizes a few layers in less than 1 m thickness. This fauna consists of highly euryhaline foraminifera and freshwater to oligohaline ostracod assemblages, indicating a temporary salinity reduction to 5–10 ‰. No similar freshwater fauna has been reported from the Sarmatian of the Central Partethys so far. The eventual disappearance of the foraminifera from the paleontological record coupled with a complete turnover in the ostracod fauna indicates the transition from the marginal marine Sarmatian Sea to the brackish Lake Pannon, marking the Sarmatian/Pannonian boundary (11.6 Ma).

A new deepwater lobster, Metanephrops serendipitus sp. nov. (Crustacea, Decapoda, Nephropidae), from lower Miocene of Meljski hrib (Maribor, Slovenia)

A new species of nephropid lobster, Metanephrops serendipitus sp. nov., is described from the lower Miocene (Ottnangian/Karpatian) in beds of Central Paratethys. Metanephrops serendipitus sp. nov. represents the first fossil representative of the genus from the northern hemisphere and thus extends its known palaeobiogeographical distribution. Additionally, the palaeoecology of the new species is discussed. Similar to extant species of Metanephrops, the fossil nephropid inhabited a fine-grained deep-sea environment, associated with frequent brittle stars.

An overlooked diversity—the Costellariidae (Gastropoda: Neogastropoda) of the Miocene Paratethys Sea

The Miocene Costellariidae of the Paratethys Sea are revised. In total, 39 species in seven genera are recorded herein, documenting an unexpected diversity of these marine gastropods. The α-diversities of up to 14 species and basin-scale γ-diversities of up to 28 coeval species reveal the Central Paratethys as a unique diversity hot-spot for Miocene Costellariidae. Palaeobiogeographic relationships with the adjacent Proto-Mediterranean Sea are low at species level, and absent for Eastern Paratethyan and north-eastern Atlantic faunas. Antithala nov. gen. and Fedosovia nov. gen. are described as new genera. Bellardithala nov. nom. is introduced as new name for Micromitra Bellardi (1888) [non Meek, 1873]. Eight species are described as new: Antithala claviformis nov. sp., Antithala filipescui nov. sp., Bellardithala kovaci nov. sp., Bellardithala baluki nov. sp., Bellardithala fedosovi nov. sp., Bellardithala dacica nov. sp., Pusia confunda nov. sp., and Vexillum transalpinum nov. sp. Lectotypes are designated for Mitra badensis Hoernes & Auinger, 1880, M. brevior Friedberg, 1911, M. fuchsi Hoernes & Auinger, 1880, M. intermittens Hoernes & Auinger, 1880, M. januszkiewiczi Friedberg, 1928, M. lapugyensis Hoernes & Auinger, 1880, M. laubei Hoernes & Auinger, 1880, M. michelottii Hörnes, 1852, M. moravica Hoernes & Auinger, 1880, M. neugeboreni Hoernes & Auinger, 1880, M. partschi Hörnes, 1852, M. pseudorecticosta Boettger, 1906, M. sturi Hoernes & Auinger, 1880, M. szobbiensis Halaváts, 1884.

40Ar/39Ar geochronology of Burdigalian paleobotanical localities in the central Paratethys (south Slovakia)

The Lipovany and Mučín paleobotanical localities contain important floral associations within the tuff horizons, which were used for determination of subtropical to tropical climatic conditions during the Early Miocene. Based on the combination of results from plagioclase and biotite 40Ar/39Ar dating, the age of the tuff deposition is around 17.3Ma. For the Lipovany locality, single-grain 40Ar/39Ar convergent ages of 17.49±0.54Ma and 17.28±0.06Ma, for plagioclase and biotite were obtained, respectively. The Mučín locality only provide an imprecise convergent age of 16.5±1.4Ma due to the small size of the analyzed plagioclase crystals. The results thus allowed to include the fossil subtropical flora of the studied localities in the late Ottnangian regional stage (upper part of the Burdigalian). Additionally, these age data indicate that deposition of the overlaying Salgótarján Formation starts much later than originally thought (during Ottnangian-Karpatian boundary).