GROWTH AND MORPHOGENESIS IN THE CANADIAN FOREST SPECIES: V. FURTHER STUDIES OF WOOD GROWTH IN BRANCHES AND MAIN AXIS OF PINUS RESINOSA AIT. UNDER CONDITIONS OF OPEN GROWTH, SUPPRESSION, AND RELEASE
The analysis of growth in trunk and branches of four red pine trees of different situations is extended to include internode length, circumference, annual wood volume increment, and cambial area. The effects of position of a branch on the tree and of suppression and release are examined in reference to these growth indexes.Increase in circumference responds in much the same way as radial growth, and branch position is important. The greatest annual increase in girth of any branch internode occurs when it is near the periphery and near the top of the tree.Terminal growth of branches is less affected than radial or tangential growth by branch position if the tree is in the open but is suppressed by surrounding trees. Upon release of a tree only those axes whose local environment was actually improved showed an increase in apical growth.The interpretation of annual wood volume increment is complicated by the fact that it is determined not only by internal and external conditions during the current growing season, but also by past growth, both apical and cambial. It is subject to the effects of suppression and of branch position, chiefly through the influence of these factors on the cambial growth component.