Wood density in Norway spruce: changes with thinning intensity and tree age

2005 ◽  
Vol 35 (7) ◽  
pp. 1767-1778 ◽  
Author(s):  
Tuula Jaakkola ◽  
Harri Mäkinen ◽  
Pekka Saranpää

The effect of thinning intensity on growth and wood density in Norway spruce (Picea abies (L.) Karst.) was investigated in two long-term thinning experiments in southeastern Finland. The stands were approaching maturity, and their development had already been studied for 30 years. The intensities of thinning were low, normal, and high (i.e., the stand basal area after the thinning was, on average, 40, 27, and 24 m2·ha–1, respectively, in Heinola, and 30, 28, and 17 m2·ha–1 in Punkaharju, respectively). Compared with the low thinning intensity, the normal and high thinning intensities increased the basal-area increment of individual trees by 52% and 68%, respectively. Normal and high thinning intensities resulted in a relatively small reduction (1%–4%) of mean ring density compared with low thinning intensity. The random variation in wood density between and within trees was large. About 27% of the total variation in wood density was related to variation between rings. Our results indicate that the prevailing thinning intensities in Norway spruce stands in Fennoscandia cause no marked changes in wood density. At least, the possible reduction in wood density is low compared with the increase in individual tree growth.

Forests ◽  
2020 ◽  
Vol 11 (4) ◽  
pp. 416 ◽  
Author(s):  
Oskars Krisans ◽  
Roberts Matisons ◽  
Steffen Rust ◽  
Natalija Burnevica ◽  
Lauma Bruna ◽  
...  

Storms are the main abiotic disturbance in European forests, effects of which are expected to intensify in the future, hence the importance of forest stand stability is increasing. The predisposition of Norway spruce to wind damage appears to be enhanced by pathogens such as Heterobasidion spp., which reduce stability of individual trees. However, detailed information about the effects of the root rot on the stability of individual trees across diverse soil types is still lacking. The aim of the study was to assess the effect of root rot on the individual tree stability of Norway spruce growing on drained peat and mineral soils. In total, 77 Norway spruce trees (age 50–80 years) growing in four stands were tested under static loading. The presence of Heterobasidion spp. had a significant negative effect on the bending moment at primary and secondary failure of the tested trees irrespectively of soil type. This suggests increased legacy effects (e.g., susceptibility to pathogens and pests due to fractured roots and altered water uptake) of storms. Damaged trees act as weak spots increasing the susceptibility of stands to wind damage, thus forming a negative feedback loop and contributing to an ongoing decline in vitality of Norway spruce stands following storms in the study region in the future. Accordingly, the results support the importance of timely identification of the decayed trees, lowering stand density and/or shortening rotation period as the measures to counteract the increasing effects of storms on Norway spruce stands.


2005 ◽  
Vol 35 (1) ◽  
pp. 113-121 ◽  
Author(s):  
Kjell Karlsson ◽  
Lennart Norell

The probability that an individual tree will remain in even-aged Norway spruce (Picea abies (L.) Karst.) stands subjected to different thinning programmes was modelled, using data from a thinning experiment established in 25 localities in southern Sweden. A logistic regression approach was used to predict the probability and the Hosmer–Lemeshow goodness-of-fit test to evaluate the fit. Diameter at breast height (DBH), quadratic mean DBH, thinning intensity, thinning quotient, basal area, number of stems per hectare, stand age, number of thinnings, and site index were used as explanatory variables. Separate analyses for stands thinned from below, stands thinned from above, and unthinned stands were performed. The modelled probability graphs for trees not being removed, plotted against their diameter at breast height, had clear S-shapes for both unthinned stands and stands thinned from below. The graph for stands thinned from above was bell-shaped.


2010 ◽  
Vol 40 (5) ◽  
pp. 843-849 ◽  
Author(s):  
John B. Bradford ◽  
Anthony W. D’Amato ◽  
Brian J. Palik ◽  
Shawn Fraver

Growth dominance is a relatively new, simple, quantitative metric of within-stand individual tree growth patterns, and is defined as positive when larger trees in the stand display proportionally greater growth than smaller trees, and negative when smaller trees display proportionally greater growth than larger trees. We examined long-term silvicultural experiments in red pine ( Pinus resinosa Ait.) to characterize how stand age, thinning treatments (thinned from above, below, or both), and stocking levels (residual basal area) influence stand-level growth dominance through time. In stands thinned from below or from both above and below, growth dominance was not significantly different from zero at any age or stocking level. Growth dominance in stands thinned from above trended from negative at low stocking levels to positive at high stocking levels and was positive in young stands. Growth dominance in unthinned stands was positive and increased with age. These results suggest that growth dominance provides a useful tool for assessing the efficacy of thinning treatments designed to reduce competition between trees and promote high levels of productivity across a population, particularly among crop trees.


Holzforschung ◽  
2010 ◽  
Vol 64 (1) ◽  
Author(s):  
Tuula Jyske ◽  
Seija Kaakinen ◽  
Urban Nilsson ◽  
Pekka Saranpää ◽  
Elina Vapaavuori

AbstractThe effects of thinning intensity on wood structure and chemistry of Norway spruce [Picea abies(L.) Karst.] have been studied in a long-term thinning experiment established in the 1960s in south-western Sweden. The experiment comprised datasets of unthinned control trees and trees of four thinning treatments, in which thinning operations of different intensities were carried out. These thinning operations were applied either once, three, or five times over the course of the stand rotation. Thinning intensities varied from 0% to 60% based on the stand basal area (BA) before thinning: (A) 5×20% (i.e., 20% of the BA removed five times); (B) 3×40% (i.e., 40% of the BA removed three times); (C) 1×60% (i.e., 60% of the BA removed once); and (D) 5×40% (i.e., 40% of the BA removed five times). Altogether 60 trees that were 67 years old were sampled 38 years after the treatment onset. Radial growth rate, wood density, tracheid properties, and the chemical composition of wood were studied in detail. All four thinning treatments increased growth rate and decreased wood density; however, treatment D showed the greatest change. Single, heavy thinning in treatment C resulted in unfavourable intra-stem variation in ring width and wood density. No significant differences in tracheid properties and wood chemistry were found between the treatments. We conclude that only a remarkable increase in growth rate would induce detrimental changes in tracheid properties and wood chemistry in the context of thinning.


1996 ◽  
Vol 26 (10) ◽  
pp. 1838-1848 ◽  
Author(s):  
P. Lejeune

A model predicting circumference growth of individual trees has been developed for mixed irregular stands dominated by Fagussilvatica L. in southeastern Belgium. This model integrates simultaneously tree, stand, and growing site characteristics without considering distances between individual trees. It has a determination coefficient of 40.1% and a residual standard deviation of 0.45 cm/year. Forest mensuration variables considered in the model are circumference, tree social position represented by the total basal area of trees greater than the subject tree, stand basal area, and some index of stand structure corresponding to the ratio of the variance over the mean circumference. The site component is essentially expressed by the length of the growing period. The low accuracy obtained for individual tree growth is relative because the results are used mainly after individual trees have been distributed into size classes. We have been able to show that the introduction of crown description in the model can increase its accuracy. But this modification requires a submodel describing crown changes over time.


2015 ◽  
Vol 45 (8) ◽  
pp. 1006-1018 ◽  
Author(s):  
Sonja Vospernik ◽  
Robert A. Monserud ◽  
Hubert Sterba

We examined the relationship between thinning intensity and volume increment predicted by four commonly used individual-tree growth models in Central Europe (i.e., BWIN, Moses, Prognaus, and Silva). We replicated conditions of older growth and yield experiments by selecting 34 young, dense plots of Norway spruce (Picea abies (L.) Karst.), Scots pine (Pinus sylvestris L.), and European beech (Fagus sylvatica L.). At these plots, we simulated growth, with mortality only, to obtain the maximum basal area. Maximum basal area was then decreased by 5% or 10% steps using thinning from below. Maximum density varied considerably between simulators; it was mostly in a reasonable range but partly exceeded the maximum basal area observed by the Austrian National Forest Inventory or the self-thinning line. In almost all cases, simulated volume increment was highest at maximum basal area and then decreased with decreasing basal area. Critical basal area, at which 95% of maximum volume increment can be achieved, ranged from 0.46 to 0.96. For all simulators, critical basal area was lower for the more shade-tolerant species. It increased with age, except for Norway spruce, when simulated with the BWIN model. Age, where mean annual increment culminated, compared well with yield tables.


2014 ◽  
Vol 44 (3) ◽  
pp. 173-181 ◽  
Author(s):  
Matthew G. Olson ◽  
Spencer R. Meyer ◽  
Robert G. Wagner ◽  
Robert S. Seymour

Understanding the response of tree regeneration following commercial thinning treatments can improve planning in managed forests dependent on natural regeneration. We used long-term commercial thinning experiments in eastern spruce–fir stands of Maine, USA, to test two hypotheses: (1) commercial thinning increases the density of tree regeneration and (2) tree regeneration density increases with increasing thinning intensity. A decade after thinning, densities of softwood and hardwood regeneration were 10 times greater in thinned stands than unthinned stands. The abundance of small softwood (0.11–0.60 m tall) was highest in lower intensity thinning treatments, whereas medium (0.61–1.40 m tall) and large (≥1.41 m tall to 8.90 cm diameter at breast height) softwoods increased proportionally with thinning intensity, a pattern related to a higher rate of recruitment in more open stands created by heavier thinning. Hardwood density generally increased with thinning intensity and developed into a significant component of the large size class. Softwood regeneration density was higher in older spruce stands than younger fir stands, which may be due to greater abundance of advance regeneration, higher residual stand mortality, and greater harvest disturbance in older spruce stands. However, acceptable softwood stocking was achieved in all replicates of thinning treatments. Therefore, in addition to providing higher individual-tree growth and merchantable yield, commercial thinning in eastern spruce–fir stands also increases regeneration density. The rate of recruitment also increased as thinning intensity increased, thus stimulating understory regeneration similar to that of a shelterwood establishment cut.


Forests ◽  
2021 ◽  
Vol 12 (7) ◽  
pp. 841
Author(s):  
Iveta Desaine ◽  
Annija Kārkliņa ◽  
Roberts Matisons ◽  
Anna Pastare ◽  
Andis Adamovičs ◽  
...  

The increased removal of forest-derived biomass with whole-tree harvesting (WTH) has raised concerns about the long-term productivity and sustainability of forest ecosystems. If true, this effect needs to be factored in the assessment of long-term feasibility to implement such a drastic forest management measure. Therefore, the economic performance of five experimental plantations in three different forest types, where in 1971 simulated WTH event occurred, was compared with pure, planted and conventionally managed (CH) Norway spruce stands of similar age and growing conditions. Potential incomes of CH and WTH stands were based on timber prices for period 2014–2020. However, regarding the economics of root and stump biomass utilization, they were not included in the estimates. In any given price level, the difference of internal rate of return between the forest types and selected managements were from 2.5% to 6.2%. Therefore, Norway spruce stands demonstrate good potential of independence regardless of stump removal at the previous rotation.


2021 ◽  
Vol 13 (12) ◽  
pp. 2297
Author(s):  
Jonathon J. Donager ◽  
Andrew J. Sánchez Meador ◽  
Ryan C. Blackburn

Applications of lidar in ecosystem conservation and management continue to expand as technology has rapidly evolved. An accounting of relative accuracy and errors among lidar platforms within a range of forest types and structural configurations was needed. Within a ponderosa pine forest in northern Arizona, we compare vegetation attributes at the tree-, plot-, and stand-scales derived from three lidar platforms: fixed-wing airborne (ALS), fixed-location terrestrial (TLS), and hand-held mobile laser scanning (MLS). We present a methodology to segment individual trees from TLS and MLS datasets, incorporating eigen-value and density metrics to locate trees, then assigning point returns to trees using a graph-theory shortest-path approach. Overall, we found MLS consistently provided more accurate structural metrics at the tree- (e.g., mean absolute error for DBH in cm was 4.8, 5.0, and 9.1 for MLS, TLS and ALS, respectively) and plot-scale (e.g., R2 for field observed and lidar-derived basal area, m2 ha−1, was 0.986, 0.974, and 0.851 for MLS, TLS, and ALS, respectively) as compared to ALS and TLS. While TLS data produced estimates similar to MLS, attributes derived from TLS often underpredicted structural values due to occlusion. Additionally, ALS data provided accurate estimates of tree height for larger trees, yet consistently missed and underpredicted small trees (≤35 cm). MLS produced accurate estimates of canopy cover and landscape metrics up to 50 m from plot center. TLS tended to underpredict both canopy cover and patch metrics with constant bias due to occlusion. Taking full advantage of minimal occlusion effects, MLS data consistently provided the best individual tree and plot-based metrics, with ALS providing the best estimates for volume, biomass, and canopy cover. Overall, we found MLS data logistically simple, quickly acquirable, and accurate for small area inventories, assessments, and monitoring activities. We suggest further work exploring the active use of MLS for forest monitoring and inventory.


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