Analysis of variation in the Elaphrus americanus complex of North America (Coleoptera: Carabidae)
Elaphrus americanus Dejean (sensu Lindroth, 1961) is a complex of two nearctic species: E. americanus and E. finitimus Casey. This study is primarily devoted to E. americanus, which includes two subspecies: a boreal subspecies and a newly described western subspecies, E. americanus sylvanus Goulet n. subsp., for which the type locality is Oregon, Coos Co., 16 miles north of Powers.The species and subspecies mentioned above were recognized by means of numerical taxonomic analysis. Most intuitively acceptable were the results of phenetic clustering of Mahalanobis distances followed by a series of discriminant analyses. Various analyses were carried out based directly on computed euclidean distances, e.g. principal coordinate analyses, nonmetric multidimensional scaling, and still others by computing first the variance–covariance matrix, e.g., principal component analyses. Other geographically distinct populations were recognized by univariate analysis of nominal characters.Phylogenetic relationships between various populations of E. americanus are partly reconstructed. A reconstruction of past distribution of these populations is presented and their evolutionary significance discussed.