Among the largest Plio-Pleistocene hominin skulls found to date, A.L. 444-2 is bigger, though not by much, than an average female gorilla’s skull. At first glance, A.L. 444-2 assumes a somewhat simian appearance, the outcome of a relatively small braincase combined with an inclined frontal squama and prognathic jaws. However, this apelike appearance is offset by several distinctive hominin features: a very tall face that is much less prognathic than would be expected from the skull’s general simian-like appearance; a deep, vertical mandibulosymphyseal profile; delicate supraorbital elements; and the absence of a supratoral sulcus intervening between the frontal squama and the forward-jutting supraorbital element. Nevertheless, the characteristics that account for the skull’s hominin appearance demonstrate a certain uniqueness, which is manifested in the disproportion between the considerable total height of the face and the great size of its constituent elements (primarily the zygomatic and maxillary bones), on the one hand, and the delicateness of the supraorbital element and the almost negligible degree of its anterior projection, on the other. An apparent unevenness emerges along the vertical axis of the face between its upper portion—the orbits, including the elements above and between them—and its lower portion, that is, the elements below the level of orbitale down to gnathion. Undoubtedly, part of this appearance stems from the heavy, somewhat vertical, deep, and anteriorly bulbous symphyseal region of the mandible. The corresponding region in the African apes, in contrast, is transversely pinched, as its two sides converge downward toward the midline. Furthermore, the region slopes inferoposteriorly; in anterior view, it is tucked under the alveolar element and hence is less exposed than in A.L. 444-2. The preservation of the mandible of A.L. 444-2 and its occlusion with the upper dental arcade afford a unique opportunity to evaluate some of the characteristics of an entire A. afarensis skull. Two standard measurements can be recorded: the distance between gnathion and the estimated site of nasion—a measure of the total height of the face—which is 150 mm, and the distance between gnathion and basion, estimated at 157 mm.